11
U563
CRLSSI
SMITHSONIAN INSTITUTION UNITED STATES NATIONAL MUSEUM
Bulletin
199AN ANNOTATED CHECKLIST AND KEY TO THE REPTILES OF MEXICO
EXCLUSIVE OF THE SNAKES
BY
HOBART M. SMITH and EDWARD H. TAYLOR
UNITED STATES
GOVERNMENT
PRINTING OFFICEWASHINGTON :1950
ForsalebytheSuperintendentofDocuments, U.S.GovernmentPrinting Office Washington25,D. C. - Price75cents(Paper)
ADVERTISEMENT
The
scientific publications of theNational Museum
includetwo
series,
known,
respectively, as Proceedingsand
Bulletin.The
Proceedings series,begun
in 1878, isintended
primarily as amedium
forthe publicationof original papers,based on
thecollections oftheNational Museum,
thatsetforthnewly
acquiredfacts inbiology, anthropology,and
geology,with
descriptions ofnew forms and
re- visions of limited groups.Copies
ofeach
paper, inpamphlet
form, are distributed as published to librariesand
scientific organizationsand
to specialistsand
others interestedinthedifferentsubjects.The
dates at
which
theseseparatepapers
arepublished arerecorded in the table ofcontents ofeach
ofthevolumes.
The
series of Bulletins, the first ofwhich was
issued in 1875, con- tainsseparate publicationscomprising monographs
of large zoologicalgroups and
other general systematic treatises (occasionally in several volumes), faunalworks,
reports of expeditions, catalogs oftype
specimens, special collections,and
other material of similar nature.The majority
of thevolumes
areoctavo
insize,but a quarto
sizehas been adopted
ina few
instances inwhich
large plateswere regarded
as indispensable.In
the Bulletin seriesappear volumes under
theheading
Contributionsfrom
the United StatesNational Herbarium,
inoctavo
form, publishedby
theNational Museum
since 1902,which
containpapers
relating to the botanical collections of theMuseum.
The
presentwork forms No.
199 ofthe Bulletin series.Alexander Wetmore,
Secretary,
Smithsonian
Institution.II
CONTENTS
Page Introduction.
Itinerary and gazetteer of localities for the Walter Rathbone Bacon
Expedition, 1938-1941 4
Class Reptilia
^^
SubclassAnapsida
Order Testudines ^2
Suborder Atheca 1^
Family Dermochelyidae 13
GenusDermochelys 13
Suborder Carettoidea. 14
FamilyCheloniidae 1'^
Genus Lepidochelys 14^
Genus Caretta IS
GenusEretmochelys 16
Genus Chelonia 1"^
Suborder Trionychoidea 1^
Family Trionycbidae 18
Genus
Amyda
18Suborder Cryptodira 18
Family Dermatemyidae 19
Genus Dermatemys 19
Family Chelydridae 20
Genus Chelydra 20
Family Kinosternidae 21
GenusKinosternon 21
GenusClaudius 26
GenusStaurotypus 27
Family Testudinidae 27
GenusGopherus 28
Family
Emyidae
28Genus Geoemyda 29
Genus Pseudemys 31
Genus Chrysemys 33
Genus Terrapene 34
Genus Malaclemys 36
Genus Clemmys 37
SubclassLepidosauria 37
OrderSquamata.. ^'
SuborderAmphisbaenia 37
Family Bipedidae 37
GenusBipes 38
rn
Class Reptilia
—
ContinuedSubclass Lepidosauria
—
ContinuedOrder
Squamata —
Continued PageSuborderSauria 39
Family Gekkonidae 40
Genus Coleonyx 41
GenusGonatodes 45
Genus Phyllodadylus 46
Genus Thecadadylus 49
Genus Hemidadylus 49
GenusAristelliger 51
GenusPeropus 51
GenusSphaerodadylus 52
Family Iguanidae 53
GenusAnolis 55
Genus Corythophanes 68
Genus
Laemandus
69Genus Basiliscus 71
Genus Iguana 72
Genus Ctenosaura 73
Genus Enyaliosaurus 75
Genus Dipsosaurus 77
Genus Sauromalus 78
GenusHolbrookia 81
GenusCallisaurus 85
Genus
Uma
89Genus Petrosaurus 90
Genus Streptosaurus 91
GenusCrotaphytus 91
Genus Gambelia 93
Genus Phrynosoma 94
GenusSceloporus 105
GenusSator 139
Genus Urosaurus 140
Genus Uta 147
FamilyXantusiidae 151
Genus Lepidophyma 151
Genus Gaigeia 153
Genus Xantusia 154
Family Scincidae 155
Genus
Mabuya
155Genus Scincella 156
Genus Eumeces 160
FamilyAnelytropsidae 170
Genus Anelytropsis 170
FamilyTeiidae 170
Genus Ameiva 170
Genus Cnemidophorus 174
Genus Gymnophthabnus 192
Family Helodermidae 192
GenusHeloderma 192
CONTENTS V
Class Reptilia
—
ContinuedSubclass Lepidosauria
—
ContinuedOrder
Squamata —
ContinuedSuborder Sauria
—
Continued PageFamily Anguidae 194
Genus Celestus 194
GenusAbronia 196
Genus Barisia 198
Genus Gerrhonotus 203
Genus Elgaria 205
Family Xenosauridae 207
Genus Xenosaurus 207
Family Anniellidae 208
Genus Anniella 209
SuborderSerpentes.'
Subclass Archosauria 209
Order Loricata 209
Family Crocodylidae 210
Genus Crocodylus 210
Family Alligatoridae 211
Genus Caiman 212
Species inquirendae 212
Statelists 215
Index 231
'SeeU.S.Nat.Mus.BuU.187, 1945, p. 12.
AN ANNOTATED CHECKLIST AND KEY TO THE REPTILES OF MEXICO, EXCLUSIVE OF THE SNAKES
By HoBART M.
Smith andEdward
H.Taylor
INTRODUCTION
This volume
constitutes the last of a series of checklistsand keys
to the herpetologicalfauna
ofMexico. The
first, ''AnAnnotated
Checklistand Key
to theSnakes
ofMexico"
(U. S.Nat. Mus.
Bull.187),
was
publishedon October
5,1945.The
second,"An Annotated
Checklistand Key
to theAmphibia
ofMexico"
(U. S.Nat. Mus.
