A JOURNAL ON TAXONOMIC BOTANY,
PLANT SOCIOLOGY AND ECOLOGY
REINWARDTIA
A JOURNAL ON TAXONOMIC BOTANY, PLANT SOCIOLOGY AND ECOLOGY
Vol. 12(2): 129-204.22 November 2004
Editors
ELIZABETH A. WIDJAJA, MIEN A. RIFAI, SOEDARSONO RISWAN, JOHANIS P. MOGEA
Correspondence and subscriptions of the journal should be addressed to
REINWARDTIA Vol 12, Part 2, pp: 135 – 140
MISCELLANEOUS
NOTES
ON
MAINLY
SOUTHEAST
ASIAN
GRAMINEAE
J.F. VELDKAMP
Nationaal Herbarium Nederland, Universiteit Leiden branch, P.O. Box 9514, 2300 RA Leiden, The Netherlands. E-mail: veldkamp@nhn.leidenuniv.nl
ABSTRACT
VELDKAMP, J.F. 2004. Miscellaneous notes on mainly Southeast Asian Gramineae. Reinwardtia 12(2): 135 —
140. — During the preparation of a revision of the herbaceous grasses of SE Asia a few nomenclatural and taxonomic novelties were detected. New combinations are proposed in Aristida L., Kengia Packer, Rytidosperma
Steud., and Urochloa P. Beauv. Aristida novaeguineae Ohwi is re-instated as a distinct species. A key to the Malesian species of Aristida L. is provided. Moorochloa Veldk. replaces Brachiaria auct., s.s.
Key words: SE Asia, Gramineae, Aristida, Brachiaria, Cleistogenes, Kengia, Moorochloa, Rytidosperma, Urochloa.
ABSTRAK
VELDKAMP, J.F. 2004. Aneka catatan tentang Gramineae, terutama di Asia Tenggara. Reinwardtia 12(2): 135 —
140. — Telah ditemukan beberapa hal baru berkaitan dengan tata nama dan taksonomi rerumputan dari Asia Tenggara. Kombinasi baru diajukan untuk Aristida L., Kengia Packer, Rytidosperma Steud., dan Urochloa P. Beauv. Aristida novaeguineae Ohwi diposisikan ulang sebagai jenis tersendiri. Kunci identifikasi jenis-jenis
Aristida di Malesia diungkapkan. Moorochloa Veldk. menggantikan Brachiaria auct. s.s.
Kata kunci: Asia Tenggara, Gramineae, Aristida, Brachiaria, Cleistogenes, Kengia, Moorochloa, Rytidosperma, Urochloa.
INTRODUCTION
During the ongoing study of the Gramineae for the Flora Malesiana, the editorship of the Newsletter Grass Literature, and specimens to identify passing my desk, some notes were collected, which are presented here. When I offered some remarks on Aristida L. (Veldkamp, 1992) a key and additional information on the curious distribution of some of its species in Malesia should have been given, an omission which is corrected here.
There has been an extensive discussion on the generic delimitation of Brachiaria (Trin.) Griseb. against Urochloa P. Beauv. which caused me to make a proposal (1996) to lectotypify the first in the way as it is still generally accepted, and thus retain this well-known name for at least a few species. However, the Nomenclatural Committee for Spermatophyta (Brummitt, 1998) refused it and suggested that a new name (actually a new genus) should be published, with which wish is complied here. A species formerly included in Brachiaria is transferred to Urochloa.
The delimitation of Rytidosperma Steud. is still in doubt, which necessitated a new combination.
