SAMOAN INSECTS
AKD RELATED ARTHROPODS:
CHECKLIST AND BIBLIOGRAPHY
Kaiin S. Kami and Scott E. Miller
Bishop Museum Technical Report No. IS
Honolulu, Hawaii
1998
Published by Bishop Museum Press
lS25Bemice Street Honolulu, Hawaii 96817
Copyright © Bishop Museum, 1998 All Rights Reserved Printed in the United States of America
ISBN 0-930897-74-9 ISSN 1085^55X
iDtroduetíon
This checklist and bibliography provides a preliminary synthesis of the literature on insects and related aithiopods of the Samoan Islands. It is a parallel product U) the checklist (Nishida, 1997b) and bibliogrBphy(Nishida, 1997a) of Hawaiian insects and their relatives produced by Bifiiop Museum's Ha^A^ Biological Sxnvey and it complements the catalog of Samoan oonmarine snails and slugs (Cowie, 1998). To provide a contextual fiamework for future work in Pacific entomology. Miller (1997) reviewed the general status of understanding of insects of the Pacific Islands.
Hie present list includes 2523 species of insects and 251 species of other arthropods.
Thsse numbers are probably an overestimate of the number of published species records
(because of unresolved synonymies), but certainly underestimate the total number of species present in Samoa, because many have not yet been recorded in the literature.
Many new unrepoited records are present in collections, especially at Bishop Museum (Honolulu) and Landcare Research (Auckland). Nevstheless, this list provides a foundation for future research.
This List is based on card files of Pacific insects assembled up to úie mid 1960s by Edward H. Biyan, Jr., and Amy Suehiro at the Bishop Museum and a critical review of a large amount of Uterature on Pacific entomology, supplemented by checks of major online hterature databases and hljrary catalogs. We constructed a database of aU citations to all names mentioned as occurring on any Samoan island. The checklist and bibhography presented here are summaries of thai comprehensive citation database, which we expect to both make available on Internet and deposit at American Samoa Community Collie.
We have attempted to provide the current valid usage of all the names that have been associated with Samoan insects. We have not attei^Jted to validate the original identificalions, although in some cases it is clear the names were mis^iplied. Resolution of residency status (e.g., endemic, indigenous, alien) was b^ond the scope of this project, but we hope to add residoicy status to the database in the future. We have followed the names used in the hterature in reporting species and subspecies, but it appears Üial many . of the subspecies names appUed to Samoan insects should be reviewed. Similarly, many of the generic concepts EçipHed earlier in tins century need critical review. The moths include some notable examples of monotypic endemic genera that have not been critically evaluated in the last 50 or more years.
Geogr^hic coverage includes all the Samoan iclflTid^, including Swains Island. The checklist columns for "Samoan Islands" (SI), "American Samoa" (AS), and "Westem Samoa" (WS) denote records that did not specify particular islands. We retain usage of Westem Samoa for the country now known as Samoa to avoid confusion.
Títere have been four major periods of contribution to the hterature on Samoan
arthropods. (1) The early German work, especially Rechinger (1907-1914), resulting
fit>m collections in 1905; (2) the seminal Insects of Samoa series (9 volumes, 1927-1935)
pubUshed by the British Museum (Natural History), resulting &om expeditions by the
British Museum (especially collections made by F.A. Buxton and GJi.E. Hopkiiis in
I924-I925) and Bishop Museum (E.H. Bryan, Jr. and associates); (3) the work of the Hawaiian Sugar Planters' Association and Bishop Museum in 1923,1940, and other years (Swezey, 1942); and (4) from 1974 to 1978, the United Nations (United Nations Development Programme and ihc Food and Agriculture Organization) sponsored an agricultural pest and disease survey that included Samoa, coordinated by the New Zealand Department of Scientific and Industrial Research (now LandCare Research) (Maddison, 1989). Unfortunately, most of the results of this last survey have not yet been published.
The information contained herein reflects the information available to us as of 15 December 1997. But it is an interim product, subject to continued improvement in resolution of taxoDomic identities and geogrsqihic distributions. We will appnaate notification of additions and corrections.
Acknowledgements
This project was supported by the Cooperative Research, Education, and Extension Service, United States Department of Agriculture through project number EIPM055OO, admjitistered by the American Samoa Community College, Pago Pago, American Samoa, and arranged by Don Vargo. He project also benefited fitim synea^ with the Hawaiian insect checklist thai was supported in part by the John D. and Catherine T. MacArthur Foimdation. Tlie libraries of Bishop Museum and the University of Hawaii provided most of the literature. Our colleagues at Bishop Museum helped in many ways, especiaUy Neal Evoihuis, Gordon Nishida, G. Allan Samuelson, and Sabina Swift Other specialists who contributed literature, assisted with general reviews, or resolved specific problems included William Knight, Jacqueline Nfillex, Lee Milkr, Eugene Munroe, Vojtech Novotny, Dan Polhemus, Malcolm Scoble, Mit^iael Shaffer, Roy Snelling, Timothy New, Kevin Tuck, Agnes Vargo, Don Vargo, and Maia Vaswani.
Literature Cited
Cowie, RÜ. 1998. Catalog of the norunarine fFrwil« and slugs of the Samoan Islands.
Bishop Museum Bulletin in Zoology 3: viii+122 pp.
Maddison, PA. 1989. UNDP/FAOSPEC survey of agricultural pests and diseases in the
South Pacific. Technical report volume 1. General introduction and account of the entOTnological survey. Dej^rtment of Scientific and Industrial Researdi, Auckland, NewZealand. 113 pp.
Miller, S£. 1997. Biogeograpby ofPacific insects and othCT terrestrial invertebrates: a
status report, pp. 463-475 in A. Keasi and S.E. Miller (eds.). The origin and evolution of
Pacific island biotas. New Guijiea to Eastern Polynesia: Patterns andprocesses. SFBAcademicPublidiing,Amsterdam. viii+531pp.
