2A.A. Palmer, 0//r Small Eastern Strikes. [july
OUR SMALL EASTERN SHRIKES.
BY
WILLIAM
PALMER.Three
Shrikes are universally understood to occur in NorthAmerica
east ofthePlains.The
NorthernShrike{La>iiusIwfealis)^a wintervisitant in our eastern States; the
Loggerhead
(Z. ludo- vicianus), which is considered a fairlycommon
bird overmost of the region betweenMaine and
Floridaand Ohio and
Illinois to Louisiana;and
theWhite-rumped
(Z. /. exciibitoroides), which issupposed to inhabit Canada, Michigan, and westwards.
An
examination of considerable material, 176 specimens, com- pelsme
to relegate excnbitoroides to the Plains region west of the immediate Mississippiwooded
drainage area; Indovicianiis to the SouthAtlantic and Gulf coasts, and Florida, and to recognize anew
form as occupyingmuch
of theremaining region of the East.Historical Synopsis.
The name
Laniusludoviciamcswas
first givenby
Brisson^ to a birdfromthe region thenknown
asLouisiana.On
hisdescription Linnseus"-based hisbinomialname composed
of thesame
words.Vieillot^ describes two Shrikes, one Laniusborealis, the other Z. ardosiaceus,
whose
habitat he gives as Georgia, Florida, and Louisiana.Wilson
knew
buttwo Shrikes,one the northern,which he called Z. excnbitor, thinking itidentical with theEuropean
bird,and
his Laniuscarolinensis''which he found on his visit to South Carolina and Georgia.Of
this he says: "This species inhabits the rice plantations of Carolina andGeorgia, where it is protected for itsusefulness in destroying mice."
We may
be sure that Wilson'Orn.,II,1760, 162,pi. 15,fig.2.
23. N., I, 1766, 134.
30is.Am.Sept., I, 1807,Si.
^Am. Orn., Ill, 1811, 57,pi. 22, fig.5.
/oi.xvn Pat-micr, Our SiiHill Eastern Shrikes. ^d-S
either
knew
nothing of a small Shrike occurring north of the Carolinas or that he confused it with the Northern.Bonaparte seems to have had no special acquaintance with the Loggerhead. In his 'Observations on the Nomenclature of Wil- son'sOrnithology,' hestates,^ " 34. § L.carolmensis, vol.iii.,p. 57.
This species is peculiar to the southern parts of North America.
Vieillot's
name
of ardosiaceus has thepriority,and
will, therefore, be adopted."He
then gives "L. ludovicianus? Linn" as a syn-onym
of " L. ardosiaceus Vieill." In later publications he adopts Lanius liidoviciainis asthe correctname
forthe Loggerhead.Swainson evidently
knew
nothingby
practical experience of theLoggerhead
Shrike. Apparently hehad
no specimens, but used the descriptions of his predecessors in distinguishing his Laiiius cxcnbitofoidesfrom Z. ludoviciamis?-Audubon's knowledge
of these birdswas
superiorto all others but hefell far shortof the real facts.He
tells us,"The
Logger-head
Shrike is partially migratory in Carolina.A
fewmay
be found through the winter; but thenumber
is ten times greater insummer."^ He
also quotes Wilson as above.Audubon
appears neverto have suspected that his bird bred in Louisiana, for he says, "Seldom
reaching farther eastward than North Carolina, or farther inlandthan the State of Mississippi, inwhich latter, as well as in Louisiana, it appears only during the winter months.Its chief residencemay, therefore, be looked
upon
asthe Floridas, Georgia, and the Carolinas.""*He
also says: "This bird appears in Louisianaonly at intervals,and
seldom remainsmore
thana fewweeks
inDecember
or January."^The
original ofAudubon's
platewas
procured byhim
in Louisiana.Professor Baird gave the range ofthe Loggerliead as "South Atlantic
and
Gulf States,"and
of Lanius excubitoroides as"Missouri plains
and
fur countries to Pacific coast. Eastward into Wisconsin, Illinois,and Michigan
(.'').'"' This is the first'Journ.Acad. Nat. Sci. Phila., Ill, V.
-FaunaForealiAmericana,II, 1S31, 115,pi.34.
'BirdsofAmerica, Vol. IV, 1S42,p. 137.
V,r.,p. 135.
5/.<:., 136.
