fact that they are part of a series of typical variant,
show
that they are probably this species.The
specimens in the Missouri series are uniquein havingtheband
considerably reduced onthe proximaland
distalmargins,butits
maximum
widthis stillgreaterthanthe length ofthe marginal vein. TheseHopkins
lots, however, agree in having thehead rounded
in front, face receding below antennal fossae, setaeon submarginal vein usually 11 to 14,and
in theirgeneral color pattern.Dr. A. C.
Kinsey
provided five femalesfrom
the galls of Plagio- trochus chrysolepidicola(Ashmead),
var,undetermined,from pugnvs
Kinsey, in one instance on Q. dumosa^Paso
Robles, Calif.,and
againfrom
thesame
gall atNapa,
Calif.; alsofrom
thesame
gall at Exeter, Calif.,on
Q. lohata^and
othersfrom
Manitou, Colo.,from
P. frequens frequens (Gillette)on
Q. UTidulata.The
rearing dates for the cynipid gallmakers
of these Kinsey lots are in Feb- ruary,March, and
April, 1920.Several other California specimens were received through the courtesy of Dr. Isabel
McCracken from Leland
Stanford Junior University collection (Lot554, Sub. Ill)and
rearedby
herfrom
the galls of Disholcaspis eldoradensis (Beutenmueller)on
Q. lohata^ on the Stanford campus, April 1915.I have not been able to
examine
specimens ofDecatoma
kelloggi Fullaway. DoctorMcCracken
kindlycompared some
ofmy Kinsey
specimens with the type of kelloggi
and
reports that these differ incolorfrom
thetype. D. va7'iansWalsh,
however,variesextremely in color,Fullaway
states that the head of kelloggiis "fairlythick anterior-posteriorly,"and
the frontwing
has "abrown
gourd- shaped cloud extending across the middle of thewing
two-thirdsits width."
His
specimenswere
rearedby
Mrs.Rose
PattersonBlakeman from
a "twig swelling " gall on Quercus chrysoleyis^the galls collected at Stevens Creek, Santa Clara County, Calif. It
is probable that this gall is a close relative, if not a
member,
of the genus PlagiotrochiLS,from which what
I regard as variansWalsh
has been reared.The head and wing
characters quoted above are essential features of varians,and
altogether it is very likely that D. kelloggiFullaway
is identicalwith Walsh's species.The
outstanding features of varians are:The
greatly variablebody
color, the hindfemora
usually mostly ochreous-yellow out- wardly, cheeksreticulate punctate, not polishedand
smooth, usually 13 to 15 setaeon
the submarginal vein,and
the finely granulose sculpturing ofat leastthe sidesof the posterior half of theabdomen.
It is also remarkable for its wide geographic distribution,
and
the broad range of genera of cynipid gallson
oakfrom which
it has been obtained.38 PKOCEEDINGS OF THE
NATIONAL,MUSEUM
vol.79There
issomuch
variation inthecolorofthisspeciesfrom
Illinois, the type locality, that specimensfrom
theWestern United
States readilycome
within the extremes of the color differences.At
firstinclined to look
upon
thewestern lots,as well assome from
Illinois, as representing several distinct species on account of both host and color differences, Iam
at last compelledby
the lack of structural characters toregardthem
all asone. Thissame
inconstancy of colorand
absence of morphological charactersmake
it impossible to recognize varieties. These differences cut across the generic lines of the cynipid hostsand
do not correlate withany
geographic boun- dary,lines.13. DECATOMA DUBIA Walsh Plate1, Figubb12
Decatomavariansvar. duhiaWalsh,Amer. Ent. and Bot., vol.2, no. 10, p. 300, 1870.
Readily separated
from
D. variansWalsh
withwhich
it has been frequently confused,by
its larger size, generally darker color, the narrowerand
longer curved submarginal band,and
the distinctly relatively shorterand
widerhead which
ismuch
less produced in front than in vanaiis.Female.