Nishîda, G.M. 1997a. Hawaiian tenestrial arthropod bibliography. Bishop Museum
Technical Report 11 : iii + 255 pp.|Thttp://www.bishopJiawaii.org/bishop/HBS/bibib/aithbibJïinil]
Nishida* G.M. 1997b. Hawaiian terrestrial arthropod checklisL TTiird edition. Bishop
Museian Technical Report 12: iv + 263 pp.Pittp:/Avww.bishop.hawaii.org/bisbop/HBS/arthrosearclLhtinl]
Rechinger, K. (ed.). 1907-1914. Botanische uod zoolo^sche Ergebnisse dner wissenschaitlidien Foischungsielse nach den Samoa-Ioseln, dem Neuguinea-Aidûpel und den Salomons-Inseln von Maíz bis Dezember 1905. Denksch-ifien der Kaiserlichen
Akademie der Wissenschaften, Ma^ematische-Naturwissenschi^liche Klasse 81:197--317; 84: 385-562; 85: 175-432; 88:1-65; 89:443-708; 91:139-213.
Swezey, 0 JI. 1942. Notes on food habits of Lepidoptera in Samoa. Proceedings of the
Hawaiian Enîomologicaî Society 11: 202-216.SnmnMiï by Class and Order
Group No. of Families No. of Genera No. of Species
AltACHNIDA 72 151 223
Acari , 33 <0 79
Aimfiae 33 S4 135
PseudoscoipiODida 4 5 7
Scorpiooida 2 2 2
INSECTA 2» 1379 2523
Aiiopluia 4 5 6
Coleopm 30 314 S36
CoUembola 3 S 5
DemuqXEn 5 10 12
Didyoptoa S 16 23
Dqiluiii 2 2 2
Dipttia 42 210 472
EfÂaneroptera 1 1 1
i&lmipEn 21 93 134
22 120 239
Hyme&optera 29 14« 237
isopoat 3
e
10Lc^idopleia 37 317 34«
Mallophagi 2 20 «6
Mantodea 1 3 3
NèiuujjlBid 3 9 20
Odomita 3 19 3«
Oiämpuad
«
36 61Phflsmatodca 1 2 5
8 13 26
Siphonsptera 1 3 3
3 22 32
Tlyunura 2 3 4
'níchoptoa 1 1 1
Zoiaptaa I I 1
CRUSTACEA 4 5 9
Malacostiaca
Isopoda 4 5 9
CEGCLOPODA 4 4 6
DIPLOPODA 4 9 9
SYMPHYiA 1 1 1
TAKDIGRADA 2 2 3
TOTALS 343 1551 2T74
Table of Contents ARACHNIDA
Acari 1 Aimeae 4 Pscudoscotpionida 79 Scorpionida 80 INSECTA
Anophna 4 ColeiçHEra 8 CoUonbola 24 Dermaptera 24 Dicryoptoa • 25 Diplura 2Í Díptera 2«
Ephemeroptoa 39 Heteroplera 39 Hamoptera ^3 HymcDoptem 50 Isoptera 58 Lcfâdcfilaa 58 MiUiçiaga 74 Mancodcä - 75 Neuropiara 76 Odonata 76 Orthoptera 77 Phasmatodea 79 Psocoptsra 79 Sqibon^iteia 80 Tbysanoptera 81 Thysanura 82 Trichoptera 82 ZoapBesi 82 CRUSTACEA
Is(Ç)odB • 55 CHILOPODA 8 DIPLOPODA 2«
SYMPHYLA " «1 TARDIGRADA 81 BibUography 83
SAMOAN ARTHROPOD CHECKLIST
Higher Tsxon Fanjily
Genus
Species pesprilper. Year MTM OFII TATT TliT AIIN ROS SWA UPO MNO APO NIIH SAV SI AS
. 19IS) ACAKI
Austrach j ptcri idac PaiBlaniellobatcs ceylonicus (Ouden BiBChychth oniidae
Brschychthonius pocifícus HamiDa, 1973 LJocbthODius
oceanicus Hammer, 1973 Carabodidse
Canbodes
luteoamnis Hammer. 1972 samoensis Balogh &. Balogb, 1986 CeraiomidâÊ
cranalameJlatus Hammer, 1973 AnelLozetes
discifei Hanuner, 1973 Donucolidac
Gressittolus morgiiiatus Balogb, 1970 Eremaeozetidac
octomaculatus Hammer, 1973 Ercmulidae
Eremulus cuTviseta Hammer, 1971 Eupelopjdae
Nesopelops samoacDsis Hammer, 1973 Euphthincftrídae
AustroQñtia Icbroaaeci Jacot, 1934 Rhysonicia
aiduaiC-L, Koch, 1841) Gfllumnidae
Acrogalumna venmlis (Wilhnami, 1931) Allogalumna
upolueosis Hammer. 1973 Galumna
ilabellifen Hnmncr, 1958 Neogalumna
antemûgo Hammer, 1973 PergalumDB
bryani (Jacot. 1934) Haplozrtidac
Perxylobates
venuiseta (Balogh & Matamka, 1968) Rostrozelcs
foveoLams Sellcick. 1925 Xylobaies
capuciQus (Berlesc, 1908)
Upo Upo
Upo Upo
Upo
Upo
Upo
Upo
Upo Upo Upo Upo Upo
Upo Upo Upo
MUA-Mmna. OFUOfi», TAU^au, TUT-TuDiil*, AUN=Aimnu. ROS^Rose, SWA-Swàoï. UPOUpolu. MNO-Manooo, APO-Apolima, TJÜU-NuuMk, SAV-Savùi. SI-<Kaou Islmdi, AS-Americu Samen. WS=Wc3mn Sanios
Higher Tsxon Family
Gnus
SntdaiDe^criberVear MITA OPTÍ TATl TTIT AITN ROS SWA UPO MNO APO Nini SAV SI AS W<!
Hennaimíellidae PLasmobates
acutirostmm Hamms, 1973 Upo Hydrozetidae
Hydrozetes
lamnac (de Coggi, IS99) Upo Hypocbthoniidae
Malacoai^dia
ranigeraBerlese, 1913 Upo WS Ixodidae
Amblyomma
cordifcnim Neumann, 1899 Upo SI Rhipiceptialus
s (Lamille, 1806) Upo SI Laelapidae
Aspidilaclaps
mirabilis Tragardh, 1949 Upo Coleolaclaps
sp.