A SYNOPSIS OF MALESIAN ARISTIDA
1. a. Column absent (Sect. Aristida)..…..….…… 2 b. Column present (very short in A.
novae-guineae)………....….… 5 2. a. Lemma 6.25—15 mm long, callus 0.5—1.5 mm
long, lateral awns 9.5—35 mm long. — Lower glumes shorter than the upper ones…..……. 3 b. Lemma 1.5—5.6 mm long, callus 0.1--0.4 mm long, lateral awns 0—9 mm long………. 4 3. a. Annuals. Blades 0.5—0.8 mm wide. Panicle
axes scaberulous. Lower glume 4.75—5.8 mm long, mucro up to 0.5 mm long; upper glume 5.5—6.7 mm long, mucro up to 0.5 mm long. Lemma 6.25—7.5 mm long, callus c. 0.5 mm long, central awn 12--16.2 mm long, lateral awns 9.5—12.7 mm long. Anthers c. 1.5 mm long. — Savu, Sumba, Timor ……….…..…... ………. 1. A. adscensionis
b. Perennials. Blades 1—3 mm wide. Panicle axes smooth. Lower glume 9—17 mm long, mucro 2—5 mm long; upper glume 11—20 mm long, mucro 2—5 mm long. Lemma 9—15 mm long, callus 0.75—1.5 mm long, central awn 20—42 mm long, lateral awns 20—35 mm long. Anthers 2—2.5 mm long. — S Thailand (Satun), expected in N Malaya ……….. 11. A. setacea
136 REINWARDTIA [VOL.12
4. a. Perennials. Panicle few-spikeled, 20—30 by c. 5 cm. Lower glume 5.25—8.7 mm long, longer than the 4.75—6.4 mm long upper glume. Lemma 4—5.6 mm long, callus 0.3—0.4 mm long, central awn straight, lateral awns 4.25—9 mm long. Anthers 1.1—1.5 mm long. — Celebes, Mindanao……..…… 2. A. chinensis
b. Annuals. Panicle many-spikeled, 3—10 by 0.8—4 cm. Lower glume 1.4—2.5 mm long, shorter than the 2—3.5 mm long upper glume. Lemma 1.5—3 mm long, callus 0.1—0.2 mm long, central awn recurved, lateral awns 0(!)— 4 mm long. Anthers 0.1—0.6 mm long. — Celebes, Luzon, Mindanao, New Guinea (Irian Jaya: Kebar, Baliem, Swart Valley; Papua New Guinea: W Highlands, Morobe Prov.) ………… ………. 4. A. cumingiana
5. a. Lemma column articulated at base. — Basal internodes glabrous. Blades not coiled, 6—25 cm long. Panicle contracted, few-spikeled, 0.8—1 cm wide. Lower glumes 7.5—22 mm long. Lemma margins overlapping, callus 1—2 mm long, column 15—38 mm long, lateral awns well-developed, 24—70 mm long. Anthers 0.75—3 mm long. (Sect. Arthratherum)...… 6 b. Lemma column not articulated at base. —
Panicle axes glabrous. Glume apex acuminate to mucronate. Lemma 5.7—11 mm long…... 8 6. a. Panicle axes glabrous. Glumes apex acuminate to mucronate; lower glumes 7.5—14 mm long, shorter than the upper ones. Lemma 8—10 mm long. Anthers 1.5—3 mm long. — Luzon, Buru, New Guinea .…...…...…...…...….. 7 b. Panicle axes sparsely pilose. Glumes acute; lower glumes 13.5—22 mm long, longer than the upper ones. Lemma 4.5—6 mm long. Anthers c. 0.75 mm long. Culms 0.15—0.4 m long. Central awn straight, 50—70 mm long. — Sumba……..…….……… 10. A. polyclados
7. a. Culms 0.15—0.4 m long. Glumes mucronate. Central awn straight, 55—72 mm long. Anthers c. 1.5 mm long. Lower glumes 9—14 mm long. — Luzon….... 5. A. holathera var. holathera
b. Culms 0.6 — 1 m long. Glumes gradually acuminate. Central awn recurved, 30 — 50 mm long. Anthers c. 3 mm long. Lower glumes 7.5—10 mm long. — Buru, New Guinea (W Prov.) ……….………… 12. A. superpendens
8. a. Basal internodes glabrous. Blades not coiled. Lower glumes shorter than the upper ones. Lemma margins overlapping, lateral awns well-developed, 8—26 mm long. (Sect.