«B.N.A., 135S, y-i, 326.
2A.6 Palmer, Ottr Small Eastern Shrikes. |_July
appearance of this last
name
for a bird east of the Mississippi River, a practice only tooreadily followed afterwards, for its use extendedtoMaine and Canada
and even to the Carolinas. Pro- fessor Bairdhad
five specimensof ludovicianusfrom Georgiaand
an imperfect series from Wisconsin, Illinoisand
Michigan, besides anumber from
theWest,and
necessarily considered his southern specimens as distinct fromthe others.Dr. Coues, with all the
work
of his predecessors before him.and
his SouthCarolinaand
western experiences, readily fell into the view of considering theUpper
Mississippi birds as inter-mediate betweenL. ludovidamis
and
Z.excubitoroides,aconclusion which has since remained unchanged.^ Hiswork had
the effect ofbroadeningthe ornithological fieJdwork ofnumerous
observerswho
based their identifications primarilyupon
his results.Some
peculiar Shrike literature
was
thus encouraged, the effects of whichwill be noted later.The
only Shrikes mentionedby Dr.Gadow
'^that are probably from the eastern States aretwo, a '^juv. sk." from Louisiana, and an ''ad. sk." from "N. America," whichmay
be anything. All the othersmentioned are fromMexico
and Western America, yet all are placed under onename
L. ludovicianus, though it is stated thatCanadian
examples "are very distinct from the extreme southern form, which is confined to the southern Statesand Mexico
(Z. ludovicianus).''''We now come
to a phase of literature due to our increasingknowledge
ofthe range of these birds, for, as the taste for orni- thology increased through' the middleand
northeastern States, so accounts of these birdsbecame
numerous.At
first they were ascribed to the largerspecies Z. borealis,,forwe
find records of this bird)breedingin theNew
England Statesand
Pennsylvania, which were afterwardschanged
to ludovicianus or excubitoroides.Then
notices of the capture of L.excubitoroidesbecame common,
though inmany
cases awrong
identificationwas
later admittedand
changemade
to ludovicianus. But the records increased,iReytoN.A. Birds, 1872, 125; 1890,338; B.N. W., 1S74, 103; B.C. V., 1878, 563.
2Cat. BirdsBrit.Mus.,Vol. VIII, 1S83,246.
^'iScjs^] Palmer, Our SmallEaslem Shrikes.
247
both
names
being variously used, even inone case, both of them, for the birds of amated
pair! untilnow
it is considered that except in Michigan, Wisconsin, Minnesotaand
Canada, ludovi- cianusis the onlyform found.Dr.
Wheaton
^ gives Lanius ludovicianus as, a "Common
sum-mer
residentin Middle, lesscommon
in Northern and SouthernOhio
Firstascertained to occur inOhio by
myself in 1874, a female specimen, takenMay
31, 1873, O'^ whichmy
note in Coues'Birds oftheNorthwestwas
inpart based, provinga nearly typical specimen of this variety.Her mate was
an equally wellmarked
individual of var. excubitoraides."On
page 311, he says ofthis latter, "Rare
in southernand
middle Ohio, probablymore common
in northern Ohio.Summer
resident fromMarch
toSeptember. Breeds."
On
page 312 he says of excuhiforoides:"
Thus
it appears that this variety has extended its range east-ward
from the Mississippi Valley mainly along the basin of the Great Lakes."Raymond
VV. Smith,^ speakingof the birds ofWarren
County, Ohio, gives Z. ludovicianusas "uncommon summer
resident," and of L.1. excuhiforoides as " resident, probably breeds."He
also says: "The
Shrikes of this locality are just on the border line between theLoggerhead and
theWhite-rumps,and
inmany
casesit is almostimpossible todistinguish the variety."
The
A.O. U. Check-List of 1886 gavetherange of ludovicianus as "Florida, the Carolinas,and
the Gulf States east of Texas."In the list of 1895 this
became "Eastern
United States, west to the Plains; northtonorthernNew
England. Breedsfromthe Gulf States to Virginia and casually north, on the Atlantic Coast tosouthern
New
Jersey; in the interior, northward to the Great Lakes,and
through western Pennsylvaniaand New York
toNew
Hampshire,
Vermont,and Maine."
Itwould
seem
that, generally, the test of whether a particular bird belongedto eitherludovicianus or excubiioroides.^depended
on the presence of dark, orwhite(=
pale) upper tail-coverts, the1Birds ofOhio,1880, 309.