Bull. 194),
appeared on June
17, 1948.In
this, the thirdvolume,
the turtles,amphisbaenians,
lizards,and
crocodilians are treated.The preparatory work
thatpreceded
these checklistswas pursued
astime permitted from
1932 to the present, a period ofabout
17 years. Ithas
involveda number
ofexpeditions toMexico
atour own expense
^and
thecollecting ofmore than
50,000 herpetological specimens.Except
in rare instances the materialshave
all passedunder
the scrutiny ofone
or the other of us,and
the detailed results of these studies areembodied
inmore than
ahundred
short or longerreports.Our own
ideashave
oftenchanged with
the acquisition of materialsnot
originally available.It
was hoped
that the long-awaitedwork on
turtlesby
Stejnegerand Hartweg
w^ould see the light before theappearance
of the turtle checklist,but
since there is stillno
certainty of thecompletion
of thiswork
in theimmediate
future it doesnot seem
wise to delayon
this account.We have
refrainedfrom
anticipating certainforms not now
recog- nized thatarepresenteven
inour own
collection; thespecimens
repre- sentingthem
are referredwithout comment
to the recognizedform they most
closelyresemble.A number
ofsuch forms belong
toAnolis,no adequate
revision ofwhich has
yetbeen made, although
theseniorauthor
iscontemplating such
awork.
Certain other groups, especiallyCnemidophorus and
Uta, meritreinvestigation.There
aremany
otherunsolved problems
inMexican
herpetology;much more
explorationremains
tobe
performed.Probably no
single'From1938 to 1941 1wasfinancedbytheSmithsonianInstitutiononseveral expeditions,through tenure ofaWalterRathboneBaconTravelingScholarship.—H.M.S.
1
BULLETIN"
199,UNITED STATES NATIONAL MUSEUM
problem
of distribution ismore demanding
of explanationthan
theextreme dearth
ofPlethodontidae
intheGuerrero
"island"and
in thehigh western
plateau,and
theapparent replacement
of that familyin the latter region, largelyby
theAmbystomidae.
The treatment
offorms
inthisvolume
differslittlefrom
that ofthetwo
precedingvolumes,
except that thename
of the collector ofeach type
isadded.This name
often is asimportant perhaps
as thename
ofthe
author
of aspecies. Unfortunately, the collector is frequentlyunknown
or uncertain.This
is true of the collectionsfrom Mexico
studiedby Wiegmann,
inwhich
casewe have given Deppe
credit for obtainingmost
of the reptiles.The
othertwo men
constituting the collecting party,Count Von Sack and
Schiede, were,we
believe,engaged
primarily in the collection of orchids.However,
it isnot
at allimprobable
thatsome forms
accredited toDeppe were
actuallytaken by one
of the others.We
refrain herefrom
considering the implications thatmay be deduced from
a perusal of thecombined data
of the threevolumes, such
asan
investigation of faunal origins, lines ofmigration, relative age oftaxonomic
groups,and
faunal areas of ecological relationships.However, we
are fullyaware
thatsuch
considerationwould
doubtless yield valuable information.The number
offorms
recognized in theherpetofauna
ofMexico,
including thoseforms
described since the fii'st of these checldistswas
published, is relatively large.The
following table represents thenumbers
in the various groups:Table
1.—
Number
offorms recognizedCHECKLIST OF REPTILES OF MEXICO 3 with
it,almost
inevitably, occasional overzealous reduction ofspecies to subspecies.Such changes
aremade mthout due demand
forfacts,but
insteadwith
speculations that oftenmerely
serve asan
excuse totamper with an
acceptedarrangement
thatmay be
equallyas plausible.One
ofthemost
exasperating taskshas been
the unravelingofcertain purely nomenclatorial knots. Severalnames,
forexample,
are of controversialorthography. Worthy
of specialmention
are Agkistro-don
vs. Ancistrodon,Cnemidophorus
tessellatus vs. C. tesselatus, Cory- tophanes vs. Corythophanes,and Kinosternon
vs.Cinosternon.Opinion
3G of the InternationalCommission on
ZoologicalNomenclature
interprets the permission
given by
Article 19 to correctany
"error of transcription" tomean
also "error of transliteration,"whether
the originalauthor was aware
of hiserroror not.By such
procedure, thenames
Ancistrodon, tessellatus, Corythophanes,and
Cinosternonwould be recommended. On
the otherhand, Moore,
Weller,and Knight
(Journ. Paleont., vol. 16, 1942, pp.250-261) maintain with
excellentreason
thatonly
invery
clear-cutand
exceptional cases isany
modification of the originalorthography
of a genericname
justified.
Their view
isstrengthened by
Blackwelder,Knight, and Sabrosky
(Science,vol. 108, 1948, pp. 37-38).Maintenance
of originalform would perpetuate
Agkistrodon, tesselatus, Kinosternon,and
Corytophanes. Since theproper procedure
isnot
universally oreven
generallyagreed upon, we
see littleadvantage
in deviationfrom
currentlyaccepted form untU some means
of real standardization isprovided by
the InternationalCommission on
ZoologicalNomen-
clature.
For
the firsttime
in these checklistswe
hereadopt
the practice of placing acomma between
scientificname and author
in citations ofall references except theoriginal.We wish
toacknowledge
the assistance ofW.
LeslieBurger
in the laborious task ofchecking and
rechecking various points in the manuscript,and
in criticizing certain parts,and
ofDr. Doris M.
Cochran,
zoologist, division of reptilesand amphibians. United
StatesNational Museum, who devoted much time
to curatorial dutiesconnected with
thevoluminous
collectionand
inmaking
available forstudy
theNational Museum
herpetological collections.We
are grate- ful likewise for the correctionsand improvements
suggestedby Dr.
D.
F. Hoffmeister,Karl
P.Schmidt, and Dr.
FrederickShannon. The
Graduate Research Boards
of the University of Illinoisand
the University ofKansas have
generouslyprovided
financialsupport
for researchand
clericalwork
involved incompletion
of the present project,and
theWalter Rathbone Bacon
Scholarship of theSmith-
sonianInstitution furnished a substantialsum
for traveland
research inMexico.
4
ITINERARY AND GAZETTEER OF LOCALITIES FOR THE WALTER RATHBONE BACON EXPEDITION,
1938-1941As
previously stated, the seniorauthor was enabled
to collect inand study
materialfrom Mexico by
tenure of aWalter Rathbone Bacon Traveling
Scholarshipfrom 1938
to 1941.The
collectionobtained during
those yearshas formed an important
addition to other materialfrom Mexico and
meritsan
elaboration of the routes traversedby
the expeditionand
the localities atwhich specimens were
secured.Such an
itinerary was, as amatter
of fact, tohave accompanied
astudy
of the lizards of theBacon
collection, as stated inour
introduction to thesummary
of theamphibians
(Proc.U.