Macrocladae)……..………. 9 b. Basal internodes appressed pilose. Blades
becoming coiled with age. Lower glumes longer than the upper glumes. Lemma margins involute, forming a longitudinal furrow, lateral awns much reduced, 0—4 mm long. Culms 0.6—1 m long. Blades 7—21.5 cm long.
Panicle loosely contracted, few- spikeled, 15— 28 by 1—1.5 cm. Lower glume 6.25—10 mm long. Callus c. 0.5 mm long, column 4—6.25 mm long, central awn straight. Anthers c. 1 mm long. — New Guinea (C Prov.) (Sect.
Streptachne)…….…… 13. A. utilis var. utilis
9. a. Lemma column 3.5—21 mm long……..…. 10 b. Lemma column 0.5—3 mm long. Culms 0.4–0.7 m long. Blades 5.5—15 cm long. Panicle few-spikeled. Lower glume 5—8.2 mm long; upper glume 7—11 mm long. Central awn patent to recurved, 22—30 mm long, lateral awns 8.7–22 mm long. Anthers c. 1.5 mm long. – New Guinea (Manokwari, C Prov.) 8. A. novaeguineae 10. a. Panicle loosely contracted to lax, 10—60 by
2—8 cm…….………..… 11
b. Panicle contracted, 4–9 by c. 1 cm. Culms 0.35–0.4 m long. Blades 9–17 cm long. Panicle few-spikeled. Lower glume 5.75–10 mm long; upper glume 8.75—13 mm long. Callus 1.25— 1.5 mm long, column 7.5–8.5 mm long, central awn strongly recurved, 17–18 mm long, lateral awns 9–14 mm long. – New Guinea (Milne
Bay) 14. A. warburgii
11. a. Blades 7–80 cm long. Panicle 15–60 cm long. Lateral awns 8–27 mm long…….… 12 b. Blades 7—12.5 cm long. Panicle 10—13 cm
long. Lateral awns 28--30 mm long. Culms 0.25—0.45 m long. Panicle few-spikeled. Lower glume 6—7 mm long; upper glume 10—12 mm long. Callus 1—1.5 mm long, column 5.5—8 mm long, central awn straight, c. 30 mm long. Anthers c. 1.25 mm long. — New Guinea (W Prov.) 9. A. papuana 12. a. Callus 0.5—0.75 mm long, column 3.5—8 mm
long, central awn straight. Anthers 2—4 mm long……..…………...……...…………... 13 b. Callus 1—1.5 mm long, column 12—21 mm
long, central awn recurved. Anthers 1.25—1.5 mm long. Culms 0.5—1.2 m tall. Blades 15— 30 cm long. Panicle few-spikeled. Lower glume 6—9 mm long; upper glume 11—15 mm long. Central awn 25—32 mm long, lateral awns 15—27 mm long. —New Guinea (Merauke W Prov.)...………... 7. A. meraukensis
13. a. Culms 1—1.75 m tall. Blades 35—80 cm long. Panicle many-spikeled, 30—60 cm long. Lower glume 7.5—12 mm long; upper glume 9—13 mm long. Central awn 22—33 mm long, lateral awns 20--26 mm long. Anthers c. 4 mm long. — Malaya (Perlis), Philippines (Busuanga, Culion)……….………. 3. A. culionensis
b. Culms 0.35—1 m tall. Blades 7–21 cm long. Panicle few-spikeled, 13–24 cm long. Lower glume 2.8—5.7 mm long; upper glume 5.5–8.8 mm long. Central awn 8—18.5 mm long, lateral awns 8—15 mm long. Anthers 1.5–2 mm long. –– New Guinea
2004] VELDKAMP: Misc. notes on mainly Southeast Asian Gramineae 137
The species of Aristida spp. in Malesia often have curious, unexplainable distributions, which are enumerated below.
