SPECIES WRONGLY PLACED IN GENUS AND UNRECOGNIZABLE SPECIES

SPECIES.

Polysphincta acutaProvancher,Nat. Can.,voL12,1880, p. 44, female.

S.A,Rohwer,

who

has examinedthe type,says that

what

^is left ofit (the

abdomen

and apices ofthe antennaearegone) isverylike Olisfojjyga canadensis Provancher.

But

the shape of the

abdomen

aud the long ovipositor as described

by

Provanclier exclude ^itfrom that genus.

From

Polysphincta its long ovipositor and short first

No.2326.

THREE TRIBES OF ICHNEUMONINAE—CUSHMAN.

37 tergite differentiate^it. Exceptfor the lack of an areoletit might be an Epiurus; oreven withouttheareoletit

may

bea Tromatobia, thoughhere the shortfirsttergite isoutof place.

(PolyspMncta cingulataProvancher)=Tromatobia rufovariata(Cresson).

PolysphinctanigricepsWalsh,and

PolyspMnctanigrita\VAJ.s-R,Trans.Acad.Sci.,St.Louis, vol.3, 1873, p. 144.

The

typesof thelasttwospecieshave beendestroyedand it ispos- sible oidy to conjecture

what

they were.

From

Walsh's careful descriptions, however, it is evident that they were botli species of Zatypota andcloselyallied tonigrocephala (Davis).

PolyspMnctapimploidesWalspi, Acad.Sci.St.Louis,vol.3, 1873, p. 145, If this isproperly referred to the genus, asseems very doubtfid,

it isverydistinct

by

reasonoftheverylongovipositor. I

know

of no speciesthatapproaches itin this respect.

{PolyspMnctapleuralisProvanchek)=^(jBgssms)^Diplazonpulchripes (Provancher)

{PolyspMncta rubricapensis Provancher)

=

(Pimpla) Zaglyptus incompletus (Cresson).

{PolyspMncta spiaosaDAxif^)= {Pimpla)Zaglyptus incompletus (Cresson).

PolyspMncta rufigasira (Ashmead)=(Pm/,Za) Zaglyptus arrimosus ^(Cresson).

Isotypic,Ashmeadha^dngredeecribed Cresson's type.

ZABKACHYPUS, new genus.

Differs from Polysphincta principally

by

the characters used in the foregoingkey togenera. Its short, stout

body

\vithstoutlegs, rery short tarsi, deep sternauli, and incomplete pentagonal areolet renderitveryeasily recognizible.

Head

distinctly narrower than thorax; malar space shorter than basalwidth of mandible; antennae stout; thorax nearly ashigh as long: propodeimi veryshort, ^vithout either carinaeorlongitudinal groove;sternaulideep;legs stout,tarsi veryshortand thick;second intercubitus indicated

by

thickenings in cubitus and radius, the areolet irregularly pentagonal in position but open behind; first tergite broad with sides widely divergent; other tcrgites strongly transverse, elevations and impressions weak; exserted portion of ovipositorsub equal inlength to firsttergite.

Genotype.

—

Zahrachjpus

prlmm Cushman, new

species described below.

ZABRACHYPUSPBIMUS,newepecies.

Female.— Length6.5

mm

^„,antennae4.25

mm.,

ovipositor 1.0

mm.

Head

polished, ratherdenselypilose; face medially elevated and weakly punctured, about aslong as wide; clypeus half as long as wide, arcuately subtruncate, rather weakly convex; malar space barelyhalf aslongasbasalwidth, ofmandible;eyeslarge,convergent below, sinuate within; diameter of lateral ocellus, postocellar line,

38 PROCEEDINGS

OF.

THE NATIONAL MUSEUM.

^{vol. 58.}

and

ocell-ociilar line equal; temples convexly sloping; antennae stout, taperingslightly toward apex; thorax suhpolished, weakly punctate; notauli deep, complete, prescutum rather low; meta- pleura and

propodeum more

coarsely and densely punctate, the latterpolished medially; stigma narrow, radius inmiddle; nervulus postfurcal; nervellus perpendicular, stronglybrokenin middle;first tergite as broad atapex aslong, with anteriorbasinbroad and set off

by

rather strong carinae, with^'deep oblique'apical impressions

;

following tergitesrather densely, finelypunctured, elevationstrans- verse, apical tergites subpolished; exserted portion of ovipositor rather longer than first tergite.

Black; palpi white; clypeus, mandibles, and scapebelowpiceous;

antennae brown;tegulae yellow;wingsfaintlybrownish;legsmostly testaceous, coxae piceous, those of front and middle legs reddish toward apex; hind tibiaeand tarsifuscouswithsubbasalannuhis on tibia andbasal

anmdus

on basitarsus yellowish; apices offront and middlefemora, basesof their tibiae,and alltrochanters

orless yellow.

Typelocality.

—

^Montana.

Type.—Cat. No. 19172, U.S.N.M.

One

femalewithoutotherdata.

TribeTHERONIINI^Cushmanand Eohwer.