S.Nat. Mus.,
vol. 95, 1945, p. 521). Itnow seems
unlikely that acom-
plete
survey
of theBacon Hzards can appear
for anumber
of years, for a satisfactorycompletion
ofsuch
astudy
involves revision of anumber
ofvery
sketchilyunderstood
genera,such
as Anolisand Cnemidophorus. In
themeantime
aneed
foran account
oftheBacon
itinerary
and
localities continues to exist.We
have, accordingly,taken
thisopportunity
to present thisinformation
in this the final checklist.The
expedition personnel consistedofthe seniorauthor and
his wife.They
receivedhearty support
in fieldwork from
anumber
of indi- viduals,through whose combined
effortsabout
22,000specimens were
securedduring
a period oftwo
years.As
closel}''ascan now be
deter-mined
(the lizardsnot
yethaving been
fullystudied),500
speciesand
subspecies (146amphibians,
160lizards, 1amphisbaenian,
170 snakes,20
turtles, 3 crocodilians)were
secured.Among
thosewho
contributed to the success of the expedition areDr. Alexander Wetmore, who saw
to it that the expeditionwas
prop- erlyplanned and
could functionsmoothly; Dr. Doris M.
Cocliran,who must have
nearlyequaled
the expedition personnel intime
ex-pended
forthecollection,inasmuch
asthe laborious taskof cataloging thespecimens
restedwith
her; Dr.Linton
P. Satterthwaite,who provided
thefacilities forour
stay atPiedrasNegras
inthemidst
of a Httle-known, remote,and
austere area;Mr. and Mrs. Dyfrig McH.
Forbes,
who provided
for nearlytwo
years a base of operations inVeracruz and who were
a constant source ofcompanionship,
inspira- tion,and
material;EiziMatuda, who very
generouslyprovided
facil- ities for atwo-month
stay at his fincaand who
sympathetically aided us inevery
possibleway
tosample
theherpetofauna
of the area asthoroughly
as possible;Thomas MacDougall, who
inTehuantepec
secured for usnumerous specimens and arranged
forour
travel into areas otherwise difficult of access;Ernest
Rateike, ofPalenque,
Chiapas, who
accepted us as a guest in hishome
for amonth: and
aCHECKLIST OF REPTILES OF MEXICO 5
host of local residents, far toonumerous
tomention, who
materiallyaugmented our
collectionsand made our
visitspleasantand
successful.Inasmuch
aswe
traveledchieflyby
car,our
routesarebest followedon
aroad map
ofMexico. We
enteredMexico on October
5, 1938, atCiudad
Juarez,Chihuahua, and
followed theChihuahua-El Paso highway southward
toVado (October
9).We then turned back
toAhumada and
followed apoor
trailthrough
Carrizaland Rancho Nuevo
to Progreso(October
10).We camped
beside theRio Santa Maria about
amilesouthwest
ofProgreso
for 5 days, interruptedonly by one
hurriedround
trip toCasas Grandes
for thebenefit ofa snake- bittencowboy. At
thiscamp we were
aided greatlyby
10 or 15cowboys who brought
in material as fast as it couldbe
preserved.Our specimen
containers full,we determined
togo
to theborder
to shipspecimens and hkewise
to obtain higherwheels
for the car (a half-ton panel truck),with which we had encountered numerous
diffi- cultiesbecause
ofdeep
rutsand high
centers.We
left theProgreso camp on October
15, passingthrough Casas Grandes and
Ascencidn,and reached Las Palomas
thesame
day. Aftertwo days
inColumbus and Deming, N. Mex., we
retracedour
steps(October
20)from Las Palomas
toCasas Grandes. As
theweather had become
cool,we continued southward
along theroad through San Buenaventura and Carmen
to themain El Paso-Chihuahua highway,
thencesouthward
toCiudad Chihuahua. We
left thereon October 27
forTorredn,
arrivingOctober
29.We
collected inthe vicinity ofTorre6n October 30 and
31,and on November
1 started forCiudad Durango. We
could get
no
fartherthan
Pedricena,however,
and, after exploringwith
littlesuccess asideroad
toNazas,
returnedtoTorre6n on Novem-
ber4,
We continued
toSan Pedro
thenext day and
collected in the vicinity ofthattown
untilNovember
10. After a side trip to Parras off the Torredn-Saltillohighway, we made an attempt
to find Jaral, Coahuila, a localitymade famous
zoologicallyby
Hellerand
Barber.This turned
out tobe an abandoned ranch near
Hip61ito, and, asno
likely habitat for the
montane
species recordedfrom
Jaralappeared
tobe
nearerthan
20 miles orso,we continued
to Saltillo(November
14).
We
collected in the vicinity of Saltillo, Arteaga,and Mount
Zapaliname
untilNovember
17,when we
left forMonterrey.
On November
18we
arrived atHacienda La Clementina
(near Forl6n),and we remained
there untilNovember
28,when we drove
toLaredo
to shipanother
lot of specimens.We
returned toLa
Clementina on December
3and
leftDecember
5 forMexico
City.We
stayed atHuichihuayan
for several days, leavingDecember
9.Our
route,with
briefstops at ornear
various towns,then
ledthrough
Mexico City
toTehuacan,
Puebla, thencethrough Orizaba
to Potrero Viejo, Veracruz,where we
arrivedon December
11.6 BULLETIN
199,UNITED STATES NATIONAL MUSEUM
Potrero Viejo
remained our headquarters
forover
amonth. We
made numerous
short tripsduring
that time; especiallynoteworthy were
those toPalma
Sola (on theVeracruz-Orizaba highway),
toTezonapa,
Veracruz,and
Cosolapa,Oaxaca, and
toCuautlapan,
Orizaba,C6rdoba,
Acultzingo,and
otherlocalitiesalong theC6rdoba- Tehuacan
road.We
left Potrero Viejoon January
16, 1939,and
after a brief stop in
Mexico City made headquarters
inCuernavaca, Morelos,
for nearlytwo
weeks,with
side trips toPuente de
Ixtla (Morelos),Cacahuamilpa
(Guerrero),and Zempoala (Morelos and Mexico)
Leaving Cuernavaca February
2,we continued toward Acapulco,
reaching thereFebruary
5.We
collected in the vicinity ofAcapulco, with
side trips toCoyuca,
untilFebruary
11.Our
route,with
fre-quent
stops,then
lednorthward
again toMexico
City,where we
arrived
on March
1. Aftertwo days
in search of axolotlsand
otherambystomids
inthevicinityofMexico City
(withside trips toTexcoco, Zumpango, and Chimalhuacan)
,we
started(March
4)on
theroad
toGuadalajara. Our
first deviationfrom
this routewas on March
9and
10, to Patzcuaro.On March
11we reached Uruapan on another
side tripand continued southward
toApatzingan,
returning toUruapan on March
19.An attempt was made
to find Crotalus foly- stictusinthemarshes
ofthe easternend
ofLake Chapala, but without
success, sincemost
ofthemarshes have been
drainedand
arenow under
cultivation.We reached Guadalajara on March 24 and returned
to Potrero Viejoon March
26.On March 30 we
left Potrero Viejo enroute toLaredo with another shipment
of specimens, arriving atLaredo on
April 3, 1939.After nearly a
month
intheUnited
Stateswe
leftLaredo on
April29, 1939,and
arrived in Potrero Viejoon May
2.From
therewe went
to
Veracruz by
trainand
securedpassage
forAlvaro Obreg6n,
arrivingon May
13.A
riverboat was then
taken, viaCiudad
delCarmen,
to
Tenosique, Tabasco,
thehead
ofnavigationon
theRio Usumacinta.