— Length
3.2 to 4.3mm.,
usually of the larger size, fairly robust,mostly black above,some
partsofbody
yellow to rufousand brown;
head black, exceptnarrow
circumocular band, templesand
cheeks mostly,malar
space, antennalscrobeand
an oblique barfrom
it to eye,
and
face at times only below antennaeand
at times toand
includingmuch
of vertex, yellow or rufous-yellow,head
almost entirely black in afew Michigan
specimens; scape light ochreous- yellow, pedicelsame
butmore
orless fuscous above, ringjointyellow, flagellum chestnut-brown; sidesofpronotum
yellow likethat ofhead, topinmost
cases black,but at times onlymedian
third,or even only posterior two-thirds of this area, black, rest yellow; mesothorax black,lowerparts ofscapulaeand
axillaeand
the pleura deepyellow, rufous, orbrown
to partly black; metathoraxand propodeum
black;
legs mostly yellow, front coxae immaculate to black on anterior surface, middle
and
hind coxae sometimes rufous-yellowand
sparselymarked
black,femora
yellow, disks of outer surfaces sparselyand
feebly to broadlyand
deeply brown-black, tibiae light yellow, fore tibiae at times immaculate, at times faintly brown, or front pair faintly,and
others increasingly brown, the hind pair deeply so except baseand
apex, tarsi faint yellowish white; stigma almost black, submarginalband
deep dark brown, usually abouttwo and
a half times as long as wide, itswidth
almostuniform
throughout, rarely broader than length of stigma, crooked in form, bending distinctlytoward
body, reaching two-thirds across wing, its apexART.28
CHALCID
FLIESNORTH OF MEXICO BALDUP 39
rounded,but sometimesnarrowed and
roughly tapered; petiole black,abdomen
mostly black, only about cephalo-ventral fourth brownish to mostly yellow.Head
mostly just one half as long as wide, sometimes slightly longerand
narrower,transversely ovate-rectangular, frontextending butfeeblyaheadofeyes,occiputonly slightlyconcave, vertex laterad of posterior ocelli finely wrinkledand
punctate, malar spacesmooth
wrinkled, notpunctured,restofhead coarselyreticulate witha small umbilicate puncture in each reticulation,and
moderately hairy;scape of antenna as long as pedicel,
Fl and
half ofF2
combined, pediceland Fl
about equal in length,F2
toF4
equally longand
thick, length of each one-fifth greater than diameter, club
compact and
slightly dilatedand
as long asF4 and F5
combined;pronotum
one-seventh narrower at its widest point thanhead and
mesothorax,upper
surfaces of prothoraxand
mesothorax denselyand
rather coarselyumbilicatepunctate;propodeal groovedeep,broad,rounded, crossed in front
by
a prominent carina; surface ofpropodeum
coarselyrugosereticulate,more
so in front of the carina;submarginal vein with 13 (rarely) to 19, frequently 18, setae;abdomen
shiny,smooth
polished, except sometimes thirdand
fourth segments partly coriaceous, bare, exceptbeyond
third segment, latter with a transverserow
of hairsinthe middle, posterior surface ofabdomen around
ovipositor sheath moderately hairyand
punctate, sheath extending prominently above tip ofabdomen.
Male.
— On
the average smallerthan the female, butin color,vesti- ture, sculpturing,and form
like the female, except the usual differ- enceintheshapeof theabdomen and
theantennae;Fl
aboutone-sixth longer than pedicel,F2
barely or not at all longerthanF3 and
F4,F2
toF4
each one- fourth longerthanthick,club feebly dilatedand
as long asF3 and F4
combined.Remarks. — The
originalWalsh
specimens of duhia are perhapsdestroyed.
One
female in theUnited
States NationalMuseum
col- lection is labeled ^^Decatovia dubia Walsh," probably in Walsh's handwriting, but there isno
date, locality, or host given toshow
whether or not itwas
used in preparing the original description.Only
five femalesand
onemale
were originally athand when
theform was
described.They
were rearedby
DoctorWalsh from
the gallsof Disholcaspismamma
(Walsh) onthemossycup
oak {Qiiercus macrocarpa).