Hypoaspis sp.
Laelaps
echidnioa Berlese, 1SS7 Ros Upo Duttalli Hiist, 1915 Ros Upo Liodidac
L i odes
bataviensis (Sellnick, 1925) Upo Neol iodes
sp.
Lohmanniidae AustrecBTUs
tahiticnsü (Hammer, 1972) NcotñchacBTus
phmiosus Hammer, 1973 Upo Malaconotbridac
MaJacoDotlmu
ccniums Hammer, 1973 Upo hexaseiúsus Hammet, 1972 Upo
>Jaiiliermamüidae Masthermamiia
mamillarís <Beilese, 1904) Upo Nothñdae
Nothrus
occanicus SeÜnick, 1959 Upo Oppiídae
Amerioppia
octocoma Hammer, 1973 Upo wooUeyi Hammer. 1968 Upo Macbuella
linéala Hammer, 1973 Upo Oppiella
nova (Oudemans, 1902) Upo Ranusella
cfaulmnaniensis Hammer, 1958 Upo chulumaníensis cuitipilus Hammet, 1971 Upo Onbatellidae
Oribatella
scboucedenl Balogh, 19S9 Upo
Higher Taxen Fsmib'
G«aus
Spyiies Dffcriber. Year MITA am TAU T\n ATTN ROS SWA ITPO MNO APO Nlrtl SAV Sl AS WS
Upo
Upo
Upo Onbaiidae
Eríoppim problctnuica pacifica
Balogh & Balogh. 1986 Pbtttincaridae
Hop lap hure 11 a
¿lauca Hammer, 1972 scspellou Aold, 1965 PlaticreiiLacidac
Pedroconcsclla gymndnoDis Hunmer, 1966 Quadroppüdae
Quadroppia circumita (Hammer, 1961) Sarcoptidae
SarcQptcï
scabici bominis HcriDg, 1839 Schcloribatidae
Fijibates rosttanu Hammer, 1971 Pcncheloribalcs
clzvafiis Hammer, 1973 Scmoabaies
acoüruütium Hammei, 1973 Schelonbates
ñmbríatus Thor, 1930 funtKÎBtus javensis Willmami, 1931 pracmcijns (Bcriesc, 1910) pracmcisus fijiensis Hammer, 1971 praeiiicisus inierrupCus Berlcse, 1916 sphaeroidcs Hammer, 1973 tubuaiessis Scllnick, 1959 Sbiaiobaics
tuberculams Hammer, 1973 Tuberemaeus
uideiuanis Hammer, 1973 perfon&i5 [WiUmami, 1931) thienemaum (WiUmaim, 1931) Suctobclbidac
Disco tu ctobelba varioselosa (Hsmmer, 1961) Suctobelbclla
insulana (Hammer, 1972) semiphmiosa BaJpgh & Mahunka, 1967 semiplumosa tabitiensis Hammer, 1972 Tectocepheîdac
Teciocepheus minor Berlese, 1903
«clstus (Michael, 1880) Trhypocbúi9aüdac
A&oDOümis sulcams Hammer, 1972 AlloDOthnis
Tusseolus WallwoTk. 1960 russcolus retículatus Hammer, 1972 schuiliiigi van der Hammen, 1933 Archegozetes
magnus (SeUnick, 1925) WS Trhypochthonius
CKcaivaius (Willmaiin, 1919) Upo
MUA-Manue. OFl^Ofu. TAU-Tu. TUTTutaila. AUN^Arauo. ROS-^DK, SWA-Swaini, UtO^Jpelu, MNO^faDono. APOApolima. NUU-^Uulcle.
SAV^ivaü, 5I=SvDWD Iilmdi, AS=AiiiericsD sñoa. WS=Wc9Bn Samoa
Upo Upo Upo Upo Upo Upo
Upo Upo Upo
Upo Upo Upo
Upo Upo
Upo Upo Upo
Higher Taxoo Family
GCDUS
Snecie^irtescriherYear MT!A OFIJ TAtI TtJT A1TK' RQS SWA UPO MNO APO Ntni SAV SI AS WS Uropodidae
Uropoda
samoae Hirst, 1927 Upo ANOPLURA
Haematopinidae Hannatoplous
sais adventicius Neumaim, 1912 Upo SI Hoplopleura
pacifca EwiBg, 1924 SI PedicuJidae
Pediculus
hinnaniis Linnaeus, 17SS Upo SI humanus capiiis DeGeer, 1778 Upo
Polyplacidae Polyplax
spiDulosa (Bunneistcr, 1839) Upo SI Pbtbirídae
Pbthinis
pubis (Linnaeus. 17S8) Upo SI ARANEAE
Araneidae Aoqisia
rbomboidcs L. Koch. 1867 Tut Upo Mno Sav Araneus
capitalis (L. Koch. 1871) SI spcculabundus (L. Koch, 1871) Upo
Cyclosa
littoralls (L. Koch, 1867) Tut Upo Mno Sav SI Cyrtopboia
motuccensis (Doleschali, 1857) Tut Upo Ncoscooa
maculaticeps (L. Koch, 1871) Upo
tbdsi (Walckenaer, 1841) Tut Swa Upo Mno Apo Sav Poicilopachys
australasia (GiifEth & Pidgeon, 1S33) Tut Arcbacidae
Dolichognatha
richardi (Maiples, 1955) Upo Baiychdidae
Idioctis
helva L. Koch. 1874 Upo Clubion Idas
Cbeiiacantfaiiun
fiirax L. Koch. 1873 Tut Upo
gilvum L. Koch. 1866 Upo SI voTBX Maiples, 1955 Upo
Clubiona
alveolflla L. Koch. 1873 Tut Upo Mno SI samocnsis Bcrland, 1929 Tut Upo SI upolnensis Maiples, 1964 Upo
zimmamani Maiples, 1964 Tin Oedignatha
mogamoga Maiples, 1955 Upo Corinnidae
Corinna
cetrata (E. Simon, 1888) Upo Cryptottelidae
Crypto the le
vemicosa L. Koch, 1872 SI WS
MUA-Mmus, OFU-Ofii, TAU-Tau, TUT^TtmiUa. AUN=Aunuii, ROS=*i)se. SWA^Swainj. UPOUpolu. MNO=Manono, APO=Aj»liina. NUU-^mittle, SAV'Ssvaii, Sl^Samom Islands, AS=Ainerican Sanoa, WS=We3ten] Samoa
Higher Taxon Family
Genus
Species Describer. Year MIIA OFTT TAII TITT ATTN ROS SWA ITPO MNO APO Nini SAV Desidae
Paralheuma
armaia (Marples, 1964) Swa Dictynidae
Banaidja
bi&sciaia(L. Koch, 1872) Tut Upo Sav Epeindae