Our
finaldestination, PiedrasNegras, Guatemala, was reached on May
21 after
two days by mule. We
collected in theimmediate
vicinity ofPiedrasNegras
untilJune
23,when we
left forTenosique.
From Tenosique (June
30)we continued downstream
toEmiliano Zapata and
thereprocured mules
for a2-day
trip toSan
Juanito,a ranch
halfa
milefrom
the village ofPalenque. There we remained, with
a side trip only to the ruins ofPalenque
several miles distant in the hills, untilAugust
6,when we
left forAlvaro Obreg6n
and,immediately
thereafter, PotreroViejo,where we
arrivedon August
14.There we were
joinedby
the juniorauthor and with him
lefton
August
18 forMexico
City,where we
arrived, via the usual routethrough Tehuac§,n and
Puebla,on August
22, afternumerous
briefCHECKLIST OF REPTILES OF MEXICO 7
stopsen
route. After afew days
spentin the environsofMexico City
searching forambystomids, on August
29we took
theAcapulco road and
followed itwith
onlyone
deviation to Tixtla, Guerrero, arriving atAcapulco on September
3,We
returned toMexico City by
thesame
route, arrivingon September
6.Agaui we
collected in theimmediate
environs ofMexico
City, chiefly investigatingmontane
faunas.On September
10we
left forGuadalajara but turned back near Sahuayo. We
then,on September
15, startednorthward on
thePachuca Road
toEl Chico National Park
inHidalgo. Afterone day
therewe
returned,September
18, toMexico
City,where Dr. Taylor
left us.
We then
returned,September
19, to Potrero Viejo. Aftera few
local tripswe
again returned to theGuadalajara
road, leavingSep- tember
29.We took
the sideroad
to theNevado de Toluca (October
2)
and then
retracedour
steps toMexico
City, arrivingOctober
3.After a
few days
spent in local trips,we turned northward
toLaredo with another
load ofspecimens, arrivingatLaredo October
17.Our only
sidetrip enroutewas
toGaleana, Nuevo Le6n,
via theroad from
Linares,Tamaulipas (October
13, 14).After nearly
two months
in theUnited
States,we
again crossed theborder
atLaredo on December
10, 1939,and headed
directly for Potrero Viejo, Veracruz, arrivingDecember
15. After afew
local tripsnear
Potrero Viejo,we
left(December
27) forTehuantepec by
rail.
We
arrivedon December 30 and remained
untilJanuary
28, 1940.A number
of side tripswere
taken,by
truck toCerro
Arenal,by
rail toMatias Romero, Oaxaca, by
rail to Salina Cruz,and by
rail toTonala.
Illness necessitated return to Potrero Viejoand
ulti-mately
toMexico
City.We remained
thereuntilMarch
15,A\henwe
returned to Potrero Viejo.
On March
17we
startedon
a side trip, lasting untilMarch
24,which
ledby highway through Tehuacan, Tecamachalco,
thencenortheastward on
theJalapa
road, toTeziutlan (Puebla)from El Limdn,
Totalco,and eastward
toPuente
Nacional.Shortly thereafter
we
left forTehuantepec by
rail again, arriving April 1.We
stayed onlyone day and
traveledby
railtoAcapetahua, whence we went by bus
to Escuintla.From
therepack animals were used
to transportour equipment
toLa Esperanza,
a fincaabout
5 miles northeast of Escuintla,We
arrived thereon
April 4and
re-mained
untilJune
5, 1940.Various
short side tripswere taken
tonearby
fincas.On June
5we
returned toTehuantepec
and, afterone
day, toPotrero
Viejo,arrivingJune
9.About one month
later,on
July6,we
crossed the
border
atLaredo with no
deviationsfrom
the directroutefrom
Potrero Viejo to the border.8
We
returnedimmediately
toMexico City by way
of thePan American Highway and remained
there untilAugust
7, except for a tripby
railtoGuanajuato, Guanajuato, on July
19-21.On August
8we
againdrove
to PotreroViejo.Within
afew days we made
abriefand
finalforayalong thehighway from
AcultzingotoTehuacan
beforepacking
allequipment
thathad been
stored at Potrero Viejo,We
left
our headquarters
there for the finaltime on August
8 and,with but
a briefstop inMexico
City, traveledslowlynoithward
along thePan American Highway, with
aheavy
load, crossing theborder
atLaredo on August
24.The
followinggazetteer includesonly
thoselocalitiesrepresentedby specimens
in theBacon
collection that arenot
tobe found on
theNational Geographic
Society's1939 map
ofMexico, Central America, and
theWest
Indies, scale 1:5,702,400.Most
of the localities are plottedon
other, larger-scalemaps, but inasmuch
as these arenot always
readily availablewe
include all missingfrom
theGeographic
Society'smap, which
is readily available to all investigators.The
localities are
arranged
alphabetically within the states,which
arethemselves
alphabetically arranged.CHIAPAS Acacoyagua: 3 milesnorthwestofEscuintla.
Ag^uacate: 7 milesnorth-northeastby northofPalenque.
Belen: 20milessoutheastofEscuintla.
Cerro
Obando:
3 milesnortheastofLa
Esperanza (whichsee).Colonia Hidalgo: 7milesnortheastofAcacoyagua (whichsee).
Colonia Soconusco: 12 milesnortheastof
La
Esperanza (whichsee).Cruz
delaPiedra: 2 mileswestofLa
Esperanza (whichsee).FincaJuS.rez: 10 milesnortheastof
La
Esperanza (whichsee).Javarinero: 13 miles eastofPalenque.
La
Esperanza: 3 miles east of Acacoyagua (whichsee).La
Magnolia: 3 milesnorthwestofLa
Esperanza (whichsee).Las
Nubes:On
Cerro Obando, 2 miles northeast ofLa
Esperanza (which see).Motozintla: 25mileseast-northeastbyeastofEscuintla.
Bancho Las
Gradas: 2 mileswestofLa
Esperanza(whichsee).Saltode Agua:
On
CerroObando, 1milenortheastofLa
Esperanza (whichsee).San
Juanito: 1 mileeast-northeastof Palenque.Santa
Bosa: NearComitdn.CHIHUAmjA Carmen:
32miles east-northeastbyeastofGaleana.Carrizal: 10 mileswest-southwestby southof
Ahumada.
Ciudad
Delicias: 5 milessouthof Meoqui.Ojo deFederico:
A
smallranch8 milessoutheast ofAscenci6n.Progreso: 35milesnortheastofGaleana, near Rfo Santa Maria.
Rancho
Nuevo:A
smallranch 10 mileswest-northwestby westofCarrizal.Samalayuca:
25milessouthofCiudadJudrez.CHECKLIST OF REPTILES OF MEXICO
COAHUILA.
