34; It is proposed to publish a series of reports, giving an account of the new discoveries in science, and of the changes which have been made from year to year in all branches of knowledge. ". The genus Coptocarpus Chaudoir of the Australian region with notes on Related African species (Coleoptera: Carabidae: Oodini), Smithsonian Contributions to Zoology, number 184, 25 pages, 33 figures, 1 table, 1974. The Australian New Guinean genus Coptocarpus Chaudoir has been taxonomically revised.
This led to a preliminary morphological analysis of the oodines in general and to the discovery of the apparently close relationship between certain African species and Coptocarpus species. The only practical revision of Coptocarpus, that of Sloane (1910), covers seven of the 13 species now known, provides a key for the identification of specimens, and provides brief specimen descriptions with little regard for species variation. However, it should be pointed out that Oodes, as now conceived, is a "catch-all" genus and needs to be studied from a worldwide point of view, particularly with regard to the location of the Australian species.
METHODS.—This study is the result of the examination of 117 specimens of Coptocarpus and hundreds of other Oodini specimens. Furthermore, Anatrichis has only two male probasitarsomeres with squamate setae below, while members of Oodes have three enlarged ones, but the third article of Oodes has squamate setae on fully half of the article, while members of Coptocarpus have only one third. wearing these sets. DESCRIPTION (Figure 19).-Form broad and de- TYPE- SPECIMENS.-The holotype male is depressed, with a very small head about one-third CSIRO; it was collected by P.
ETYMOLOGY.—The genitive patronymic, philipi, honors my friend and mentor who collected the types of this species, Philip J.
Coptocarpus chimbu, new species
Coptocarpus yorkensis, new species
Lettes is recognized by the almost smooth elytra and the lack of punctures on abdominal sternum II. Mesothorax: Elytral interneurons very weakly impressed, transversely striated; scutellar interneur almost obliterated; intervals flat, interval 3 with two weakly impressed setigerous pores posterior to middle; otherwise elytra as in C. ETYMOLOGY.-The adjective, yorkensis, refers to Cape York, the general area where these beetles live.
LOCAL REGISTRATIONS (Figure 18). I have seen two specimens, both from Cape York, Queensland; one from the vicinity of Tozers Gap (CSIRO) and the other from the Iron Range (MCZ). TYPE SPECIMENS. - The male holotype and allotype are in CSIRO; both were collected by P. DESCRIPTION (Figure 22). Shape very shelf and convex with a transverse pronotum, which is slightly narrower than the inflated elytra, and with a very large head, which is wider over the eyes than half the eyes. width of the pronotum.
Head: Very broad across eyes, wider than half width of pronotum; mental tooth broadly truncate, otherwise head as in C. Mesothorax: Elytral interneurs striate, striae well impressed, numbers 1, 4-7 not quite extended at basal margin; scutellar interneur one row of unconnected punctures removed from interneur 1; intervals nearly flat, interval 3 with two small compressed pores behind middle; humeral angle with well-developed, acute tooth projecting beyond border; otherwise as in C. However, all the character states studied are constant, and the male genitalia with all its intricate internal details are constant.
ETYMOLOGY - Latin adjective grossus, meaning "large", given the width of these beetles. LOCAL RECORDS (Fig. 17).—I have seen 12 specimens from the following localities: AUSTRALIA: WESTERN AUSTRALIA: Margaret River (CSIRO, MCZ, MHNP, USNM), Manjimup (MCZ), 30 miles SW Nannup (CSIRO). DESCRIPTION (Fig. 23).—Form broad and convex with a transverse pronotum equal in width to the width of the elytra, and a moderately large head slightly less than half the width of the pronotum across the eyes.
Easily recognized by slightly convex elytral intervals, continuous lateral pronotal whorl, laterally dotted prosternum, and smooth met- and mesepisterna. DESCRIPTION (Figure 24).—Form broad and moderately convex with a transverse pronotum nearly as wide as the elytra, and with a moderately large head, which is slightly narrower than half the width of the pronotum. Best known by non-punctate venter, complete pronotal lateral whorl, well-impressed striato-punctate interneurae, and flat intervals.
Coptocarpus chaudoiri Macleay
Coptocarpus gibbus Chaudoir
Coptocarpus fuscitarsis (Blanchard)
Coptocarpus championensis Chaudoir
Male genitalia with bent endophallus, left side profile; anterior endophallus hook, left oblique aspect; and apex, ventral aspect of Coptocarpus championensis, Geraldton, Australia.
Coptocarpus impar Sloane
Coptocarpus thoracicus (Castelnau)
The width and general shape of the pronotum varies in relation to the elytra, with the pronotum being narrower, sub-equivalent or wider than the elytra and the microsculpture of the elytra can be exceedingly rough or rather smooth. LOCALITY RECORD (Figure 18).—I have seen 19 specimens from the following localities: AUSTRAL-IA: WESTERN AUSTRALIA: King George Sound (MHNP); Manjimup in Karri Forest (CSIRO); Margaret River (CSIRO, MCZ, MHNP, USNM); Swan River (MHNP).
Coptocarpus australis (Dejean)
DESCRIPTION (Figure 30).—Form broad and convex with a transverse pronotum nearly as wide as the elytra, and with a large head which, over the eyes, is less than half the width of the pronotum. VARIATION.-The remarkable size difference between individuals led to confusion among the early students of the genus. Members of the other four groups are different apomorphies of shape, they are wingless, and they have some or all of the objective synapomorphies 1, 2 and 4 (see Table 1).
The members of the philipi and chimbu groups are most similar to Orthocerodus and Hoplolenus members, respectively. Perhaps with the evolution and spread of the more successful Oodes members, this primitive group of Coptocarpus-like animals was replaced in the northern Ethiopian and Oriental regions, leaving the relict groups in the southern Ethiopian and Australian regions. Due to the general research on oodine already mentioned, I do not want to recast the term "thryptocerin".
I also want to point out that if the above alternative scenario is correct, we can expect to find a relict Coptocarpus genus or two in the eastern tropics. It is very possible that further study of the Indian Holcocoleus latus LaFert6 will prove the key to this problem. Bombardier beetle reclassification and taxonomic revision of North and Central American species (Carabidae: Brachinida).
Studies of the subtribe Tachyina (Coleoptera: . Carabidae: Bembidiini), part I: A revision of the Neotropical genus Xystosomus Schaum. Studies of the subtribe Tachyina (Coleoptera: . Carabidae: Bembidiini), part II: A revision of the new world-Australian genus Pericompsus LeConte. Manuscripts for serials are accepted by the Smithsonian Institution Press, subject to actual review, only through departments of the various Smithsonian museums.
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