This species is well represented
from
areas rangingfrom
Arizonaand Texas
through the Mississippi Valley to the AtlanticOcean
from
the northern to the southern limits of theUnited
States.But
considering the distribution of its variety doaneiFuUaway,
duhia occurs also in California areas of the country.Whenever
host recordsaccompany
the specimens they indicate that duhia inhabits40 PROCEEDINGS OF THE NATIONAL MUSEUM
vol.79mostly the galls (only stem gall
known
as hosts to date) of various species of Disholcaspison
differentoak
species of the Leucobalanus group.A few
recordsfrom
Belonocnetna are at hand. Specimens that Ihave identified as this speciesare presentas follows: 6 femalesand
3 malesfrom
stem galls of Disholcaspis inavirria(Walsh) on
Q. macrocarpa^Urbana,
111., April 20 to June, 1923,and May
20 toJune
18, 1924,and from
Catlin, 111., 1 femalefrom
the stem gall of D. globulus (Fitch) onswamp
chestnutoak
{Q,. michauodi)^ reared April 15, 1929; also 4 femalesand
4 malesfrom Urbana,
111., butfrom
stem gall ofD.
globulus (Fitch) on Q. alba,May and
June, 1923; the aforementioned materialwas
rearedby
myself; 1 of each sex probably rearedby
DoctorWalsh
at Rockford, 111.,and
1female labeled "through A. Bolter, 1890." L.
H. Weld
sentme
4 of each sex bearing No. 38and
rearedby him from D. mmnma
(Walsh)
at Evanston, 111.,and
4 femalesfrom
thesame
hostand
with thesame number
reared at Manistee, Mich.,June
15, 1907; 7 femalesand
2 males (U.S.N.M. coll.) were originally obtainedfrom
theMichigan
Agricultural College,some
rearedJune
28and
29, 1887,from
" red-oak lesser globular leaf-gall," Ace. Cat. 738,and
others with Ace. Cat. Nos. 857(March
23, 1888)and
900(May
28, 1887)came from
the galls ofD.
maiivina(Walsh) and D.
globulus (Fitch), respectively,and
one with Ace. Cat. No.720A
(June 23, 1887) isfrom
"bastard-oak apple gall,"and
another bearing data"Mich.,903" withouthost recordsisthis species; 2 females (189'"'^) that issued in April, 1895, 1 labeled "
on
Holcaspis globulus "were reared
by
J. G.Barlow
at Cadet, Mo., both probablycoming from
thestem gall ofD.
globulus (Fitch);from
thesame
placeand
collector, are 3 specimens (389''°^) datedJuly 26, 1883, with
no
host records available; 1 female (2610°*), probably this species,was
reared onMay
4, 1882,from
a cynipidoak
gall receivedfrom H. K.
Morrison,Fort Grant,Ariz.,
on March
23 of thesame
year; 1 femalefrom
"Fitch's Collection,"and
2 of this sexon
one point labeled"Westcott" are without question dubia; 4 females
and
1male
(165''°-), reared
June
28, 1883,from
the gall of Belonocnetna fossoriaWeld
[Dishol^easpisvirens(Ashmead)] on
Q. virens receivedfrom H.
G.Hubbard,
Crescent City, Fla.; 2 maleson
one pin (165) reared onMarch
13, 1875,from
galls of B. fossoriaWeld
collectedby Muhleman from
Q.virens (?) on October 31, 1874; 3 malesfrom
thesame
species of gall but on Q. stellaia (obtu^iloba), the galls collectedby
Patton at Selma, Ala., in October, 1880,and
the insects issuingon December
16 of that year; 1male
(165''°) isfrom Heavue,
Tex., another without locality or hosts(165")
with label"
on
oak.Mar.
13, 1875,"and
a thirdwas from an
"undeterminedoak
gall "; 2 femalesand
1male from Texas
(Belfrage coll.) ; 10 fe-males
and
3malesrearedby
C. N.Ainsliefrom
thegallsofD.munima
AST.28
CHALCID
FLIESNORTH
OFMEXICO BALDUF 41 (Walsh)
onmossy cup oak
{Q. maorocarpa) ; I have 3 additional femalesfrom Mr.