Hip61ito: 25milesnorth-northeastofGeneral Cepeda.
Zapaliname:
A
mountain3 milessouthofSaltillo.DISTRITO
FEDERAL
Atzacoaico: 10 milesnortheastof MexicoCity.Cafiada deContreras: 5 milessoutheastof Mexico City.
DURANGO
La Qoma:
12 mileswestofLerdo,onthesouthside of theRfoNazas.La Loma:
12 mileswestofLerdo,on the northsideofthe Rfo Nazas.Pedricefla: 25 mileseast-southeastofNazas.
GUATEMALA
Desempefia: 4 milessoutheastofPiedras Negras.Pozo
de laJicotea: 2 milessoutheastofPiedras Negras.GUERRERO Agua
Bendita: 13 milessouthofTaxco.Agua
delObispo: 8milesnorthofDos
Caminos.Cacahuamilpa:
10 milessoutheastofTetecala, Morelos.Cojruca: 7 milesnortheastofAcapulco.
ElTreinta: 20 milesnorthofAcapulco.
Garrapatas: 5 milessouthwestofDosCaminos.
JulianBlanco: 4 milesnorth of
Dos
Caminos.Ocotito: 3 milesnorthof DosCaminos.
Omilteme:
3 milesnorthwest ofChilpancingo.Paso
delLimonero:
9 milesnortheastofAcapulco.TierraColorada: 3 milessouthwest ofGarrapatas (whichsee).
Xaltianguis: 31 milesnortheastofAcapulco.
HIDALGO
Barranca
delosHorcones: 6milessouthof Durango(which see).Chapxilhuacan: 19 milessouthwestofTamazunchale, San Luis Potosl.
Colonia: 6 milessoutheastofPachuca.
Durango:
15 mileswest-southwestbysouthofJacala.ElChico
Parque
Nacional: 15milesnortheastofPachuca.Maguey
Verde: 7 milessouthof Durango.Tianguistengo: 10 milesnorth-northeastofZacualtipdn.
MEXICO
Chalco: 13 milesnortheastofXochimilco.Chimalhuacan:
12 miles east ofMexico City.Lerma:
10 miles eastofToluca.Nevado
de Toluca: 32 milessouth-southwestbysouthofToluca.RioFrio: 18 mileswest-northwestofTexraelucdn, Puebla.
Santa Magdalena:
15 miles eastof Mexico City.Zempoala:
6mileswest-southwestofTres Cumbres, Morelos.MUSEUM
michoacAn
Carapa(n): 23milesnorthofParacho, ElTemazcal:
20miles eastof Morelia.La Palma:
10 milesnorth ofSahuayo.Puerto
Hondo:
23 miles east of Moreha.Bancho San
Jose: 24miles eastofMorelia.Tacicuaro: 5 miles eastofQuiroga.
NUEVO le6n Mamulique
Pass: 45milesnorth ofMonterrey.Santa
Caterina: 7 mileswestof Monterrey.OAXACA
Caj6n
dePiedra: 12 milessouth-southwestofSalinaCruz.Cerro Arenal: 16 mileswestof
Tehuan
tepee.Cerrode Chipehua: 16milessoutheastofSalinaCruz.
Cerro de
Huamelula:
34mileswest-southwestbysouth of Tehuantepec.Cerro Quengola: 10 mileswest-northwestbywestofTehuantepec.
Cosolapa: 8 miles northwestofAcatMn.
Coyol: betweenSan Antonioand Las Cruces.
El Lim.6n: 25milessoutheastofTehuantepec.
Escurano: 15 mileswest-northwestofTehuantepec.
La
Concepci6n: 32 mileswestofTehuantepec.Lachiguiri: 29milesnortheastofTehuantepec.
La
Gloria: 8milessoutheastofChimalapa.LasPilas: 4 milesnorthwest ofTehuantepec.
Las Vacas: 40mileswestofTehuantepec.
LlanoOcotal: 20mileswest-southwestby westofTehuantepec.
Matias Romero:
34miles northofSan Ger6nimo.Mixtequilla: 4 milesnorthwest ofTehuantepec.
Palmar:
37mileswest-northwestbywestofTehuantepec.Pixixi: 6 milessouthofTehuantepec.
Portillo
Quayabo:
16 miles westof Tehuantepec.Portillo
Las
Vacas: 40mileswestofTehuantepec.Portillo
Los Nanches:
32 milesnorthwestofTehuantepec.Rincon San
Pedro: 16milesnortheastofTehuantepec.Rio Grande: 20 miles west-southwestby south ofTehuantepec.
Bio
Hondo:
41 mileswest-northwestbywestofTehuantepec.San
Francisco Guichina:61 mileswest-northwestbywestofTehuantepec.San
JoseManteca:
61 mileswest-northwestbywestofTehuantepec.San Mateo
delMar:
14 mileseast-northeastbyeastofSalinaCruz.San
Pedro Quiechapa: 10 mileswestofYautepec.Tenango:
24mileswest-southwestbywestofTehuantepec.Tres Cruces: 43miles west-northwest by west of Tehuantepec.
Yerba
Santa: 10 miles west-northwestby west ofTehuantepec.PUEBLA
Alchichica: 10 miles southwest of Perote, Veracruz.
Cacaloapam:
8 milessoutheastofTlacotepec.El Seco: 27milesnortheastofTepeaca.
La
Virgin: 2 miles north of Cacaloapam.CHECKLIST OF REPTILES OF MEXICO 11
Ozumbilla: 4 mileseast-southeastof Morelos.
PajaroVerde: 300feetwestofPuebla- Veracruzstateline,about 7 miles south- east of Morelos.
Tecaraachalco: 15 miles northwest ofTlacotepec.
SAN LUIS POTOSf Pujal:20milesnorthofTancanhuitz.
TABASCO El Retire: 13 milessouthof Tenosique.
LosRieles: 10 miles south of Tenosique,
Santo Tomas:
12 miles southeast ofTenosique.TAMAULIPAS
Ciudad
Mante (=
VillaJudrez): 18 miles east-northeast by north of Antiguo Morelos.Forlon: 18 miles east-southeast of Llera.
Hacienda La
Clementina:13 mileseast-southeastofLlera.TLAXCALA
Apizaco: 8 milesnorth-northwestbynorthofTlaxcala.
VERACRUZ
Acultzingo:9 miles east-southeastbyeastofMorelos, Puebla.
Atoyac:4 mileseast-northeastofPotreroViejo (whichsee),
Cerro Gordo: 20 miles east-southeastofJalapa.
Cuautlapan: 4 miles southwest of Fortin (which see).
Cruz
Blanca: 8 miles northeast of Perote.ElLiraonTotalco:10 milessouthwestofPerote.
El
Maguey:
8 miles east-southeastbyeastofPotreroViejo (whichsee).Encero: 8 miles east-southeast ofJalapa.