*P--dZ-
•'Journ.Am. Soc.N. Hist., 1891, 122.
248
Palmer, Our Small Eastern Shrikes.iJulv
universal ignorance of the real excubitoroides being sufficient to determine theissue. Also,identifications of excubitoroidesfromeast of the Plains have been based on breedingbirds, usually, some- times on winterspecimens taken at the most northern partoftheir winter habitat. In no instance do these
seem
to have been com- pared with typical specimens of the formwhose name
theytook.Taxonomic Differences.
Lanius ludovicianus ludovicianus
Linn.Loggerhead
Shrike.Subsfecific characters.
—
Adult$
: Above dark slaty; beneath almost immaculatewhite;bill large and stout, swollen toward tip; hook large andcoarse, gently curveddownwards;taillongerthanwing.Adult $: Similar, butsmaller. Typelocality, "Louisiana."
Lanius ludovicianus migrans,
siibsp. nov.Migrant
Shrike.Subsfecific characters.
—
Adult$
: Above bluish gray; beneath paleslaty; throat white; bill smaller, i-egularlytapering; hook delicate and sharply bent downwards; tail shorter than wing. Type, No. 163077,
$, Kingston, Ontario, April4, 1S98; Dr.C.K. Clarke.
Adult $ : Duller,especiallybeneath,and smaller.
Distribution.
From
middle Louisianaeastwardalongthe Gulf Coast and its indenta- tions;throughoutFlorida,andeastward intoNorthCarolina. Extending from thisrangetoan indeterminate distance up thevallej's, though gen- erallyconfined below the lOO-foot contour line.' Non-migratoryexcept atitsmorenorthern and itshigherhabitat....
ludovicianus.^Itwould seem desirable that thelifedistributionof forms should be con- sidered in relation to contour unes, altitudeand the influences of thevarious kindsof forest distribution being the principal factors affectingthem. The
relativequiescenthumidityof swamps anddenseforestswiththeirveryslight interactions resulting fromrapid general climate changes,plus the minimum amountof sunlight,producesresults differentfromthedryerandmore exposed elevations or depressions. Theamount of sunlight,the character of the food, andthe influences of a limited habitat seem moreimportant in theirresults thandegrees of temperature. Increase or decrease of the radiatingpowersof the ground surface destroys or drives out forms and necessarily influence thosethat remain, or replacethem,and thisradiatingpoweris determined by influences otherthanmere temperature.
°8g3 J Pai.mkk, Our Small Eastern Shrikes.
24O
Froin Maine,Vermont, andCanadatoMinnesota; southwardsintoNorth Carolina and tlieOhioVailejtothe Plains. Absent in winter from its
more northernand higherliabitatsand migrating in tlieautumntoward the Athmtic Coast and intothe Carolinas,Tennessee,and lower Missis- sippi valley. Breeding almostentirelyabovethe 500-foot contour in the valleys,casually up toabout2000 feet, and to within about 50 miles of the coastin Maine.
From
Canadaandthe edgesof the plains intergrad-ing into exciibitoroides .
...
. . migrans.From
the above it will be seen that hulovicianus is a residentof theseaward edgeofthecoastal plain, ranging up thevalleysabove the 100-foot contour in suitable places, especiallywhere civiliza- tion has prepared a way.On
the otherhand, mig7-ans is a resi-dent of the Transition
Zone
between the Carolinian and the Canadian, affected insome
places by the opening up of suitable breeding placesby
the agency of man.In
Maine
the Migrant Shrike does notseem
to beuncommon.
Mr. O.
W.
Knight has recently recorded it fromnumerous
local- ities^and
informsme
that he can always findthem
insummer.
He
issure of itsbreeding atHampden,
fivemiles south ofBangor and
about 55 miles from thesea.Most
of the records I have are breeding records, but the bird does notseem
towinter. InNew
Hampshire and Vermont
the bird is locallydistributed in suitable places, nearly all of the ten localities I have being of breeding birds. Except one breeding record in northwesternMassachu-
setts- therecords from that State
and
from Connecticut,Rhode
Islandand
New
Jerseyare either ofwintervisitantsor of migrants, though Mr. Stone mentions its probable breeding in southernNew
Jersey,^ but thismay
be the southern form. InNew
V^orkit breeds in most of the middle
and
western counties, being a migrant or winter resident in theHudson
Valleyand
onLong
Island. In Pennsylvania it breeds in the counties borderingthe western boundary, but it is only a migrant or winter visitant in the rest of the State, apparently. I do not
know
of its breeding in Maryland, though Mr.Kirkwood
informsme
that in themanu-
'Bull. Univ.Maine, 1S97.
