Vo1
'1920
CYI1
] Chapman, Proposed Neiv Race of the Killdeer.
105
DESCRIPTION OF A PROPOSED NEW RACE OF THE KILLDEER FROM THE COAST OF PERU.
BY FRANK
M.CHAPMAN.
Collections
received during 1919,by
the AmericanMuseum
from its Peruvian representative,
Mr. Harry
Watkins, contain fourteenspecimensofa Killdeerwhichbreeds onthe coastofPeru, at least fromLima
to near the Ecuadorian boundary. Killdeerhave
been before recorded from Peru,1 from Colombia,2 Ecuador,3 Paraguay,3and
Chile,3 but it has beenassumed
that these birds were winter visitants fromNorth
America. It seems, however, not improbablethat they wereresident birds, as,beyond
question, are our specimens from Peru. These represent adults at the beginningand
theendofthe post-nuptial molt,and young
infresh juvenalplumage
andin thedown.This discoveryplaces theKilldeer, distributionally, in the group of Plover to which Octhodromiis wilsonius
and
Mgialitis collaris4 belongand
suggests thatournorthern Killdeerisderivedfrom the SouthAmerican
form.Of
Mgialitis collaris,Ridgway
remarks thatSouthAmerican
specimens"much more
often(infact usually) have thecinnamon
on head and neck presentand
alsomore
pro- nounced,"5and
it is in the greater extent of the rustymargins of the upperparts that the Peruvian Killdeermay
be distinguished from theNorth American and West
Indianforms. Thisdifferenceissufficientlypronouncedand, so far asour material goes, constant to warrant the recognition of the Peruvian bird as a well-marked raceforwhich Ipropose the
name
'Scl.&Salv., P. Z.S.,1S6S,p. 176(TamboValley,southwesternPeru); Taczanowski, Ibid.,1S79.p.244 (Pacasmayo).
2Scl.&Salv.,P.Z.S.,1S79,p.547(Medellin).
sSharpe, Cat. Bds.Brit.Mus.,XXIV,pp. 247, 742.
4In default of material to consider the conclusionsinregard togenerareachedby Ridg- way(Bull.U.S.N. M.,50,Pt.VIII),Ifollowhere thenomenclatureofthe BritishMuseum
Catalogue.
5Bull.U.S.N. M.,50,Pt.VIII,p.141.
106
Chapman, ProposedNew
Race of the Killdeer. [j"nOxyechus
vociferusperuvianus new
subspecies.Subsp. Char.
—
Smallerthan Oxyechusvociferusvociferus,agreeinginsiiewith the West Indian Oxyechus vociferus rubidus, but in post-nuptial plumagedifferingfromthembothinthegreaterextentoftherustymargins oftheplumageoftheupperparts and,particularly, ofthelesserand median wing-coverts.
Type.
—
No. 163,083,Am.
Mus. Nat. Hist., <? ad. (fresh post-nuptial plumage,theouterprimaryofthe nuptialplumagestillpresent),Paletillas, alt. 1550ft., northeast of Payta, Prov. Piura, Peru, June 22, 1919; H.Watkins.
Description of Type.
—
Similar in color to Oxyechus vociferus vociferus but all thebrown feathers of the plumage, including crown, nape, back, scapulars, tertials, etc. margined with rusty orochraceous; exposed por- tions of lesser and median wing-coverts rusty. Wing, 160; tail, 91;tarsus, 31.5; exposed culmen, 19
mm.
Description of freshJuvenalPlumage.
—
9 juv.,Bequeta, nearLima, Peru, Jan. 22, 19T9. Similar to corresponding plumageof Oxyechus vociferus vociferus.DescriptionofNatal Doivn.
—
(Twospecimens notmorethanfour daysold, Paletillas, Peru, June 22, 1919). Similar to corresponding plumage ofOxyechusvociferusvociferus.
Specimensexamined.
—
Oxyechusvociferusperuvianus. Peru: Paletillas, Piura, 1&
ad., 2pull. (June 22, 1919); Samate, Piura, 2 c? <? ads.(May
30, 1919); Chilaco, Piura,1
d
1 ad., 1 9 ad.(May
27-25); Pilares,Piura, 1 <? ad.,1 9 ad.(June16,1919); Bequeta, Prov.Lima,2& &
ads.,2 9 9 ads., 1 9 juv. (Jan.22-24, 1919).Oxyechusvociferus rubidus. WestIndies: (Cuba, Haiti,Jamaica, Porto Rico,Grenada), 18 adults representing both sexesand every month but August.
Oxyechusvociferusvociferus.
A
largeseriesfrom throughouttheUnited States,takenineverymonth andrepresentingallplumages.Remarks.