Fortin: 5 miles eastof C6rdoba.
Matacabestro: 7 milesnorth-northwest bynorth ofJoaqufn.
Mata
de Cafia: 18 miles east-southeastofJalapa.Medellin:10 milessouthofVeracruz.
Metlac: 1milewestofFortfn (whichsee)
.
Ojo de Agua:2 miles eastofParaje
Nuevo
(whichsee),near PotreroViejo.Pan
deOUa:
8 milessouthofTeziutldn, Puebla.ParajeNuevo:2 miles eastofPenuela (whichsee).
Paso
delMacho:
5 milesnortheastofAtoyac.Pefiuela: 4 milessoutheastofC6rdoba.
PotreroViejo: 2miles eastofParaje
Nuevo
(whichsee).Presidio: 22miles southwestof C6rdoba.
San Juan
deLa
Punta: 15mileseast-southeastbysouthofC6rdoba.San
Jose de Gracia: 13 miles southeastofC6rdoba.Palma
Sola:9 miles east-southeastby southofC6rdoba.Tequejru tepee: 10 miles northeast of Jalapa.
Tezonapa: 8 miles northwest of Acatldn.
Toxtlacuaya: 17 miles northwest of Jalapa.
Xuchil:1milenorthofPotreroViejo(whichsee).
861316—60 2
12 BULLETIN
199,UNITED STATES NATIONAL MUSEUM Class REPTILIA Laurenti
ReptiliaLaurenti, Specimen
medicum
exhibenssynopsin reptilium, 1768,p. 19.Subclasses.
— Three
subclasses are representedby
livingmembers.
Alloccur in
Mexico.
KEY TO MEXICAN'
SUBCLASSES, ORDERS,AND SUBORDERS
OF REPTILIA1. Provided with ashellencasingbody above andbelow, consistingofplastronand carapace;turtles..subclass
Anapsida
orderTestudines(p.12)_ 5Not
provided with ashell 22. Anal slitlongitudinal;skull diapsid subclassArchosauria.
orderLoricata(p. 209) Anal slit transverse; skull modified diapsid; lower (quadratojugal-quadrate) arch interrupted.. subclass Lepidosauria order
Squamata
(p. 37) 3.Two
limbs only, the forelegs; body ringed with grooves.suborder
Amphisbaenia
(p. 37) Fourhmbs
ornone; bodynotringedwith grooves 44. Limbspresentor,ifnot, movableeyelidspresent;lizards.
suborder Sauria (p.39) Limbs absent;movable eyelids absent; snakes suborderSerpentes
5.
No
epidermalscutesonshell 6Epidermalscutespresent 7
6. Limbswithoutexternalevidenceof digits, oar-shaped, clawless.
suborder
Atheca
(p. 13)Limbswithdistinctdigits,3with claws suborderTrionychoidea (p. 18) 7. Limbsoar-shaped, with 1 to 3claws suborder Carettoidea (p. 14)
Limbsnot oar-shaped, atleastforelegwith 4or 5 claws.
suborder Cryptodira (p. 18)
Subclass Anapsida Williston
Anapsida Williston,Journ. Geol.,vol. 25, 1917,p.419.
Orders.
— A
singleliving order exists, the Testudines.Order TESTUDINES Batsch
Testudines Batsch, Versuch Anleit. Kenntn. Gesch. Thier. Mineral., vol. 1, 1788, p. 437.
Chelonia
Macartney,
inRoss, Transl. Cuvier'sLegons d'anatomiecompar^e...vol. 1, 1802, tab. 3.
TestudinataOppel,DieOrdnungen,Familienund GattungenderReptilien . . . ,
1811, p. 3.
Suborders.
— Five
living suborders ofTestudines
are recognized;four
occur
inMexico, and
thefifth (Pleurodira) isrestricted to south- ern portions ofboth
hemispheres.»Charactersinallkeys applytoMexican forms butnot necessarilytothoseinother partsofthe world.
CHECKLIST OF REPTILES OF MEXICO 13 Suborder Atheca Cope
Athecae Cope, Proc. Amer. Assoc. Adv. Sci., vol. 19, 1870, p.235.
Families.
— One.
Family DERMOCHELYIDAE
* Fitzinger DermatochelydaeFitzinger, Systemareptilium, 1843, p. 30.Genera.
— One.
Range. —
Tropicaloceans
of the world.Genus DERMOCHELYS
Blalnville Dermochelys Blainville, Bull. Soc. Philom., 1816, p. 119.Genotype.
—
Testudo coriaceaLinnaeus.Range. — World-wide,
in tropicaloceans and
occasionally intotem-
perate waters.Species.
— As many
as three species (or subspecies)may be
valid.The
specieshas been
recorded inMexico
onlyfrom
thewestern
coast,although
Atlanticrecords are tobe
expected.DERMOCHELYS
CORIACEA• (Linnaeus)Testudo coriaceaLinnaeus, Systemanaturae,ed. 12, vol. 1, 1766, p. 350.
Dermochelyscoriacea,Blainville, Bull. Soc. Philom., 1816,p. 119.
—
Boulenger, Catalogue of the chelonians . . ., 1889, p. 10.
—
Deraniyagala, Tetrapod reptiles of Ceylon, vol. 1, 1939, pp. 38-102, figs. 12-34.—Schmidt, Marine Life Occ. Pap., vol. 1, No.3, 1945, pp. 7-10.
Sphargis coriaceaschlegelii
Garman,
U.S. Nat. Mus. Bull. 25, 1884, p. 303 (type unknown;type locality, tropicalPacificand Indian Ocean,here restricted to Guaymas, Sonora).Dermochelysschlegelii, Stejneger,U. S. Nat. Mus. Bull. 58, 1907, p. 485.
Type
.— Unknown
.
Type
locality.— Palermo,
Sicily,by
present restriction.Range. —
Pacific coastand probably
the tropical Atlantic coast.Recorded
definitelyfrom
Sonora:Guaymas; Baja
California:Los Coronados
Islands.'ActuallythefirstfamilynameproposedforthisgroupwasSphargidaeGray(Ami.Philos., vol.10,1825, p. 212),baseduponthegenusSphargisMerrem,1820,a juniorssmonymofDermochelysBlainville, 1816;
bothgenera arebaseduponthesametjT)e,TestudocoriaceaLinnaeus. Manytaxonomists retain the oldest familynameregardlessofthe statusofitstype.
We
believethereisa certaindegreeof errorcourtedby sucha procedure,however,inasmuchasafamilynamebasedupona genericnamelaterfoundtobea juniorhomonymofanearliernameinanotherfamilywould, veryunfortunately,bepreserved;andwhatifthe seniorhomonymhadalsoserved as atypeforitsfamily? Itseemstousthatonlycurrently valid generic namesshouldbeavailable astypesforfamilynames,andthatallsynonymsandhomonymsshould be regarded unavailableforthat purpose.