Ainslie takenby him by
sweeping alfalfa at Salt Lake, Utah,and
bearingAVebsterNo.5595,but theyare smallerthan typicalduh'm\3males (66"°') issuedfrom
gallsofD.globulus (Fitch),
June
27, 1883, the galls receivedfrom
A. T. Packard, Providence, R. I.,on
April 7 of thesame
year; 2 females (66''°^) datedMarch
4, 1884, bear
no
other data; 1 female labeled "Melrose Highlands, Mass.,July7,"and
thefollowingHopkins
lots: 5 femalesand
amale(Hopk. U.
S. 13686«), reared April 5, 12,and
29, 1918,by
L.H.
Weld, from
galls of Disholcaspis spongiosaKarsch on
Q.minor
at one ormore
of these localities: Thebes, Liberty County, Ga.,and
Palestine, Trinity, orHouston, Tex.; 1 female
(Hopk. U.
S. 10774a) rearedMay
14, 1919,by
S. A.Rohwer, from
undetermined galls on Q. minor^ at Ironton,Mo.;
a pair(Hopk. U.
S. 10776a),same
collector
and
locality, rearedJune
11 (male)and
18 (female), 1919,from
galls of D. fasciata Bassett (determined S. A.Rohwer)
on Q.ruhra; 1 female
(Hopk. U.
S. 10777a) rearedby
S. A.Rohwer,
Poplar Bluff, Mo.,June
4, 1919,from
gallon
Q. minor;and
2 pairs(Hopk. U.
S. 10777b),from
undeterminedgallson Q. minor^ Poplar Bluff,Mo.,the malesissuingJune
18and
July 2,and
the females onMay
14and June
26, 1919; 5 femalesand
4 males labeled Worcester, Mass.,and
Providenceand Rumford,
R. I.,sentme by
DoctorKinsey from
theThompson
collection,had
been rearedfrom
the galls of D.Tnamma (Walsh) and
D. globulus (Fitch) ; I have a fine series of 20 femalesand
2 males rearedby
DoctorKinsey from
galls ofD.
mamina
var.on
Q. hicoloratBroadway,
N.J., onSeptember
4, 1921.From
the collection of theCanadian Department
of Agriculture, Ottawa, I have 2 femalesfrom
bullet galls onmossycup
oak \_D.mamma (Walsh) on
Q. macrocarpa'] collectedby
G. E. Sanders atTampico,
111.,on May
15and
20,and
July26, 1909; also one female labeled "on
officewindow. Mar.
3, 1906, J. F.";and
one of each sex rearedfrom
the gall ofD.
globulus (Fitch) at Guelph, Ontario,by
T. D. J.14.
DECATOMA
DUBIA RUFOSA, new yarietyPlate1, Figxjbe13
Differschiefly
from
the typicalform
ofD. dubiaWalsh
in having the headand
thorax almost or entirely reddish yellow.Female.
—
3 to 4.1mm.
long, colors chiefly rufousand
black;head
rufous, sometimes vertexand
also occiput abovemore
or less dark brown, antennae chestnut-brown; prothorax goldenbrown
to light reddish yellow; mesothorax concolorous withhead
but sometimes front of prescutum, lower edge of axillae posteriormargin
ofscutel,and
margins of dorsal sclerites narrowly black;metanotum and
propodeum
rufous to almost black; legs entirely very pale yellowish42 PROCEEDINGS OF THE NATIONAL MUSEUM
vol.79brown,
hind
tibiae at timesdusky brown
as in dubia,' submarginalband
in color, size,and form
as in dubia;abdomen
darkbrown
to black, except that antero-ventral part is in small part lightbrown
to brownish yellow.
The
dimensions of the head, themeasurements
of the antennal joints, the sculpturingand
the vestiture of the parts, including thenumber
of setaeon
the submarginal vein, agree with the typical dubia.Male.
—
Differsfrom
the female only in being slightly smaller on the average,and
in the dimensions of the antennal joints, the latter agreeing with those of dubia.Type
locality.—
Charleston, S. C.r2/;?e.—Female,
U.S.N.M.