The

genera included in thistribeare TherordaHolmgren, Neothe- ronia Krieger, and Einmecoideus

Ashmead. The

firsttwohavehere- toforebeen placedinthe(Pimphni)Ichneumonini, while Aslmiead's genus

was

described in the Lissonotini. Pseudacoenites Kriech- baumer,

synonymous

with Theronia,

was

considered

by

its author to berelated toAcoenites Gravenhorst, but wasplaced

by Ashmead

in theLissonotini.

The

closest relative ofthe Theroniini

among

tlie tribesisprobably theE])hialtini, and from that tribe itis easily distinguished

by

^the form of the ovi])ositor, the slit-like propodeal spiracles with their prominentsurroundingcarinae, theshort,deep, parallel notauli, and thepolish.ed, usuallylight-colored body.

Sofar as definiteinformationinregardtothehost-relations ofthese insects is athand, theyare apparentlysecondary parasites through otherIchneumonidae,recordsof theirhaving beenrearedfromother Ichneumoninae and Ophioninae having been^published.

Description.

—

^Entire

^body

almost unsculptured, smooth andpol- ished; head transverse, narrow behind eyes, temples

or less

convex;face broaderthanlong,narrowed towardclypeus; eyeslarge,

or less distinctly emarginate or sinuate opposite antennae;

ocelli large; frons slightlyconcave; malar spacevery short;clypeus truncateorroundedatapex; antennae^long^filiform, scapeshort.and

No.232G.

THREE TRIBES OF ICHNEUMONINAE—CUSHMAN.

³⁹

strongly obliqueatapex, pedicel aboutaslong as scape,basaljoint offlagellum nearly twice aslong assecond, those

beyond

middle as broadas long;thorax rathershort,highanteriorly;notauli shortand parallel, deepanteriorlywith a ridgerunning backalong themargin of the lateral lobe; prepectus broad, the carina turning sharply toward the promesothoracic suture about half

way up

the pleura;

mesopleural suture angulate opposite the punctiform fovea;

scutellum strongly convex, margined only atbase;

propodeum

with at least avery strong apical carina, spiraclelong and slit-like, the surrounding carina high; wings large, stigma narrow lanceolate, radius before middle, areolet broadly sessile, nervellus strongly reclivousand broken farabove the middle;legs, especially posterior femora, stout, last tarsal joint inhindlegs aslong asfirst;

abdomen

fusiform, rathernarrow atbase;first tergitelongerthan broad and longerthansecond, others transverse, apical tergite short;ovipositor shorter than abdomen,subcylindrical, the lance nearlystraightdor- sally toapex;hypopygidiumretracted.

Generic characters.

— The

three genera are separable

by

^the

structural and colorcharacters used inthe followingkeyto genera.

Thesearethe only charactersofany importance^thathave i)een dis- covered;andoftheseonly two, the propodeal carinaeandtheinfuma- tion ofthewingapices, arewithoutexceptions withinoneoranother of the genera as represented in the material

exammed.

Krieger's very long and minute description of Neotheronia^{^} applies almost equallyas well,with thesetwoexceptions, to Theronia.

Even

these two characters are

orlesscomparative,for

some

specimensof Neotheronia have the

propodeum

flattened medially with obsolete ridgesinthe normalpositions ofthecarinae, and the infumationof the wings ⁱⁿ that genus varies in intensity, sometimes almost to absence.

KEYTOGENERA.

1. Propodeumbefore apical carinadi\idedintofiveareas,thelateralcarinasome- timesweak;prepectal carina onlyweakly curvedatsternauli,not subangulate whereitturnstowardthe promesothoracic suture, the sternauliweak; wings immaculate,stigmaandcosta usually red; facemoreor lesselevated in middle, withoutlongitudinal furrows;clypeua usuallyconvexbasallyand impressed

atapex ^Theronia Holmgren.

Propodeumwithlongitudinal carinaewantingbefore apical^carina;wings moreor lessinfumateat apex, costa blackish, stigma usually^so;face usuallyflatwith twomoreor less distinct longitudinal furrows; clypeus^flat,broadlyrounded

atapexand witha reflexedmargin ^2.

2. Prepectal carinaasin Theronia;firsttergitenearlyparallel-sided,aboutfour times as longaswideat apex,dorsal carinaeobsolete;wingsyellowwithbrilliant goldreflections,stigmaexceptcostalmarginred; face^slightlyelevated medially

and with very weak furrows EpimecoideusAshmead

1Krieger, R., Zeitschr.Uym.Dip., vol. 5,189.5,p. 3S9.

40 PROCEEDINGS OF THE NATIONAL MUSEUM.

^{vol. 58.}

Prepectal carinadeeply curved at sternauli and usually subangulatewhere it turnstowardpromesothoracicsutui-e;firsttergitebroaderwithsidesdivergent, dorsalcarinaedistinct;wingsatmostslightlystained,stigmausually blackish;

face usually flatwithstrong furrows, rarelyelevated mediallyand without

furrows Neotheronia Krieger.

Genus

THERONIA

Holmgren.

TheroniaHolmgren, Ofvers.Vet.-Acad. Forh., vol. 16, 1859, p. 123. Geno- type.