1. ARISTIDA ADSCENSIONIS L.
DISTRIBUTION. (Sub)tropics in the New and Old World, not in Australia; nearest in Asia: SW Thailand (Kanchanaburi); Malesia: Lesser Sunda Islands (Sawu, Sumba, Timor). In view of the disjunct distribution most likely introduced, but compare a similar one for Acrachne racemosa
(Heyne ex Roem. & Schult.) Ohwi,
Desmostachya bipinnata (L.) Stapf, and
Elytrophorus spicatus (Willd.) A. Camus.
2. ARISTIDA CHINENSIS Munro
DISTRIBUTION. Thailand to S China (Guang-dong, Hong Kong), Taiwan. Malesia: Celebes (Palu), Philippines (Mindanao: Cotabato).
3. ARISTIDA CULIONENSIS Pilg.
DISTRIBUTION. S Thailand (Phuket, Satun, Songkla) to S Vietnam. Malesia: Malaya (Perlis), Philippines (Busuanga, Culion). Note the disjunction here.
4. ARISTIDA CUMINGIANA Trin. & Rupr.
DISTRIBUTION. Tropical Africa to China (Jiangsu) (not in Pakistan) and N Australia (Queensland). Malesia: Celebes (Central, Towuti Lake), Philippines (Busuanga; Culion I.; Luzon: Benguet, Bontoc, Ilocos Norte, Lepanto, Nueva Ecija, Nueva Vizcaya, Rizal, Zambales; Mindanao: Bukidnon), New Guinea (Irian Jaya: Kebar, Baliem, Swart Valley; Papua New Guinea: W Highlands, Morobe Prov.).
NOTE. Occasionally the lateral awns are lacking, e.g. in the Philippines in BS 41306 (Ramos) (P, SING; 3-awned in BRI) from Culion I., BS 26092 (Fénix) (BRI, L, MO, SING, US) from Tanculan, Mindanao, and Co s.n. (L) with unknown provenance. A collection by the indefatigable J.F. Maxwell (CMU) was made Central - Thailand (Maxwell 01-397, CMU, L, from Nakorn Nayok Prov.) as well.
Such forms with longer glumes and simple awns have been distinguished in Africa as A. diminuta (Mez) C.E. Hubb. Its occurrence in Thailand and the Philippines suggests that it cannot be upheld at any rank (Veldkamp, 1992: 227).
5. ARISTIDA HOLATHERA Domin VAR. HOLATHERA
DISTRIBUTION. Australia (W Australia to Queensland), Malesia: Philippines, Luzon (Bataan, Ilocos Norte, Ilocos Sur, La Union, Pampanga, Zambales).
6. Aristida macroclada Henrard var. queens-landica (B.K. Simon) Veldk., stat. nov.
Aristida macroclada Henrard subsp. queenslandica B.K. Simon, Austrobaileya II (1984): 95, t. 3b. — Type: S.T. Blake 21741 (BRI, sh. 65923; CANB, MEL, NSW).
DISTRIBUTION. Australia (NE Queensland), Malesia: New Guinea (Irian Jaya: Merauke; Papua New Guinea: W Prov.).
NOTES. Simon (1984, 1992) regarded this taxon as distinct from the typical one because of its well-developed column (5—8 mm long i.s.) with several spirals, against a short column (1—2 mm long i.s.) with half to one spiral in the latter, and a subspecies because these taxa appear to be allopatric.
Indeed the length of the column is distinctive, and the only difference found in this survey.
The typical variety has been found in the Northern Territory and NW Queensland; the citation for New Guinea refers to A. novae-guineae (see below). The var. queenslandica
occurs in Australia in NE Queensland and in New Guinea in the adjacent areas of Merauke and the Western Province.
However, to regard allopatry as an argument to confer automatically the status of subspecies is
an a priori reasoning often applied, but
inapplicable to taxonomy. The difference in the length of the column could well be due to a single mutation, in which case at most the rank of forma should be given. Populations ought to be checked whether the forms do not occur together. The apparent allopatry both in Australia and New Guinea may also be due to the fact that in Malesia all species of Aristida tend to be rare and widely scattered, as will be evident from the present account, while the areas where A. macroclada s.l. has been found in Australia are badly under-explored.