Epeira
maculsDcqis L. Koch, 1871 Tut
pogisa Marples, 1957 Tut Upo Filistatidae
Wandella
bakCTi (Berland, 1938) Upo Mno Sav Gnaphosidae
Odontodrassus
javanus (Kulczynski, 1911) Upo Poecilochroa
rollini Berlaad, 1929 Mno Hahniidae
Alistra
berlandi (Maiples, 1955) Tui Upo Mno Heteropodidae
Heieropoda
nobiUs (L. Koch. 1875) Mno suspiciosa (L. Koch, 1875) Upo
venatoria (Linnaeus, 1758) Upo Sav Linypbiidae
Bffliiyphantes
paradoicus Berland, 1929 Upo Cyitaracboe
bispinosa (Keyseriing, 1865) Upo Mno vetnicosa L. Koch, 1871 Upo Linypbia
tuasivia Maiples, 1955 Upo Sav NeoDcsiotes
rcmifonnis Millidge, 1991 Tot Prise i palpus
palmarius (Maqdes, 1955) Upo Lycosidae
Lycoîa anna (Strand, 1913)
tongalabuensis (Stnod, 1911) Upo LycoîcUa
ihonlli Berland, 1929 Upo Mysmeaidae
Mysmena
phyUicola (Maiples. 1955) Upo samoensis (Maiples. 1955) Upo Mno Tamasesia
ocuminaa Marplea, 1955 Upo
iDtunda Marples, 1955 Upo Sav OcbyiDceratidae
Apiacera
minuta Marples, 1955 Upo Mno Cenileocera
lansfoTdi Maiples, 1955 S«ra Upo Mno Speocera
minuta (Marples, 1955) Up«
Higher Taxon Family
Genus
SpgcigsPwcnbCT.Ycar MUA OFTJ TAU -nrr AITN ROS SWA ITPO MNO APO Nim SAV SI AS WS Oecobiidae
Oecobius
navus Blackwall, 1859 Upo Oonopidae
loricata (L. Koch, 1S72) Ischsothyreus
lanutoo Maiples, 1955 Oonopinus
occanicus Maiples, 1955 Orch estilla
vainuia Maiples, 1955 Ptaolcidae
Alterna
maurícisna Waldcenaer, 1837 Holocneminus
maculatus Maiples, 1955 Ptaolcus
ancoralis L. Koch, 1865 sp.
Physocyclns
globosus (Taczanowski, 1873) palUdus (Blackwall, 1856) sp.
Pisauridae Dolomedes
äcetus L. Koch, 1876 Salticidac
Ascyltus
ptcrygodes (L. Koch, IS65) similis (Beny, Beally, & Proszymki, 1997) Athamas
wtiitmaei P.-Cambridge, 1877 Bavia
lon, 1877 maculatus (Keyserling, 1883) Cytaea
{ñscula (L. Kodi, 1S67) piscula subsiliens (Kulczynski, 1910) tiispiiiUcra Maiples, 1955 E&le
albobicinctus Berland, 1938 Euryattus
lautus (Keyserling, 1881) Ftacillula
minuta (Berland, 1929) Mencmenis
bivhtanis (Etafour, 1831) MoIHka
pusilta (Strand, 1913) Plcxippus
folistus (L. Koch, 1846) paykulli (Audouin, 1827) petersü (Karsch, 1878) Raiahu
nitida Berland, 1929
Upo Upo Upo Mno Apo Upo
Upo Mno Sav
Upo MBO Sav
Upo
Upo MDO Sav
Upo
Upo Mno Sav
Upo Mno Sav
Upo MDO Sav
Upo Sav
Upo Upo Upo
Mno Apo Sav
Upo Upo
Upo Sav
Upo Mno Apo
Upo
Upo Mno Sav
S>.a Upo Upo
Mso Apo
Upo Upo Mno Apo Higber Taxon
Family Genus
Sneci«DKCTbet Year MlfA OFU TAU TTTT AirN RQS SWA IIPO MNO APO Nini SAV SI AS Salticidae (CODL)
Savsiie
puDCtata Maiples, 1957 Ssv Tfaorelliola
cnsifcra (Thorell, 1877) Tin Triic
longipalpis Marplcs, 1955 ZcDodonis
micToplnhalmus (L. Koch, 1881) Tut Scyiodidae
Scyiodcs longipes Lucas. Ift45
striaüpes (L. Koch. 1872) Tktt Sicanidoe
Loxoscdes
mfcsccns (Dufour, 1820) Upo SP-
Sympbytognathidae Paiu
sastooisís Maiplcs, 1951 Upo Tetrab lanmidsc
xp. G., Maiples, 19S5 F«lli^teiiuDB
CAitaoaini (Maiplcs, 1955) samoeosis Maiples, 1964 TetiagnBlhidw
DyschiriogDBtbs nígromaculaia Bcriand, 1935 oceánica Berland, 1929 upotueiisÍ9 Marplcs, 1955 Leucauge
granúlala (Valdceaaer, 1S41) ilolelc Maiples, 1955 bfulañs (Keyserling. 1865) prodiga (L. Koch, 1871)
tuberculaia (Keyserling. 18é5) Tut Ncphila
plumipes (LatneUe. 1804) Tetragnatha
keyseriingi Simon, 1890 Isqucata L. Koch, 1871
iiiaciko& L. Kocb. 1871 Upo Mno Sav mandibulata Waldcenaer, 1841
panopca L. Koch, 1871 Upo Sav SI pTDuma WalcJcoiaer, IS4I Upo SI Tharidiídae
Argyrodes
cometes (L. Koch, 1872) Tul grecüis (L. Koch, 1872) Tut samocnsiï Cambridge, 1880
sublimis L. Kodu 1872 Coleosoma
ailamsoni (Berland, 1934) Tut floridanom (Banks, 1900)
Conopistha unimaculaia Marplcs, 1955 Cyllognattaa
aâMs Berland, 1929 Tut
MUA-Manua, OFU-Ob. TAl^Tau. TUr=TuDiili. AUN-AUDUU. RO&'Rose, SWA*S«iiiB, UfO=Upolu, MHO=MaooM>, APOApoIims, NUU^Kuntile, SAV-^mii. SI^noMa lilanth. AS-Amoicn SOIKH. WS-Wcstem Ssnn
Apo Upo MDO Apo Upo
Upo Upo
Mno
Upo Upo Upo Upo Upo Moo
Upo Upo Upo Upo Upo Upo
Mno
Upo Upo Upo Upo Upo Upo Mno
Apo Apo Upo Upo
Higher Taxen Family
Genus
SneciKñescrihB-Vear MUA OFTI TAIT TTÎT AIJN ROS SWA UPO MNO APO NTHl SAV St AS WS Tlieridiid&B (com.)