-Auk,1SS7, iSo; Am. Nat. 1SS7,90.
^BirdsE.Penu.andN. J.,1S94, 125.
250
Palmer, OurSmall Eastern Shrikes.rtuiv
scriptsleft
by
the late E. A. Small thereis a noteof its occurrence inWashington County
insummer.
In Virginia it is a winter visitor,and
migrant over the whole tidewater region to the foot- hills of the Blue Ridge. In the valley of Virginia it is asummer
resident, wintering in mild winters in the southern portion. It
breeds
commonly
in NelsonCounty
(W. R.Robinson
^); inWar-
renCounty
(G. S. Miller, Jr.-); inRappahanock County
(Prof.F. E. L.
BeaP)
, and in FanquierCounty
(R. Ridgway^). In North Carolina it is confined insummer
to the western portion along the foothills. It is entirely absent insummer
at Raleigh, thougha winter visitor there (Brimley Bros.*), but is acommon
breeder further west at Statesville (R. B. McLaughlin*). It doubtless occurs further south, both as abreeder
and
winterresi- dent,but I have no sureinstances.From
Montreal (Wintle*) toOttawa
(Prof.Macoun
-),County
Perth (Kells), and Kingston, Ontario (Dr. C. K. Clark.*), it occurs throughmost of Michiganand
Wisconsin to Minneapolis, thence to theOhio
River,almost every record being a breeding one. It hasbeen recorded only as a migrant fromKentucky
(Fulton,Warren,and Nelson Counties).
I have seen three specimens from Tennessee
(Roane County and
Nashville), all winter birds.West
of the Mississippi itundoubtedly occursin suitable places to the edge of the Plains, but recordsare veryfew and uncertain.
The Loggerhead
is abundant throughout Floridaand
along the Atlantic Coastinto North Carolina, and probably intoVirginia on the shores of theChesapeake
region. Robinson's record for ChesterfieldCounty
^may
include this bird for thesummer
resi- dent. It occursalong the coasts ofWestern
Floridaand
of Ala-bama,
extendingup
the valleys of the latter State for a consider- able distance,probably above the 100-footcontour,WilcoxCounty
(Rev.H.
E. Wheeler"); ShelbyCounty
(C. F. Witherby");and
Butler andAntanga
Counties (Dr. D. L. Wilkinson®), I have'Inletterandspecimen.
*Inperson,
^Inperson,and Smith, Pastime,Oct., 1884, 27.
*Inletters.
^Auk, 1889,^95-
®Inletters.
^°ij^^] Palmkr, Our Siiuill Easfern S/itikes.
2^1
seen several specimens from
Hale
County, including young. Dr.Wilkinson took a set of eggs,
March
29, 1888, in Gastonbury, Wilcox County, a set of three,and
another of foureggs inHale
County,March
30, 1889.^ In Mississippiand
Louisiana it iscommon
along the coast and probably over the greater parts of these States in suitable.places along the watercourses,and
forsome
distanceup
the Mississippi Valley.The
Oxford record^ isalmost certainly this bird. It is replaced by another form in Texas.
General Differeiices.
In migrans the wing is longer than the tail, due to its migra- tory habit; in ludoviciamis the tail is longest, thus indicating its fixed habitat. In consequence the third primaryfeather of the formeris usuallythelongest,or isequaltothe fourth; in thelatter thefourth is nearly always thelongest.
The
forehead of ludovi- cianus is darklike the topof head, in migraiis it is nearly always paler. In the southern bird the underparts are usually almost immaculate, in the other the slaty of the sides of the breast extends across, especially in the breeding plumage. Usually a.fainttrace of reddish is perceptible on the breast of ludovicianics.,
butis strongerin migrans,especially in the females
and
immature..Signs of immaturity disappear quicklyin ludoviciamis, they soon
assume
adultplumage
; thereverse is true of migrans, the duller plumage,browner
primaries,and
paler edgings on the wing;covertslasting longer. Larger areas of white
marking
occur on' individuals of both formsand
are indicative of greater age butsome immature
are precocious.From
L. I. exciibitoroides,^ migrans is distinguishableby
its darker, duller plumage, especiallybeneath,by
being stouterand
longer,
and by
its largerbill, tarsiand
feet.In ludoviciamis the upper tail-coverts are almost invariably- similar tothe backin color,paleness
when
occurring being due to'Inletters.