7. ARISTIDA MERAUKENSIS Henrard
138 REINWARDTIA [VOL.12
8. ARISTIDA NOVAEGUINEAE Ohwi
DISTRIBUTION. New Guinea: Irian Jaya (Manokwari), Papua New Guinea (C Prov., Sogeri Plateau).
NOTE. Blake (1969: 3) and subsequent authors have reduced this to A. macroclada Henrard from N Australia, but specimens of that seen differ by:
1 a. Blades 15—28.5 cm by 2—3 mm. Lower glumes 3—4.5 mm long. Awns subequal, central awn straight, 8.5—17 mm long A. macroclada
b. Blades 5.5—15 cm by 0.5—1.5 mm. Lower glumes 5—8.2 mm long. Awns unequal, central awn patent to recurved, 22—30 mm long
A. novaeguineae
Therefore A. novaeguineae is re-instated as a species.
9. ARISTIDA PAPUANA Veldk.
DISTRIBUTION. Papua New Guinea, W Prov. (Morehead).
10. ARISTIDA POLYCLADOS Domin
DISTRIBUTION. Australia (W Australia to Queensland). Malesia: Sumba (Kendara, Payeti, Waingapu).
11. ARISTIDA SETACEA Retz.
DISTRIBUTION. Mascarenes, Réunion, Rodri-guez, India, Sri Lanka, Burma, Thailand (Satun). May be expected in N Peninsular Malaysia.
12. ARISTIDA SUPERPENDENS Domin
DISTRIBUTION. Australia (N Territory, Queensland). Malesia: Buru (Namlea), Papua New Guinea, Western Prov. (Weam). Note the disjunct distribution in Malesia.
13. ARISTIDA UTILIS F.M. Bailey VAR. UTILIS
DISTRIBUTION. Australia (Northern Territory, Queensland). Malesia: Papua New Guinea (N of Port Moresby, Central Prov.).
14. ARISTIDA WARBURGII Mez
DISTRIBUTION. Australia (Queensland, New
South Wales), Malesia: Papua New Guinea, Milne Bay Prov. (Sudest, Rossel Isl.). This is a curious disjunction and the species can therefore be expected on the South side of Papua New Guinea as well.
Kengia nedoluzhkoi (Tzvelev) Veldk., comb. nov.
Cleistogenes nedoluzhkoi Tzvelev, Bot. Zur. (Moscow & Leningrad) LXXXVII (7) (2002): 115. — Type: Tzvelev N. 224 (LE, holo).
DISTRIBUTION — Russia, Primorskje Prov., Partizanskaja River.
NOTE. The generic name Cleistogenes Keng is invalid under Art. 20.2 ('technical term') and has been replaced by Kengia Packer.
Moorochloa Veldk., gen. nov. for Brachiaria Auct.
As I have discussed elsewhere (Veldkamp, 1996) the genus Brachiaria (Trin.) Griseb. is to be nearly completely reduced to Urochloa P. Beauv. This idea originated with T.-Q. Nguyen (1966), was extended by Webster (1987, 1988, 1992), extensively argued by Morrone & Zuloaga (1992, 1993), and followed by subsequent authors, e.g. Terrell & Reveal (1996).
The remnant consists of 3 closely related species. The most wide-spread one is B. eruciformis (Sm.) Griseb. (incl. Panicum isachne
Roth, P. caucasicum Trin.), which occurs from S Africa to the Mediterranean, S Russia, and through the Near East to E India, and is introduced widely elsewhere, e.g. in Malesia: Java (Surakarta, Semarang, Pasuruan), Madura, Papua New Guinea (Central Prov.).
Brachiaria malacodes (Mez & K. Schum.) H. Scholz occurs in Namibia to Angola, and B. schoenfelderi C.E. Hubb. & Schweick. has been found in Namibia and Zimbabwe.
