ATOLL RESEARCH BULLETIN

No. 220 September 1978

ATOLL RESEARCH BULLETIN

220. Coval Reef Ecosystetns:

Pvoceedings of Papers Presented at the 13th Pacific Science Cotzgvcss, Vanc011 ver Edired by S. VI Smith

Issued by

THE SMITHSONIAN INSTITUTION

Washington, D.C., U.S.A.

ATOLL

RESEARCH BULLETIN

220. Coral Reef Ecnsystetrts:

Proceedings of Papers Pvesentecl at the 13th Pacific Sciertce Cortgress, Varrcol~ver Edited ^{~ I} S. V Smith ^I

Issued

by

THE SMITHSONIAN INSTITUTION Washington, D.C., U.S.A.

September 1978

ACKNOWLEDGMENT

The A t o l l Research B u l l e t i n i s i s s u e d by t h e Smithsonian

I n s t i t u t i o n a s a p a r t o f i t s T r o p i c a l Biology Program. It i s co-sponsored by t h e Museum o f N a t u r a l H i s t o r y and t h e Smithsonian P r e s s . The P r e s s s u p p o r t s and h a n d l e s p r o d u c t i o n and d i s t r i b u t i o n . The e d i t i n g i s done by t h e T r o p i c a l Biology s t a f f , Botany Department, Museum o f N a t u r a l H i s t o r y and by D . R . S t o d d a r t .

The B u l l e t i n was founded and t h e f i r s t 117 numbers i s s u e d by t h e P a c i f i c S c i e n c e Board, N a t i o n a l A c a d e ~ o f S c i e n c e s , w i t h f i n a n c i a l

s u p p o r t from t h e O f f i c e o f Naval Research. I t s pages were l a r g e l y devoted t o r e p o r t s r e s u l t i n g from t h e P a c i f i c S c i e n c e Board's C o r a l A t o l l Program.

The s o l e r e s p o n s i b i l i t y f o r a l l s t a t e m e n t s made by a u t h o r s o f p a p e r s i n t h e A t o l l Research B u l l e t i n r e s t s w i t h them, and s t a t e m e n t s made i n t h e B u l l e t i n do n o t n e c e s s a r i l y r e p r e s e n t t h e views o f t h e Smithsonian n o r t h o s e o f t h e e d i t o r s o f t h e B u l l e t i n .

E d i t o r s

F . R . Fosberg M.-H. S a c h e t

Smithsonian I n s t i t u t i o n Washington, D . C . 20560 D . R . S t o d d a r t

Department o f Geography U n i v e r s i t y o f Cambridge Downing P l a c e

Cambridge, England

TABLE OF CONTENTS

Page

Biogeochemical Budgets i n C o r a l Reef Systems

S . V . S m i t h , P. L. J o k i e l , a n d G. S. Key

. . .

¹

S p a t i a l M o d e l l i n g of C o r a l R e e f s

A. L . Dahl

. . .

^{1 3}

An E v a l u a t i o n o f C o r a l Reef Survey Methods by Computer S i m u l a t i o n

R . A. K i n z i e 111

. . .

^{2 1}

S c l e r o c h r o n o l o g y : A Data S o u r c e f o r Reef Systems Models

R . W . B u d d e m e i e r . . .

. . .

^{2 5}

Energy Exchange i n C o r a l Reef Ecosystems

A. A. Benson, J. S. P a t t o n , a n d S. Abraham

. . .

³³

D i f f e r e n t i a t i o n i n C o r a l P o p u l a t i o n s

D . C . P o t t s

. . .

55 E v i d e n c e from t h e G r e a t B a r r i e r Reef of A n c i e n t A c a n t h a s t e r

A g g r e g a t i o n s

E . F r a n k e l

. . .

⁷⁵

An I n f o r m a t i o n A n a l y s i s Approach t o Z o n a t i o n P a t t e r n s of t h e C o r a l Genus A c r o p o r a on O u t e r Reef B u t t r e s s e s

. . .

Carden W a l l a c e and M . B . Dale 95

EDITOR'S NOTE:

A s e s s i o n e n t i t l e d "Systems M o d e l l i n g and C o r a l Reef Ecosystems"

w a s h e l d a t t h e 1 3 t h P a c i f i c S c i e n c e Congress (18-30 August 1975, Vancouver, Canada). The p a p e r s i n t h a t s e s s i o n d e a l t w i t h c o n c e p t u a l a s p e c t s of r e e f s y s t e m s as w e l l as w i t h s p e c i f i c c h e m i c a l o r b i o t i c components of t h o s e s y s t e m s . It w a s d e c i d e d d u r i n g t h a t s e s s i o n t o c o l l e c t as many of t h o s e p a p e r s as p o s s i b l e i n t o a s i n g l e number o f t h e A t o l l R e s e a r c h B u l l e t i n .

T h i s number of ARB r e p r e s e n t s t h e r e s u l t of t h a t d e c i s i o n . E i g h t of t h e o r i g i n a l t e n p a p e r s a r e p r e s e n t e d h e r e i n t h e i r e n t i r e t y , i n somewhat a l t e r e d f o r m a t , o r as expanded a b s t r a c t s . The s e q u e n c e of p a p e r s i n t h i s volume i s t h e same a s t h e o r d e r i n which t h e y were p r e s e n t e d a t Vancouver. I t i s a n t i c i p a t e d t h a t t h e o t h e r two p a p e r s w i l l b e p u b l i s h e d e l s e w h e r e .

I t h a n k a l l of t h e s p e a k e r s a t t h e symposium s e s s i o n f o r t h e i r p a r t i c i p a t i o n , and t h o s e who p r o v i d e d m a n u s c r i p t s f o r making t h i s p u b l i c a t i o n p o s s i b l e . To t h e e d i t o r s of t h e A t o l l R e s e a r c h B u l l e t i n go my t h a n k s f o r e n c o u r a g i n g t h e p u b l i c a t i o n of t h e s e p a p e r s i n t h e B u l l e t i n . E d i t i n g and t y p i n g of t h e m a n u s c r i p t s w a s u n d e r t a k e n t h r o u g h t h e a u s p i c e s of t h e Hawaii I n s t i t u t e of Marine B i o l o g y , U n i v e r s i t y of Hawaii.