No. 42235.Described
from
the type, the allotype,and
five female paratypes;type, allotype,
and
three paratypesin collection of theUnited
States NationalMuseum, and two
paratypesinthe writer's collection.Remarks. —
All the type materialwas
rearedfrom
the galls ofBelonocneina fossoria
Weld
[Disholcaspis virens(Ashmead)]
at Charleston, S.C, by
J. T.Rogers and
Charles Pon.One
specimen of the series bears alabel "Rohwer
2015"and
"Pon
287."In
addition I find the following of this variety: 6 females (No.165°^^)
from
the galls ofBelonocnema
fossoriaWeld
[Disholcas- pisvirens(Ashmead)] on
Quercusstellata (obtusiloba)^ collectedby
Patton at Selma, Ala., in October, 1880,and
the chalcids issuedDecember
16 of that year; another series (165^°^) of 4 females,June
28, 1883, arefrom
thesame
cynipidand oak
hosts,from
Cres- cent City, Fla. (H. G.Hubbard);
3 females (165^"^^ ^re probablyfrom
thesame
hostsand
region; 1 females (165''°) islabeledHeavue,
Tex.; anotherfrom
post oak {Q. stellata),June
10; 1 ofsame
sex labeled "Mo., 715^°,June
6, 1883"; amale
with data " J. L. Zabris- kie,Nyack, N. Y.,Feb. 6, 1884"; 3 femalesand
amale from
eastern Florida,Ashmead
; 3 femalesand
a male,some
with undecipherable records in pencil, others withno
data; onefrom
Doctor Kinsey, rearedfrom
the gall of Disholcaspis viam/ma var.on
Q. bicolor atBroadway,
N. J.; a single female rearedby
myselffrom
the stem gall of D.mamma (Walsh)
on Q.macrocarpa
atUrbana,
111.,on June
18, 1924; 1 femalefrom
gall of D. globulus (Fitch)on
Q.alba, April 24, 1918, Falls Church, Va., reared
by William
Middle- ton; L.H. Weld
reared 1femaleofthisvariety(Hopk.
U. S. 13686«)
from
the gall ofDisholcaspis spongiosaKarsch on
Q.minor
at one of the following localities: Thebes, Liberty County, Ga.,and
Pales- tine, Trinity,and
Houston, Tex.The
rearing date isMay
15, 1918, whereas sixtypical females ofD. dubiaWalsh
issuedfrom
thesame
galls on April 12, 1918.
ABT.28
CHALCID
FLIESNORTH
OFMEXICO —
-^BALDUF43
It ispossible that this is a distinct species, but it has been reared
from
thesame
lots of gallsfrom which some
of the dubla material came,and
there issome
evidence that duhia properand
its variety rufosa intergrade in color.A
larger series, especially additional materialfrom
the States intermediatebetween Illinoisand Alabama,
is neededto determine the extent of the color variation. There are
no
recognizable differences in size, sculpturing, vestiture, or dimen- sions of thehead
or dimensions of the antennal segments.15.DECATOMADUBIA var.DOANEI Fullaway Plate 2, Figuee 14
Decatoma doanei Fuixaway, Journ.
New
York Ent. Soc, vol. 20, pp. 279, 280, 1912.Most
like the typical D. duhiaWalsh
in color, size, dimensions of thehead and
the antennal segments,and form
of the submarginalband
; it is darkerand
smalleron
the average,and
usually has fewer setaeon the submarginal vein.Female.