The most widely distributed species, B. eruciformis, generally is cited as the type of
Brachiaria (Trin.) Griseb. However, it was not included under any name in Trinius' original (1826) concept of the generic basionym Panicum
L. sect. Brachiaria Trin., and so cannot be its type.
2004] VELDKAMP: Misc. notes on mainly Southeast Asian Gramineae 139
Urochloa. Hereby the well-known name
Brachiaria has become a synonym of the latter. Because of stability in nomenclature it would have been preferable to maintain the name
Brachiaria for at least this small group. This could have been achieved easily by appointing B.
eruciformis as the conserved type and so
legalizing the traditional use.
However, this proposal was rejected by the Nomenclatural Committee for Spermatophyta (Brummitt, 1998) with the suggestion that a new name (actually a new genus) should be published. With great reluctance and after long hesitation I have finally followed this advice, and hereby dedicate this 'new' genus to that august body.
Moorochloa ab Urochloa in spiculis supra glumas disarticulatis, callo inconspicuo, lemmate superiore chartaceo ad cartilagineo lucido laevi mutico differt. — Type: Moorochloa eruciformis
(Sm.) Veldk.
1. a. Spikelets disarticulating above the glumes, callus inconspicuous. Upper lemma chartaceous to cartilaginous, shiny, smooth, muticous
Moorochloa
b. Spikelet disarticulating below the glumes, callus distinct. Upper lemma indurated, dull, coarsely to finely transversely rugose, apiculate to
mucronate Urochloa
1. Moorochloa eruciformis (Sm.) Veldk., comb. nov.
Panicum eruciforme Sm. in Sibth. & Sm., Flora Graeca I (1808): 44, t. 59. — Brachiaria eruciformis
Griseb. in Ledeb., Fl. Ross. IV (1853): 469 ('erucaeformis'). — Panicum cruciforme Sibth. ex Roem. & Schult., Syst. Veg. II (1817): 426 (sphalm.). — Echinochloa eruciformis Rchb., Fl. Germ. Excurs. III (1833): 45. — Milium alternans Bubani in Willk. & Lange, Nuov. Giorn. Bot. Ital. V (1873): 317, nom. superfl. — Type: Sibthorp s.n. (OXF, holo, L, photocopy, LP, photo).
2. Moorochloa malacodes (Mez & K. Schum.) Veldk., comb. nov.
Panicum malacodes Mez & K. Schum. in Mez, Notizbl. Bot. Gart. Berlin-Dahlem VII (1917): 70. —
Brachiaria malacodes H. Scholz, Willdenowia VIII (1978): 384. — Type: Antunes 202 (B, holo).
3. Moorochloa schoenfelderi (C.E. Hubb. & Schweick.) Veldk., comb. nov.
Brachiaria schoenfelderi C.E. Hubb. & Schweick. in Schweick. & C.E. Hubb., Bull. Misc. Inform. (1936): 323. — Type: Schoenfelder 584 (PRE, holo).
I take the opportunity to transfer a SE Asian species to Urochloa:
A NEW COMBINATION IN UROCHLOA
Urochloa burmanica (Bor) Veldk., comb. nov. Brachiaria burmanica Bor, Kew Bull. (V) (1950) :232. — Type: U Thein Lwin 526 (K, holo).
DISTRIBUTION. Myanmar, Yangon (Rangoon) Distr., Pyay [Prome, subdiv. of Bago (Pegu) Distr.].
Rytidosperma bonthainicum (Jansen) Veldk.,
comb. nov.
Danthonia pilosa R. Br. var. bonthainica Jansen, Reinwardtia II (1953): 258. — Notodanthonia penicillata (Labill.) P. Beauv. subsp. bonthainica
Veldk., Taxon XXIX (1980): 298. — Danthonia bonthainica Veldk., Blumea XXXVIII (1993): 217. —
Notodanthonia bonthainica H.P. Linder, Telopea VI (1996): 615. — Austrodanthonia bonthainica H.P. Linder, Telopea VII (1997): 270. — Type:
Bünnemeijer11971 (BO, holo; L).
DISTRIBUTION. Celebes (Bonthain).
HABITAT. In subalpine coppices, disturbed places, 2500—2890 m alt.