2Ragsdale,Auk, 1SS9, 224.
^Tobe treatedinanotherpaper.
2^2
Palmkr,Our Small Easter7i Shrikes. [jiilybleaching
and
wearing. In viignins the male usually has pale upper tail-coverts, bleaching in the breeding season to a dull, dirty whitish.The
stronger, duller colors of the females rarely bleaching as much.At
theend
of the breeding season theplumage
usuallypresents a very ragged, bleached condition withall the colors very
much
faded.The
great difference, usually, between the purer colors of the malesand
the darker, duller colors of the females, the difference in sizeand
the consequent varyingamount
of bleaching of the sexes is responsible for the identification, socommon,
of excuhitoroides as an eastern bird.The plumage
is always palerwhen
fresh but soon darkens, especially in 7nigrans^ where the contrast is greater. In this also the contrast between the white throatand
the darkish breast isnearly always evident, and exceedingly rare inthe southern bird.
Literniediates arid Variations.
Specimens from Greensboro, Alabama, in the
Tombigbee
River Valley, are referable to ludoviciam/s^ but represent a tendency toward migrans.^ the bill being slenderer andmore
hooked.A
specimen from Chester, SouthCarolina, is similar, as are also two
in Mr.
Kohn's
collection from Covington, Louisiana.These
last are evidently migrantsfrom amore
northernlocalit}', asbreeding birds from tli^esame
region in.Mr.Kohn's
collection are typical ludoviciajius. In Dr. A. K. Fisher's collection are six specimens from St.Helena
Island, near Beaufort, South Carolina.These
also represent a variation in the direction of migrans.
Though
havingthe large bill,thehook
ismore
curved andlonger.Certainspecimensofmigransfrom SouthernIllinoishavestouter billsthanusual, as havealsothe birds breeding aboutMinneapolis, Minnesota.
These
evidentlyrepresentgroupsof individuals with restricted habitats in the valleys of large rivers. Their having the fourth quill longer than the third, orequal to it, would indi- cate that their migratory rangeis notextensive.Measure77ients.
Measurements
weremade
of nearlyall the specimens groupedby
States.These show
that the birds taken at the most north-^°8q8^] Palmicr, Our Small Eastern SZ/iikes. 2C"J
ern parts of the range of //ligrans are the largest.
The
averages includeallthespecimens measured, noweeding
outoftheimmature or smallest being done.The
singlemeasurements
willshow
the usual range of size in fullyadult birds.Averages of hidoTiciauus.
Of
24males,wings, 3.77 ; tails, 3.89; culmens, .60; tarsi, r.07.Of
13 females, wings, 3.69; tails, 3.79; culmens, .58; tarsi, 1.05.Averages of viigrans.
Of
35 males, wings, 3.88; tails, 3.78; culmens,.54; tarsi, 1.07.Of
24 females, wings, 3.78; tails, 3.66; culmens, .53; tarsi, 1.08.General Considerations.
Shrikes are inhabitants of open, wooded, scrubby country.
The mixed
praii-ie, savanna,open
pine woods,and hummock
lands of the southern coasts afford a congenial habitat for the Loggerhead, which is an abundant bird. Similar conditions but with a greatly different vegetation, prevail about tfie prairies of the middle States
and
the farms and open country of thesummer
habitatof theMigrant Shrike.
From
the distribution here givenit will be noticed that there is a considerable hiatus^ between the breeding ranges ofthese two forms. This is evidently caused
by
the fact that the interval between the 100-footand
the 500-foot contours is a partof the great coastal plain forest region of the south, a region unsuited to Shrikes,and
in which they do not breed. It is possible that, as civilization reachedthe prairies of Indianaand
Illinois, a passage eastwardwas
afforded by which these birds extended their range eastward intoMaine and
south-'See,Ragsdale, Auk,1889, 224-226,though hisfactswereniLxed, no hiatus reallyoccurring between ttitgransand exctibitoroides,but between theformer andliidovicianits.