Cyllognalha
gracilis Marpici, I95S Upo Phol comma
soloa (Marplcs, 1935) Upo Phoroncidia
pcnoMia(L. Koch. 1872) Tut Upo rotunda (Keyserling, IS90) Upo Theridicm
tlbidum Berlmd, 1929 Upo SI
«IbosDiatma (L. Koch. 1867) Tui Upo
buxtoni Beiiaad, 1929 Upo Sav caM^jwsum Marples, I9SS Upo
cUhahna Marplcs, 193S Upo hopldnsi Bcrlaiid, 1929 Tul Upo manoDOCnse Marplcs, 19SS Mno inzaafe Marples, 19SS Upo samoensis Berland, 1929 Upo sctosum L. Kocb, 1872 Upo Therïtl i osomaii ilae
Tbeiidiosoma
lopdfiU; Maiples, 19S3 Tat Upo Thomisidac
Dia«
pmeiexia(L, Koch, 1865) Tut Upo Mno Apo Sav Titanoecidae
Pandava
lamínala (Tharcll. 1K7S) Upo SI Ulobotidu
Daramulunia
gibbosB (L. Kodi, 1871) Upo Mno SI
«ieUa(L. Koch. 1871) Ttit Upo Mno Sav SI Ulobonis
DKintifenis Marpl«• )9S5 Upo teauissimus L. Koch, 1872 Upo Zosis
genicuktus (Olivier, 1789) Upo Mno Sav SI Zodaiiidac
Storena
aleipau Marples, 195S Upo Mno CHILOPODA
Mecist0£Cphalidae Mec:su>cepbalui
maxillaris (Cervti^ 1919) Upo
tahitiensis Wood. 1862 Upo SI Oryidae
Orpfanaeus
brevilabiatus (Newport, 1845) Tut Upo Otostigmidae
Otonigmus
«stenus (Kohlrausch, 1881) SI Scolopendridae
Scolopendra
morsitaDS Lianacus. 17S8 Upo SI WS subspinipes Leaúi. 131S Upo SI COLEÓPTERA
Aglycyderidae Procerfainus
samóse Perkins, 1907 Upo SI
MUA-Mímia, OFUHDfii, TAU=Tau. TUT^omilí. AUN-Amuai, ROS-Roíe. SWA=Swams, UPO=<Jpolu. MKÍ>*laiiono, APOApolima. Nm>=Knnitíe, SAV-ficnü, SI='San>oan Islnds, AS-American Sanoa. WS^Wenern Same«
Higher Taxon Family
GcflUS
^neciesneatrihCT Year MlIA ORI TAU Tin- ATTN ROS SWA TIPO MNO APO Nini SAV ^ AS Anobiidae
Lasiodenoa
scTTicome (Fabricius, 1792) Tut Aiitliicidae
Antbicus
s Laferte, 1848 Upo
s Pic, 1908 SI Anthribídije
Araecenis
cuddiu Jonias. 1928 Mua Tau Tut Upo
fesciculaliis fDcGeer, 1773) Upo SI sniblevis Jonifm, 1928 Tut Upo SI vieillard! (Montrouzier. 1860) Tut Upo Sav SI CerambyrhjTichus
schoeaheri Montrouzier, 1855 Tut Upo Cisanihribus
oitîdus Zimmennan. 1941 Tut Deropygus
fomix ioTdaiu 1928 Tat Eucoryoïu
stevensi Pascoc, 1859 Upo McUnopMcus
dulcif Jordan. 1928 Upo glcnij j^dan, 192S Upo bopkins! Jordan, 1928 Upo Janus Jordan, 1928 Upo Notiana
bQxtoni JoTxlan, 1928 Upo vdlchi Jordan. 1924 Tut Upo Rhinotropis
cristiÎETiuFainnaire, 18S1 Upo Scinetious
pacificus Jordan, 1928 Upo Bostrích¡da£
Etiiyzop«rtha
dominies (Fabricius, 1792) Upo XylopsocQS
castaoopttn (Fainnaire, 1850)
reiigiosus (Boisduval, 1835) Ttat Upo Brcntidae
Chalybdicus
rcvcrens Kleine, 1928 Upo Cylas
fonnicarius (Fabricius, 1798) Tut fbrmiorius elegantuhis (Smnmcis, 1875) Cypbagogus
samoanus Kleine, 1928 Upo Bnprestidae
Agrílus
indignus FainDaire, 1849 Tul Upo samocQsis Blair, 1928 Upo CallistTQnu
samoeiuis (Saunders, 1874) Upo Chiysobothns
diiysonola (?) Deyrolle, 1864 Upo Cypbogastra
abdoininalis Wateitiouse, 188S Upo
MUA4bnu^ OFU^Ofu, TAl^Tau, TUT'TuQüla. AUN-AUDDU. ROS-Kose, SWA'SVùS, UPOUpohi. MNOMauoo, APO^líma. KUlMJuiaele, SAV-Savaü, SI'^iiDDn likndx. AS-Anxrícan Sñio*. WS-Wcsteni Samoa
Higher Ttxcm Family
Genus
Sneci«Desa-ibefY«T NflîA OFIJ TATl TtTT AlTN ROS SWA \WO MNO APO NUU SAV ST AS Callirhipidae
Callirbipis
femonia Waterbousc, 1877 Tut Upo SI Carabidae
AgoDum