•Acommonarrangementrestricts coriacea tothe Atlantic,schlegeliitothePacific,eitherasspecies or.sub- species.
We
areunabletofindthatanythingmorethangeographic probabilityhas ledtosuch arrangement*14
Suborder Carettoidea^ Fitzinger
Carettoidea Fitzinger,
Neue
Classificationder Reptilien . . ., 1826, p. 5.Families.
— One.
Family CHELONIIDAE Gray
CheloniadaeGray, Ann. Philos., ser. 2, vol. 10, 1825, p. 212.
Genera.
— Four.
Range. —
Tropical oceans, world-wide.KEY TO GENERA OF THE FAMH^Y
CHELONIIDAE'1.
One
pairofprefrontal scutes; costal scutes4; hornycuttingedge oflowerjaw coarselydentate, thatofupperjawstronglyribbedvertically; bonyalveolar ridge of upper jaw with a low but regularly raised auxiliary ridge behind anteriorridge, which is very strongandterminates anteriorly ina tooth at posterolateralcornerofpremaxillarypit Chelonia (p. 17)Two
pairs of prefrontals; costal scutes4-9; horny cutting edge of lowerjaw smoothor feebly denticulate, thatof upperjawwithout markedlyelevated vertical ribbing on its inner surface; bony alveolar surface of upper jaw smooth or with a single ridge; this ridge not terminating anteriorly in asharp tooth 2
2. Costal scutes 4 pairs; dorsal scutes usually conspicuously imbricate; bony alveolarsurfaceofupperjawwith asharp-crestedridge.
Eretmochelys
(p. 16) Costalscutes5 pairsormore;dorsalscutesnot conspicuouslyimbricate;bony alveolarsurface ofupperjawsmoothor with aroundedridge 3 3. Fourenlarged inframarginalscutesonbridge; dorsal colorgray to olive-green;maxillariesnotincontact,separatedbyprevomer Lepidochelys (p. 14)
Three enlarged inframarginal scuteson bridge; dorsal color brownor reddish brown;maxiUariesincontact betweenprevomer andpremaxillaries.
Caretta (p. 15)
Genus LEPIDOCHELYS
Fitzinger LepidochelysFitzinger, Systemareptilium, fasc. 1, 1843,p. 30.Caouana Gray, Catalogueofthetortoises
...
inthe BritishMuseum,
1844, p.52 (type, CheloniaolivaceaEschscholtz).
Genotype.
—
Chelonia olivaceaEschscholtz.Species.
— Two
are generallyrecognized.Range: —
Tropical oceans, world-wide.KEY TO
SPECIES OF LEPIDOCHELYS1. Inframarginalscuteswithoutpores;usually 5 pairsof costalscutes;colorusually gray; bonyalveolarsurfaceofupperjawwith a conspicuousridge.
kempii
(p. 15)Eachinframarginal scute usually with a pore at itsposterior border; usually morethan5 pairsof costalscutes; colorolive;bonyalveolarsurfaceofupper jawwith a low (not conspicuous) ridge olivacea (p. 15)
'Deraniyagala(Spolia Zeylanica,vol. 24, 1945, p. 98)suggeststhatthisgroupmaybeofpleurodirous origin.
'AdaptedfromCarr, Proc.NewEnglandZool.Club,vol. 21, 1942,pp.3-5.
CHECKLIST OF REPTILES OF MEXICO 15
LEPIDOCHELYSKEMPH
(Gairoan)Thalassochelys {Colpochelys) kempii
Garman,
Bull. Mus.Comp,
Zool,, vol. 6, 1880, p. 123,Lepidochelys kempii, Baur, Araer. Nat., vol. 24, 1890, p. 487.
—
Carr, Proc.
New
England Zool. Club, vol. 21, 1942, pp. 4,8-13, pis. 2, 4.Type.
—
Several cotypes,Mus. Comp.
Zool.Type
locality.— Gulf
ofMexico,
hererestricted toKey West,
Fla.Range.
—
AtlanticOcean and Gulf
ofAlexico.In
Alexico,the entire Atlantic coast.The only
exact recordknown
isfrom Quiatana Roo
(Islade
Mujeres).LEPIDOCHELYS OLIVACEA (Eschscholtz)
Chelonia olivacea Eschscholtz, ZoologischerAtlas, pt, 1, 1829, p. 2,pi. 3.
Lepidochelys olivacea, Girard, United States Exploring Expedition . . . , vol.
20, Herpetology,1858,p.435.
—
Deraniyagala, Tetrapodreptiles ofCeylon, vol. 1, 1939, pp. 123-163, figs. 43-65.—Carr, Proc.
New
England Zool.Club,vol. 21, 1942, pp. 4-5, pi. 5 (skull).
Chelonia dussumieri Di;m:6ril and Bibron, Erp^tologie g6n4rale . . . , 1835, p.
557 (typein Mus. Hist. Nat.Paris;typelocality, Malabar).
Careitaremivaga Hay, Proc. U. S. Nat. Mus., vol. 34, 1908, pp. 194-197 (type, U.S.N.M. No. 9973; typelocality, Ventosa Bay, Oaxaca).
Type.
— Unknown,
Type
locality.— Manila Bay,
Philippine Islands.Range. — Indian and
PacificOceans.
InMexico,
the entire Pacific coast.Reported
onlyfrom
the states of Chiapas,Oaxaca,
Guerrero,Colima, and Sonora (Tibur6n
Island).Genus CARETTA
Rafinesque CarettaRafinesqtje, SpecchioSci. Palermo,vol. 2, 1814, p. 66.Thalassochelys Fitzinger, Ann.
Wien
Mus., vol. 1, 1835, p. 121 (type, Testudo caouana Daudin).Caouana Cocteatt, in
Ram6n
de la Sagra, Historiaffsica, polftica y natural de la Islade Cuba,vol. 4, Rept., 1838,p. 31 (type, Testudo cephalo Schneider).Eremonia Gray,
Hand
list ofshieldreptiles of the BritishMuseum,
1873, p. 91 (type, Caouanaelongata Gray).Genotyye.
—
Caretta nasuta Rafinesque=(7areffa caretta caretta (Lin- naeus).Species.'
— One, with two
races, isgenerally recognized.Range. —
Tropical ocean, world-wide.KEY
TO SPECIES OFCARETTA
1. Neural bonesusually 9 ormore;iffewer, thecostalbonesusuallyinterrupting the neural seriesbycontact with eachother atoneor moreplaces.
caretta gigas (p. 16) Neural bones 7 or8, theseries rarely interrupted by median contactofcostal
bones caretta caretta (p. 16)
16 BULLETIN
199,UNITED STATES NATIONAL MUSEUM
CARETTA CARETTA CARETTA
(Linnaeus)TestudocarettaLinnaeus, Systemanaturae, ed.10, vol.1, 1756,p.197.