254
Palmer, Our SmallEaster7i Shrikes. LjulyTAuk"^
!3
<
OVol.XV"
Pai.mek, Oi/r Siitall Eastern S//rikcx.
255
"^
H
S c/i
^
:: c/5>!i;
P
Oh^
6 ^
K O
2C6
Palmer, Our Small Eastern Shrikes.ftl*^
ward
into the Carolinas along the foothills of the mountains, the early highways of the pioneers.The many
records of theabundance now
and former rarity of Shrikesmay
be thus explained, but the facts are far too fewand
too recent to be ofmuch
value in determining such a question. Dr.Ralph
informsme
that Shrikes have penetrated into theAdirondack
region bymeans
of the roads leading to settlements located inthe densewoods
atan elevation of 2,600 feet.The
two birds are perfectly distinct and readily separable, butmay meet
in the lower Mississippi Valley,and
in places where civilization haschanged
the former natural conditions, dense forests givingway
to opencountry, old fields, bushes, etc.The
Molt
of the Adult.Adults begin to change in July or August, or later, according to
summer
habitat.Some
begin to change before migrating, while inothers it is delayed until they reachtheirwinter habitat.An
adult male that I took on Smith's Island, Virginia, in fallmigration en August30, 1895, evidently
began
to molt before itstarted, but the effects of the journeyprevented its completion, though it permitted the
new
feathers to attainfull growth.The
fourmiddle feathers of the tail are nearly full grown, while the outer four on each side are the old ones. In the wings the ter- tialsare full
grown and
new,asarealsotheinner primaries,butthe outer primariesand mostofthe secondariesareold.Very
few grow- ing feathersare tobe seen. Another taken in AlexandriaCounty, Virginia,October 3^ 1889, isin full molton thewings;thetertialsandinner primariesand
some
ofthe secondaries arefullgrown and
the others are of various lengths, the two outer old primaries being still inplace.The
tail-feathers are in various stages, the outer being the shortest, about an inch,while the central are full grown.The body plumage
is nearly complete. Another taken in Maryland,November
i, isfurther advanced. Specimens takenin January, February, and Aprjl
show some
growing featherson
thethroat,and
this seems to be the extent of the springchange in the males. I have seen no molting;females.^°8q8^1 Palmick, Our Small Eastern Shrikes.
257
Molt
of the Jiiunatiirc.When
the flightfeathers of theyoung
are fullgrown new body
feathers begin togrow
on theback and
breast.Those
of theback
are dullish slaty, faintlybarred subterminally with blackish.Beneath they are whitish, tinged \vith'slaty on the sides andvery slightly with brownishontheunderbodyexcept on thethroat,each featherbeing subterminally crossed with a faint crescentofdusky.
As
the change continuesmuch
of this barringandthe dull colors wearoff, leaving the white and slatypurer.The
last of the nest- lingplumage
to disappear on thebody
is on the pileum, upper neck,and rump.The
progress of thechange oftheflightfeathers I have been unable to determine, specimens being toofew.The
nestlingwing-coverts areretainedforalongtime
and
it isprobablethat, like the flight feathers, they do not change until the next summer's molt.
An immature
female takenNovember
11, inKing George
County,Virginia, has notchanged
its flight feathers, butmany
pinfeathers are concealed under the breast feathers and molting is evident on thethroat.A
series of ten specimens col- lected in the vicinity ofWashington, D.C,
by Mr.James
Gaut, duringthelastweek
ofMarch
of this year, are all immature birds.Except two they are in various stages of molting change, both males
and
females,new
feathers appearing on the throatand
breast, on the face, and, in a few specimens, on the head
and
back.The
wing-coverts still have theimmature
light patches at their tipsthoughvariouslyworn.No
changein theflight feathers appears, andthe primaries are usually considerably weathered.Specimens Examined. •
L.I.jnigrans.
Maine i Ohio .
Vermont
...
2 IndianaNew
York 12 IllinoisPennsylvania
....
3 WisconsinMaryland 7 Minnesota
....
17District ofColumbia . . 9 Tennessee
....
3Virginia 22 Canada 3
North Carolina . . . i
Total 104
17
258
Mearns, TtvoNexv Birds from Santa Barbara Ids. rAuk LjulyL. Indoviciantis.
SouthCaro