bottuiophonis Redtcnbacbcr, 18^7 SI Anchomenus
coold(Slûme. 1894) SI CeUcoephes
parallelu$ Schnidt-Ooebel, IM6 Upo Sav $1 Chlaenius
flavigntlams Macleay, 182S Upo SI flaviguttaûu guttuiis Escfascholtz, 1&33 Upo
samoensis CsDä, 191S Upo Colpodes
anomalus Andrewes, 1927 Upo buxtoni Asdrcwes, 1927 Upo
boplcinsí AiKbewts, 1927 $1 laetus paciiicus Andrcwes, 1927 SI piceus Andrewes, 1927 Upo
Egadromi
smongdula (Fatncius, 1798) SI Endyoomena
pndierí (FaiimaiTt. IS49) Upo SI Gnsibaphaous
impressipennis (Castelnaii, 1868) Upo SI Mecyclothorax
ambjguus (Erichsoii, 1892) SI Mochthcnjs
tetmpilotus (Macleay, 182S) Upo SI Paxenfl
poiitisAmi (CbaDdoir, 1883) Upo nigríccps (Dcjcan, 1831) Upo Prothyma
bidemilabns Horn. 1934 SI Pse udodistypsi den
bidentilabris Horn, 1934 SI Steooioptius
dingo (CastelnBU, 1868) Upo Sav Tacfays
quÁdrillum Schaum, 1860 Sav sexgimalus (Famnaire, 1849) SI Cciambyddae
CcTcsiuni
maculaticolle (Blanchard, 1853) Tul Upo Sav ratículaium Aurívillius, 1928 TtiX Upo
unicolor (Fabndas, 1787} Tut Upo Sav SI DiboiRinus
acanthias (Pascoc, 1Í75) Upo Sav SI holotephnis (Boisduval, 1835) TW Upo
nisticaur (Faíaicius, 1801) TU Upo
VBStaiOT CNewman, 1847) S]
Glaocytcs
aiseosignalB AunvíUius, 1928 Upo Lampro^bn
sulcaa Aurívillius, 1928 Tut Upo Sav sukata fuscipennis Aurívilliiu, 1928 Tut
Leptocynicus
niädus Aurívillius, 1928 Upo
Higher Taxon Family
G«aus
Soede^rte^rih« Year MtJA ORJ TAT] TUT AlIN ROS SWA ITPO \fNO APO NTRI SAV Sl AS Cerambycidae (cool)
Obriuin
gynsndropsidis Faiimaire, I8SÛ Tut Upo SI Odontorhdbdus
dentípes Auriviilius, 1928 Tut Upo Qavicûnûs Auriviilius, 192$ Upo
rechingerí Auriviilius, 1913 Upo Sav teretiscapus Auriviilius, 1928 Upo
Olfithrius
tnsularís (Fainiuirc. 1830] Upo Sav SI subnitídus AuriviJUus. 1928 Tut
Oopsis
albopicta Auriviilius, 192S TBU Tut Upo
nutaior (Fsbrictus, 1787) Tut Upo SI obloogipamis Fainnaire, 18S0 Tau Tut Upo
obnsipvnois Auriviilius, 1928 Tut
saaigTBDOsa Fainoairc, 1879 Tut Upo striatelU Fainnaire, 1879 Upo variivestris Faiimairc, 1879 Tut
ParetiypaAÍus
bípunctaius Auriviilius, 1928 Upo flavovitlKlus Auriviilius, 1908 Upo
savaiicnsis Aurivüliu, 1928 Sav Piosoplus
saiDOBDus Auriviilius, 1913 Upo ventrales Auriviilius, 1928 Tut
Pterolopbia
latenpicia (Fainnaire, 1879) Tut Upo latcripicta conncta Auriviilius, 1928 Tut
Sciadella
albofasciata Auriviilius, 1913 Upo minua Auriviilius, 1928 Tut
vsriabüis Auriviilius, 1913 Tut Upo variabilis bUineaia Auriviilius, 1928 Tot Upo variabilis Dñgcnifen Auriviilius, 1928 Upo Cerylotiidae
Cerylon
tesaceum Fainnaire, 1850 Tut Upo Euxestus
erithacus (Chevrolat, 1863) SI Chiysomelidae
Aphtbona
vcrtchi veitchi Bryant, 1925 Upo ApriODOta
lucida Maulík, 1929 Tut Upo Ajgopístes
insularam (Msulik, 1929) Tut Aun Upo Sav Aulftcophon
abricü^BaJy. 188« SI mgrobnmna Meulik, 1929 Tut Upo Sav
(Fabricius, 1781) Mu* Tau Tut Upo S«v SI
«milis (Olivier, 1808) Tau Tut Upo Nuu Sav SI Brontispa
balaka£ Gressitt, 1957 Upo
\(Higissima (G«stTci, 1885) SI AS WS Cusida
diomma Boisduval, 183S Tut Upo Sav SI DiiTopidus
coiTu^tus Gressin, 1937 Upo
MLA-Mmua, OFUKJfu. TAU-Tai. TUT^Tumilfl. AUN-Awnm, ROS-*)«. SWA-^wains. UPO-Upolu, MNO-Manono. APOApoliíaa, NUU=Noiitele.