— The seven adult specimens on which this proposed
raceischieflybased,were takenintheProvince ofPiurafromMay
25 to June 22. All are essentially in the
same
stage of plumage, havingjustcompleted, or nearlycompleted, the post-nuptial molt,all but one still havingtheouterone ortwoprimariesof thenup-
tial plumage.
The
series shows little variation in colorall having theupperparts stronglymargined withrusty, as inthetype. These birds are comparable with Augustand
September specimens from the eastern United States.The
latter often have the upperparts° 1920 J Chapman, Proposed
New
Race of the Killdeer.107
margined with rusty, butnever, so far as
my
observations go, to the extentshown by
the Peruvianbird.Some
comparableNorth
American birds, on the other hand,show
almost no trace of this rusty margining, and, representing the extreme of difference between peruvianasand
vociferus vociferus,may
perhaps indicate the type of coloration towardwhichthe latter isdiverging.That
this divergence is of comparatively recent occurrence in the life ofthespecies, issuggested
by
theinconstanceofthe featureswhich characterize it,and
alsoby
the fact that in juvenalplumage
the Peruvianand North
American forms are alike.The West
Indianform appearstodifferfrom true vociferus only in size,and
consequently, is not intermediate in color betweenit
and
peruvianus.As
stated above,two downy young
butafew days from theegg, were takenatPaletillas,June 22,and we may
accept these birds, inconnectionwiththeseven adultsfromthePaletillasregionwhich are completing their post-nuptial molt, as conclusive evidence that thePeruvian Killdeernests inMay and
June.But
examina- tion of the Bequeta specimens shows thatMay and
Juneby
nomeans
constitutetheentire nesting seasonof the KilldeerinPeru.Four
of the Bequeta specimens, taken January 23and
24, are adults in the midst of the post-nuptial molt in which wingsand
tail as well as
body
feathers, are being renewed.The
remaining feathers of the nuptialplumage
aremuch worn and
practically withoutrusty margins; theincomingnew plumage
ismargined with rusty.The
fifthBequetaspecimen, takenJanuary22,isinfreshjuvenalplumage
withportionsof the nataldown
stilladheringto theends of the central rectricesand
longer upper tail-coverts.Our
col- lection contains specimens whichshow
that in the Killdeer frag- ments of the nataldown may
remain at the end of the central retrices until the spring following the bird's birth—
a surprising fact—
but the general condition of theplumage
of this Bequeta bird withitsfresh, narrowlymargined dorsalplumage, shows thatit isacomparatively
young
bird, exactlysimilar, indeed, toAugust specimens of vociferus vociferus from various parts of the United States. Consequently,just asthe Piurabirdsprovethat innorth- westernPeru the Killdeer breedsinMay and
June,sotheBequeta108
Cory,New
Species and Subspecies of Tyrannidce.[jEm
birds prove that in central western Peru the Killdeer breeds in
December
and January.The
localities in question are only about 450 miles apart. Iknow
ofnomarked
seasonaldifferencebetween them,and am
quiteat loss toaccount satisfactorilyfor this variation in nesting dates.
Possibly theuniformclimate,bothasregardstemperature
and
rain- fall, prevailingonthe coastofPeru,militates,among
Killdeer, asitapparently does
among
theCormorantsand
Pelicans of theGuano
Islandsoffthecoast,1against theestablishmentofadefinitebreeding season.
Or we may
have here a casesimilar to thatof theBrown
Pelicans in Florida which on the Gulfcoastbegin to nest in April
and
on the Atlantic coast inNovember.
Further collectionsand
field studies are required to settle this interesting question Amer.
Museum
Nat. Hist., N. Y.DESCRIPTIONS OF A NEW SPECIES AND SUBSPECIES OF TYRANNIDtE.
BY CHARLES
B. CORY.Todirostrum
beckeri sp. nov.Type from BaseofSerradaLua, nearBoaVista,RioBranco, N.Brazil
.
Male, No. 49,347,Field
Museum
ofNatural History. CollectedbyR. H.Becker, March24, 1913.
Description.
—
Similar to T. sylvia schistaceiceps (Sclater) and T. sylvia griseolumTodd, butdiffersfrom eitherinthe shorter wing, inhaving the blackloral stripe borderedbelowby a buffy streak, and aboveby a con- spicuous buffy stripe extending from the base of the upper mandible to abovetheeye. Baseofcrownand napeolivaceous; greaterwingcoverts edged withbuffyyelloworpaleorange yellow; middlewingcovertstipped withsame; lesserwingcovertsedged with olivegreen.Measurements.
—
Wing, 45; tail, 30; bill, 13mm.
1Habits and Economic Relations of the Guano Birds of Peru, by Robert E. Coker,.
U.S.BureauofFisheries,Proc.U.S.N. M.,56,1919, pp.449-511.