NOTE. Ms. M. Gies (in litt., June 14, 1992) reported the presence of axillary cleistogamous spikelets in Bünnemeijer 12260 (L, SING). I have seen the sheets, but not such spikelets.
REFERENCES
BLAKE, S. T. 1969. Taxonomic and nomenclatural
studies in the Gramineae, no. 2. Proc. Roy. Soc. Queensland LXXXI: 3.
BRUMMITT, R. K. 1998. Report of the Committee for
Spermatophyta: 47. Taxon XLVII: 869—870. MORRONE, O. & ZULOAGA, F. O. 1992. Revision de
las especies sudamericanas nativas e introducidas de los generos Brachiaria y Urochloa (Poaceae: Panicoideae: Paniceae). Darwiniana XXXI: 43—109.
MORRONE, O. & ZULOAGA, F. O. 1993. Sinopsis del
140 REINWARDTIA [VOL.12
Paniceae) para Mexico y America Central.
Darwiniana XXXII: 59—75.
NGUYEN, T.-Q. 1966. Notulae criticae de gramineis
florae vietnamensis, 1. Novosti Sist. Vyssh. Rast. III: 10—14.
PFEIFFER, L. K. G. 1872—1873. Nomenclator
botanicus: 453. T. Fischer, Kassel.
SIMON, B. K. 1984. New taxa and nomenclatural
changes in Aristida L. (Poaceae) in Australia.
Austrobaileya II: 95, t. 3b.
SIMON, B. K. 1992. A revision of the genus Aristida
(Poaceae) in Australia. Austral. Syst. Bot. V: 191, pl. III, 36, 37, t. 10, c, d.
TERRELL, E. E. & REVEAL, J. L. 1996. [Urochloa
plantaginea, U. platyphylla, U. texana first records in Maryland]. Castanea LXI: 95—96. TRINIUS, C. B. 1826. De graminibus paniceis.
Dissertatio botanica altera: 51, 125, 266. Acad. Imp. Sci., St. Petersburg.
VELDKAMP, J. F. 1992. Miscellaneous notes on SE Asian
Gramineae. VII. Blumea XXXVII: 227—237. VELDKAMP, J. F. 1996. Proposal to conserve the name
Brachiaria (Trin.) Griseb. (Gramineae) with a conserved type. Taxon XLV: 319.
WEBSTER, R. D. 1987. The Australian Paniceae
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ISSN 0034-365 X
REINWARDTIA
Vol. 12. No. 2. 2004
CONTENTS
Page
W.J.J.O. DE WILDE & BRIGITTA E.E. DUYFJES. Kedrostis Medik. (Cucurbitaceae)
in Asia .'. - 129
J.F. VELDKAMP. Miscellaneous notes on mainly Southeast Asian Gramineae... 135
PITRA AKHRIADI, HERNAWATI AND RUSJDITAMIN. A new species of Nepenthes
(Nepenthaceae) from Sumatra - 141
KUSWATA KARTAWINATA, ISMAYADI SAMSOEDIN, M. HERIYANTO AND J.J. AFRIASTINI. A tree species inventory in a one-hectare plot at the Batang Gadis National
Park, North Sumatra, Indonesia .. 145
E.A.P. ISKANDAR & J.F. VELDKAMP. A revision of Malesian Isachne sect. Isachne
(Gramineae, Panicoideae, Is.ach.neae) ' 159
JOHANIS P. MOGEA. Four new species pf Arenga (Palmae) from Indonesia 181
J.F. VELDKAMP. The correct name for Pyrrosia hastata Ching (Polypodiaceae,
Pteridophyta) ... 191
TRI MULYANINGSIH & COLIN ERNEST RIDSDALE. An additional species of
Villaria Rolfe {Rubiaceae') from The Philippines 195
ELIZABETH A. WIDJAJA, INGGIT PUDJI ASTUTI & IDA BAGUS KETUT ARINASA. New species of bamboos (Poaceae-Bambusoideae) from Bali 199
HERBARIUM BOGORIENSE BIDANG BOTANI