Caretta caretta, Stejneger, Ann. Rep. U. S. Nat. Mus., 1902 (1904), p. 715.
—
Care, Proc.
New
EnglandZool. Club,vol. 21, 1942, p. 5, pis. 1, 3.Caretta caretta caretta, Deraniyagala, Tetrapodreptiles of Ceylon, vol. 1, 1939, p.164; Spolia Zeylanica, vol. 24, 1945,p. 95.
Testudo cephalo Schneider, Algemeine Naturgeschichte der Schildkroten . . .,
1783, pp. 303-308 (type unknown; type locality, Charleston, S.
C, by
presentrestriction).
Testudocaouana Daudin, Histoire naturelle des reptiles, vol.2, 1802,p. 55, pi.
16, fig. 2 (typein Mus. Hist. Nat.Paris;typelocality, Jamaica, bypresent restriction)
.
Type. — Unknown.
Type
locality.— "About
theAmerican
Islands,"hererestricted to theBermuda
Islands.Range. — Western
AtlanticOcean;
the entire Atlantic coast ofMexico. Reported
inMexico only from Yucatdn.
CARETTA CARETTA
GIGASDeraniyagalaCarettagigasDeraniyagala, CeylonJourn.Sci.,sect. B,vol. 28, 1933, pp.61-62.
Caretta caretta gigas, Deraniyagala, Tetrapod reptiles of Ceylon, vol. 1, 1939, pp. 164-185, figs. 66-73; Spolia Zeylanica, vol. 24, 1945, p. 95.
Type.
— Presumably
inColombo Museum, Ceylon.
Type
locality.—
Ceylon.Range. — Indian and
PacificOceans and
possibly eastern AtlanticOcean. In Mexico,
the entire Pacific coast.Reported only from
Sinaloaand Baja
California.Genus ERETMOCHELYS
Fitzinger EretmochelysFitzinger,Systemareptilium,fasc. 1, 1843,p.30.Genotype.
—
Testudo imbricata Linnaeus.Species.
— One
is generally recognized,but two
species (or sub- species) are distinguishedby some
authorities.ERETMOCHELYS
IMBRICATA« (LinnaeuB)TestudoimbricataLinnaeus, Systemanaturae,ed. 12, 1766,p. 350.
Eretmochelys imbricata, Fitzinger, Systema reptilium, fasc. 1, 1843, p. 30.
—
Agassiz, Contributions to the natural history ofthe United States, vol. 1, 1857, p. 381.
—
Deraniyagala, Tetrapod reptiles of Ceylon, vol. 1, 1939, pp. 187-217, figs. 74-85.
Chelonialachrymata Cuvier, LerSgne animal,ed. 2, vol. 2, 1829, p. 13 (notype ortypelocalitydesignated;
Bermuda
Islandsbypresentrestriction).Chelonia pseudo-mydas Lesson, in Belanger, Voyage aux Indes Oriental . . .,
Zool., 1834, p. 299 (type unknown; type locality, Atlantic Ocean, here re- stricted to the
Bermuda
Islands).Chelonia pseudo-caretta Lesson, op. cit., p. 302 (type and typelocality as inthe preceding)
.
'TheAtlanticOcean specimensarefrequently regarded as Eretmochelys imbTicata(orE.i.imbricata), the PacificOceanones as Eretmochelys imbricatabissa (orE.i.sguamata).
We
areunabletodetermine charactersforsuchseparation.CHECKLIST OF REPTILES OF MEXICO 17
Carettabissa RCppell,
Neue
Wirbelthiere Abyssiniens, Amphibiens, 1835, p. 4, pi. 2 (typeunknown;typelocality,Red
Sea).Eretmochelys squamata Agassiz, Contributions to the natural history of the United States, vol. 1, 1857, p. 382 (cotypes, Mus.
Comp.
Zool. Nos. 1415, 1416; lectoholotype, No. 1416; typelocality restricted to Singapore, Straits Settlements).
Type.
— Unknown.
Type
locality.— American
seas, here restricted to theBermuda
Islands.
Range. —
Tropical oceans, world-wide;both
coasts ofMexico.
Reported from Baja
California, Sonora,Oaxaca, Campeche, and Quintana Roo
(Islade
Mujeres).Genus CHELONIA
BrongniartCheloniaBrongniart, Bull. Sci. Soc. Philom., vol. 2, 1800, p. 89.
CheloneBrongniart,
Mem.
Sav. fitrang, vol. 1, 1806, p. 610.ChelonaBurmeister,
Handbuch
derNaturgeschichte,vol.2, 1807, p.731.CheloniasRafinesque, SpecchioSci.Palermo,vol.2, 1814,p. 66.
Mydas
Cocteau, inRam6n
de laSagra, Historia Ksica, pollticay natural de laIslade Cuba,vol. 4, 1838, p. 22.
MydaseaGervais, Diet. Hist. Nat.,vol.3, 1843,p.457.
EucheloniaTschudi, Untersuchungeniiberdie
Fauna
Peruana . . ., 1846,p. 22.Megemys
Gistel, Naturgeschichtedes Thierreichs, 1848,p. 8 (typeofallabove, Testudo mydas Linnaeus),Genotype.
—
Testudomydas Linnaeus.
Species.
—
Possibly fourforms
(species or subspecies) are recogniz- able.None
ofthesearewellenough known
toustodiagnose properly.Range. —
Tropical oceans, world-wide.CHELONIA
MYDAS'
(Linnaeus)Testudomydas Linnaeus, Systemanaturae,ed. 10, 1758, p. 197.
Cheloniamydas,Brongniart,Bull. Sci.Soc. Philom.,vol.2,1800,p.
89.—
Schweig- ger, Konigsberg. Arch. Naturw. Math., vol. 1, 1812, p.412.—
Deranita- GALA,TetrapodreptilesofCeylon,vol. 1, 1939, pp.218-242,figs.86-95.Testudo viridis Schneider, Algemeine Naturgeschichte der Schildkroten . . ., 1782, pp. 299-303(typeunknown;typelocality,Charleston,S.
C,
bypresentrestriction)
.
Chelonia virgata Schweigger, Prodromi monographiae cheloniorura . . ., 1814, p.21 (typeunknown;typelocality, "SeasofTorrid Zone," here restricted to the
Bermuda
Islands).Chelonia maculosa Cuvier, Ler^gne animal . . .,ed. 2, vol. 2, 1829, p. 13 (type in Mus. Hist. Nat. Paris?; type locahty not designated, here restricted to AscensionIsland).
Cheloniaagassizii Bocourt, Ann. Sci. Nat., ser. 5, vol. 10,1868,p. 122 (type in
Mus. Hist. Nat.Paris;typelocality,
mouth
ofRfo Nagualate, Guatemala);Mission scientifique au Mexique . . ., Etudessur les reptiles, livr. 1, 1870, pp. 26-28,pi.6 (color).
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