SAV-iSiviü, SI=Saiiio«a lilDdi. AS^AmBicon Sanaa, WS^Wcsteni Sou»
Higher Tsxon Family
Genus
Species DCTcrihcr. Year MUA OFU TAU -HTT AITN ROS SWA tjpo MNO APO NTJU SAV SI AS WS QnysDmclidac (conL)
Ditropidus
lucidus lucidus (Maulik, 1929) Upo
lucidus subpalli'dus Gnssitt, 1957 Tal Sav Pl&giodem
samoaaA Maulik, 1929 Tut Upo Plesistia
bnmnea Maulîli; 1929 Tte Upo pTomecoihfica
cdfinileipennis BlAnchard, 1833 Tut Upi> SI WS reichciBnly. JS69 Tut Upo S&v SI PycnophthBlma
tutuiUna tutuilBDB Maulik, 1929 Tut Upo tutuiUna upotuans Gressitt, 1957 Upo Khyparida
aurcDvinda Gressin, 1957 Tut
subacneicollis Feiimaire, 1879 SI Stethotes
nifonigm Maulik, 1929 Upo Sav Stygnobia
aenescens Maulik, 1929 Tui
caua Weise. 1922 Mua Tau Tin Upo Sav SI cauta aimulipcs Weise. 1922 SI minuta Maulik. 19Z9 TÚ Upo
variabüis Maulik, 1929 TU Upo Sev vetmicuücollis Gressitt, 19S7 Upo
Tracby^hthona
nigra (Maulik, 1929) Tnt Upo Sav Uroplata
güaidi Pie. 1934 Upo Mno Sav Ciidae
Cis
Upo Upo
iaphanu Zinunomon, 1941 Ta
tvtuilensis Blair, 1928 1U
Scolytocis
samoeosis Blair, 1928 I«
Qeridae Cylidnis
cyancus Falmcius, 1787 Necrobia
nifipes (DcGeer, 1775) Tm Tul Tarsostenus
univiaatus (Rossi. "W) Coccínellidac
AnisorCüS affinis Cïotch. 1874 Chiloconis
lügnois (Fabrícius, 1798) Ctulonieaes
samoeoss Airow, 1927
oapnrmvîï tntiiï1^n4ÏC Armnv IQ?*? TI«
Tin Coccindls
tnnsvcnalis Fabñctus, 1781 Tm
Coelophoni
InuquBlU (Fíbriclus, 1775) Ta
CiTptolafimu montrouaeri Mulsant, 1853
Upo Upo
MÜA-Mamia, OFU-Ofn. TAU-Toi. TirP=Tuftñla, AUN=Auiimi, ROS-Rose. SWA"Sw«ins. ÜPCKJpolu. MNO*Iinono. APOApolimfl, NUU'^uulrie, SAV<<avaii. SI-SHDDMI litudt, AS=Aiiiericaii Ssmo^ WS"WestEni Soiioa
Higher Taxon Family
GCDUS
?in«i«^<P«erih^VMr MUA OFU TAU TTTT AIÎN ROS SWA UPO MNO APO TJUTJ RAV SI AS WS Coccinellida£{coDL )
Delphasms
pusillus (Le Come, 1832) Mua AS EpilachDa
cucuibitac Richards, I9S3 Upo V ig i • I) o ctopuDctata
(Fabricius, 1775) Tul Upo Sav SI vigÍDtioctopuactata
pardalis (Boisduval, 183S) 51 Neda
tricolor (Fabricius, 1787) Upo Sav tricolor Cjiensís (Crotch, 1S74) Tau Tul SI Nephaspis
bicolor Gordon, 1982 Mua AS oculatus (Blalchley, 1917) Mua AS Nephus
fijicnsis (Sicard, 1922) Tut Upo Pscudoscymiius
anómalos Chapín, 1965 AS Rhyzobius
satcllus Blackburn, 1892 AS Rodo lia
cardinalis (Mulsant, 1850) Tut SI AS Telsimia
nítida Chapín, 1926 AS Colydiidae
Bitoma
siccaiia(P«coe, 1863) Tut SI CiCODCS
•mocnus Fainnairc, 1830 Upo Colobicuí
parílis Pascoe, I860 Upo Colydodes
3 Airow, 1927 Upo S»v
s AJTOW, 1927 Upo
Hystri cones
v^ans Anow, 1927 Upo SI Keklasmcnus
latiusculus (Faiimaire, 1881) Tut Upo Nemaudiuin
posticum Fascoc, 1863 Tnt Upo Ncoiríchus
Lañusculus (Faimiaiie, 1881) Tut Upo Ocholissa
humeralis (Fainnaire, 18S0) Tut Upo SI viduaAmrw, 1927 Tut Upo
Pycnomenis
oitidicollis (Reitter, 1877) Upo Corylophidac
Meiodenis
nicidus Matthews, 1899 Upo Saäum
angusticollc Scott, 1908 Tut Crypiophagidae
Hapalíps
samocnsis Amrw, 1927 Tut Curculionidae
Acalles
samoanus Marshall, 1931 Upo SI
MUA-Mmufl, OFUOfii, TAU=Tiii, TXTT-TuniUa. AUN-Aiauai, ROS^lose, SWA=SwaiiB, UPOUpolu, MNOManono. APOApoliroa. NUU=NiiiiIe]e, SAV=Savaii, Sl=SamoaD Islands, AS=*Amnican Ssmoa, WS=WesiERi Samoa
^gher Taxas Family
Genus
S»«i«neseriber,VMr MUA OFU TATT Tirr AIIM BOS SWA IJPO MNO APQ NTJII SAV SI AS WS Curculionidae (CODL)
AcJenemis
bicoDÍfem Faimuire, 1878 Upo eiuilai5Maisball,.193] Tut Upo dúdeos signífera Marshall, 1931 Upo
varíegata Fainnaire, 1S49 Tut Upo Sav Amblycnemus
stcvcDsoni Manh&ll. 1931 Tm Upo Aaipagia
cnbreüicollis (Foinnaire. IfiSt) Tut Up« SBV St
seraisaniralis Marshall, 1931 Upo SI Anaballus
amplicoUis Fainnaire, IS49 Upo Aphanocorynes
himcnlis Maishall, 1931 Tm Aun Upo SI sevBÜemis Manhall. 1931 Sav Ctueuctetonis
nouilae Marshall, 1931 Tut Cosmopolites
sordidus (Gcrtnar, 1424) Tut Upo SI Cossonus