— Length
3.2 to 4mm.,
colors yellowand
black, mostlyblack;
head
black, circumocular spaceand
an obliqueband from
it toantennae,malar
area,and
face below antennal fossaeand
at times abovethem halfway
to vertex, yellowish to reddishbrown;
scapedark
yellow, in part lightly infuscate, pedicel yellow like scape, but mostly brownish black above, flagellum dark chestnut-brown; pro- thorax always mostly black above, its sidesand
anterior facemore
orlessyellowish brown, sometimes a black spot in thelateral depres- sion,on some
specimens a faint line of this colorfrom
near middle in front obliquely to frontend
of parapsidal grooves; mesothorax, metathorax,and propodeum
black, except mesopleura, tegulae,and narrow
area at lower edges of scapulaeand
axillae,which
are reddishbrown;
coxae black except sometimes the apex, or a stripeon
posterior lateral face, outer disks of foreand
middle femora, all of hindfemora
except ends,and
front tibiae usually brownish yellow, middle tibiae lightly to moderatelybrown
on outer face,and
hind tibiaebrown
exceptatbaseand
apex,tarsi light to whitish yellow; stigmadark
brown, submarginalband medium
brown, slightlymore
than twice as long as wide, width not quite uniform, but notmuch
broader ormore narrow
than length of stigma, apex curving stronglytoward
base of wing, distal edge reaching almost or quite as far as stigmal vein, apex rounded;abdomen
black, only antero-ventral portionmore
or less yellowish brown.Head
moderately hairy, very nearly twice as broad as long, face almost flat or slightlyconvex, vertex weakly convex, occiput slightly concave,malar
spacesmooth
in groove only, rest finely coriaceous or sculptured like face, vertex in partaround
anterior ocellus only44 PROCEEDINGS OF THE NATIONAL MUSEUM
vol.79 finely punctate, occiput, face,and
cheeks with rather coarser punc- tures than thorax, or subreticulate; antennae rather stout, scape as long as pedicel, Fl,and
half of F2, moderately dilatedtoward
base, pediceland Fl
aboutequalinlength,latterone-fifth longerthan each ofF2
to F5, latter each about one-fifth longer than itsmaximum
thickness, length of club equal to that of
F3
toF5
combined;width
ofhead and
mesothorax nearly equal,pronotum
about six-sevenths aswide
as mesothorax, dorsal punctures of thorax coarsely umbilicate, bearing small white hairs; 13 to 16 setae on submarginal vein;abdomen
mostlysmooth and
bare,segmentsbeyond
the second with afew
hairs,more around
ovipositor sheath, latter also coria- ceous, thirdand
fourthmore
orless with fine wrinkles, especiallyon
lower lateral surface.Male.
—
Like the female in color, vestiture,and
sculpturing, but in general slightly smallerand
slenderer,from
3.2 to 3.6mm.
long.The
ratio ofhead
dimensions is thesame
in both sexes.Antennae
slenderer than in the female, scape gradually dilatedtoward
base,Fl
slightly longer than pediceland
one-sixth longer than F2, F3, or F4, latter about equally long orF2
slightly longer thanF3
or F4,and
each nearly two-thirds as thick as long, club notmuch
thicker than funicle, scarcely dilated,and
as long asF3 and F4
combined.Type
locality.—
Jaspar Ridge, Stanford University campus, Calif.Type.
—
Describedfrom
one femalefrom
the gall of Disholcaspis eldoradeiisis (Beutenmueller) on Quercus dumosa., Stanford Univer- sity collection rearedby
Mrs.Rose
PattersonBlakeman.
Remarks. — The
description of themale
is basedon
one specimen kindly sentme by
Dr. IsabelMcCracken,
curator of the insect col- lection at Stanford University, California.That
collection con- tains afew
other specimens.These
lots of material bear recordnumbers
L. S. Jr.U. Lot
508, Subs. 19,45,and
508,and
were rearedby
DoctorMcCracken and
Mrs.Blakeman.
Doctor
Kinsey
sentme
the followingrearedfrom
California galls:
2 females
from
D. cam.escens (Bassett) on blue oak {Q. douglasi) atMerced
Falls,and
anotherfrom
thesame
hostsatEl
Portal; 1 femalefrom
D. eldoradensis var.on
California white oak {Q. lohata) at Diablo; 1 femalefrom
D. plumhellaKinsey
atUpland, and
anotherfrom
thesame
gall on Q. duniosa at Descanso;and
2 femalesand
amale from
B.corallina (Bassett)on
Q. douglasiatNapa
;thecynipid hosts of the Diablo lot were reared inNovember,
1923,and
those of the remaining series issued inFebruary and March
of 1920.The
collection of the
United
States NationalMuseum
contains 3 femalesfrom
the gallsofD. eldoradensis (Beutenmueller)on
Q. lohata taken at Cottonwood,Calif.,on January
22, presumably1925; 3 malesfrom
thesame
localityand
oak,and
probably thesame
gall,on January
1,1925; aseriesof4ofeachsex