—

(PimplaJlavicans Fabricius)IchneumonatalantaePoda.

PseudacoenitesKreichbaumer,Ent.Nachr., 1892,p.219. Genotype.

—

_{Pseuda.

coenites moravicus KTiech.ha,um.eY)

=

TheronialaevigataTechek, according to Krieger, Zeitechr.Hym.Dip.,vol.3,1902,p.189-190.

As

indicatedabovethedifferencebetweenthisandtheothergenera of the Theroniini is comparatively little and largely a matter of difference in degree of development of certain features.

The

cari- nationof the

propodeum

and theimmaculate wingswill apparently, however, always distinguish Theronia from the other genera.

In our fauna thegenus isrepresented

by

but two species. These areverydistinct andeasily identified

by

the charactersemployed in the following key.

Both

species were originally described in the genus

Pimpla

Gravenhorst.

KEYTO NORTH AMERICANSPECIES.

Headlargely yellow,moreorlessrufous dorsallyandposteriorly;facenot tuberculate;weaklypunctate;firsttergiteinprofileevenlycurved above

fulvescens (Cresson).

Headblack, face stronglypunctateandmediallytuberculate;firsttergiteelevated

aboA'e melmiocephalu(BruU^).

THERONIA FULVESCENS(Cresson).

Although a fairly

common

species and though originally mis- placed in the genus Pim'pla, this species has apparently escaped redescription, anditwasnotreferred toitspropergenusuntil thirty- two yearslater,

when Howard

recorded the rearingof a single male asaparasite of

Hemerocampa

leucostigmagiving the authorasBrull6.

This mistake in authorship

was

not Howard's but Ashmead's, for thespecimenonwhichthe recordisbasedisoneoftheseriesexamined and was determined

by Ashmead.

Krieger's variety americana of the European atalantae (Poda), based on a specimen from British Columbia, is undoubtedly this species; and a male labelled

by

Viereck with Kj-ieger's

name

^is ^a normal male of this species. In his latest mention offulvescens Viereck^{^}treated itas a varietyof atalantae.

The

present speciesis perhapsnothing

than a geographical raceof atalantae, but judgingfrom a comparison of the materialat

hand

with 10 specimens of atalantae it differs in having the face slightly longer, the antennae fuscous above, the discocubital vein

1Hym.Conn.,1917, p. 323.

No.2326.

THREE TRIBES OF ICHNEUMONINAE—CUSHMAN.

orlessangulateand sometimes even withashort ramulus,and the ovipositor

than half aslongastheabdomen. ^In aialantae the antennae are concolorous above and below, the discocubital vein not at allangulate, and theovipositorlessthanhalf as longas theabdomen.

Face rounded but not distinctly tuberculate medially, weakly, sparsely punctate; eyes distinctly, though broadly, emarginate opposite antennae; postocellar line and diameter of lateral ocellus subequal; epomia weak; sternauli obsoletely impressed; lateral carinaeof

propodaum weak

basally,thecostellanotat alldeveloped, thespiracle

thanhalf aslongasheightof pleural area,^itslower end very close to pleural carina, median carinae parallel; discocu- bitalvein sinuatewitha

orlessdistinct angulationinthemid-

dle, not strongly curved;hind femur with ^adistinct scrobe for the reception ofthetibia, the scrobe flankedon the outside

by

a sharp ridge which is either scalloped or entire; fii'st tergite in profile uni- formly weaklyarched above,butlittlelongerthansecond.

Head

mostly yellow, antennaereddish-brown, paler below, scape andpedicelyellow; thorax brightrefescent varied withyellow below and onscutellum, thi-ee longitudinalstripes on

mesoscutum

^darker, sometimes

orless brownish, prepectuswitha

orless distinct brownishspot on eachside, metasterum and thoracic sutures also sometimes brownish;

abdomen

darker rufescent, sometimes

orlesspiceous;wings

orless yellow;legs stramineous to pale testaceous, the lighter color on the front and middle coxae andtrochanters andbaseofhindtibia, hindfemm*atbaseand apex

or less piceous, apical joint ofhindtarsus concolorous with otherjoints.

This is a very widely distributed species and subject to great variation, especially hisize and color ofwings, and Viereck has described a variety, mellipennis, said tohave theface medially finely rugulose and the wings strongly yellowish.

The

sculpture of the face varies

somewhat

intheseriesexamined, andseveralspecimens, including one from the

same

locality as Viereck's type, have very

faint,finerugulosityin the middle;butin the opinion ^oftheWTiter neither this nor thedepth of color of the wings is of even varietal significance. However,sinceViereck's t3rpehas not been examined the variety mellipennis is treated as valid and separable from the typicalform

by

the sculptureoftheface as follows

KETTO VARIETIES.

Face notorveryfaintlyregulosemedially fulvescens (Creseon).

Face mediallyrugulose mellipennis Viereck.

Dalam dokumen THE NORTH AMERICAN ICHNEmiON-FLIES OF THE TRIBES LYCORINI, POLYSPHINCTINI, AND THERONIINI. (Halaman 30-36)