dentipes Marshall, 1921 Tut Upo limbaücoUis ManbslI, 1921 Tm
plalynhinus Maisball. 1931 Upo
quaercnsManhall. 1931 Tut Upo Sav quaerens afonus Marshall, 1931 Tut
Crauopoeus
cumtusMer^iBll, 1931 Upo Deretiodes
swcsyi Mar^iaU, 1931 Tut Deretioms
gibber Maitiiail, 1931 Upo SI scutiger Maisball. 1931 Upo SI wilden Marshall. 193! Tot SI Diacbetes
bcuncniManhall. I93t Upo briger Maisball, 1951 Upo Diocalatidra
ñumcnti (Fabiirius, 1801) Tm SI tartensis (Gueriii-Meocville, 1844) Tau Tut Upo SI Dryophthonis
ansancoUis Maïahall. 1931 Sav muscAoïs Marshall, 1931 Upo
Dynaiopechus
rubronotatus Manhall, 1931 Upo Elytroteinus
subtnmcabis (Faiimaire, 1881) Upo SI AS Elytnmis
bicolor Marshall, 192] Tut Upo Sav SI bivitmus MarshAll, lt>2l Upo
samoensis MaräulL 1921 Tut SI samociisis setiventris Maisball, 1921 Tut
Eiemouyx
Tufoplagiatus Marshall, 1931 Tut
samoanus Marshall, 1931 Tut Upo Eusccpes
postíasciuus (Fainnairc, lft49) SI Gitonischius
tubicollis MaishaU. 1931 Tut
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Higher Taxon Family
Genus
SneciMDgarrib« YMT ?>^JA OFl) TAU TUT AITN ROS SWA IIPO MNO APO NlftI SAV Sï AS WS Curculionidse (CODL)
Polytus
meilerboigi (Boheman. 1838) Upo Procces
pracustum Marshall, 1931 Tul Sav Pseud o lus
longulus (Bohcman. 1SS9) Tm Upo Rbobdoscelus
obscurus (Boisduvfd, I83S) Tm Upo SI AS WS Rhadinomenis
aiomo3p6isu3 (FaiimaiTe, 1878) Tm Rhynchaenus
samoanus Manhall, 1931 Tm Sav Rhyncolojoma
subsignanuD Marshall, 1931 Upa Rhyncolus
fuscicoDu Manhall, 1931 Tut
samoanus (Mar^iall, 1921) Tut Upo Samoacis
pianatus Zimmemum, 1937 Tut Sitophîlus
oryzae (Ltanaeus, 1763) Upû SI Spanoch«lus
planirosiris Manhall, 1931 Upo SpfaacTOThÍDUs
muians Msishall, 1931 Upo puDCticolIis Maishall, 1921 Mua Dñi Tut
StenatTupis
murdmlli Zinunennan, 1938 Tut SI myristicae Manhall, 1931 Tut
Stercodcnis
binodifrons Marshall, 1921 Upo Sav Teleodâctytus
roscidus Manhall, 1931 Upo Tcmaortumpbus
samoanus Manfaall, 1921 Upo Trigonop*
spongicollis FaómBirc, 1849 Tut Upo Sav Trigonopterus
•cneoaîveus Fainnaire, 1879 Upo Sav SI bicolor MarahBll. 1921 Tut
biooMus Marshall, 1921 Tut bryaiii Maisfaall, 1931 Tut
caesipes Manball, 1931 Upo
ampes Msidiall. 1931 Upo Sav samoanus {HeUcr, 1913) Tut Upo
serraiipes MaishaJl, 1931 S»v subftietalliciu Manball, 1921 Tut Upo Sav submctallicus mcodax MatshalL, 1931 Upo Slv Dcnnestidae
vulpinus Fa^cius, 1781 Upo SI Orphinus
terminale (Sharp, 188S) Upo SI Dytiscidee
Bidcssus
curviplicanis Zhnmemian, 1927 Upo Copelatus
marginatus Shaip. 1882 SI
MUA-Mmua. OFU-Ofu. TAU-Tau, TUT^Tutuila, AUt^AuDou. ROS"Rosc, SWA=Swtiia, UPCV-X^u. MNOMaDono, APOApolona, NUlWJuidBle, SAV-Smii, SI^anMan Islands. AS'Amcrïcan SaiiKw. ^A^=WesiEin Snaoa
Higher TaxoD Family
Genus
Sp«i«DHÇrit?CT.Ygff MUA OFU TAU TTfr AIM ROS SWA ITPO MNO APQ NTJtl SAV SI AS Dytiscidae (cont)
Cy bister
tripunctatus (Olivier, 1795) Upo Hydaticus
coftsanguineus Aube, 1838 Upo Sav SI Nolomicnis
ceneUus Clark, 1863 Upo Rhantus
Bnnectcas Sharp, 1882 SI liopceroides Zimmerman, 1927 Upo
pulverosus Stephens, 1828 Sav Elatendae
A laus
samocnsis van Zwaluwenburg, 1928 Tut Upo
speciosus (Linnaeus, 1767) Upo SI CoDodenis
pallipes (Eschscholtz, 1829) Tut Upo Sav Lacon
modestus (Boisduval, 183S) Upo Lsmononia
monticola van ZwaJuwenbetg, 1928 Upo Lanelaier
fusdpes (Fabricius, 177S) Upo SI Megapenthes
VcLfrons van Zwaluwenburg, 194S Upo Melanoxanthus
comptos van Zwaluwsibog, 1928 Tut Upo SI melanocephalus (Fabricius, 1781) Upo SI Meristhus
oceanicus van Zwaluwenburg, I94S Tut Pacificóla
compia (van Zwaluwenburg, 1928) Tut Upo Propsephus
euaeniis (Schwaiz. 1901) Tut Upo
gracîlicomis van Zwaluw^iberg, 1928 Upo SI major (CsodezB, 1878) Upo SI rufipts (ScbwBiï, 1901) Tut
tongacnsis (Casdeze, 1878) Tut Upo Sav SI upoluensis van Zwaluwenburg 1928 Upo SI Simodactylus
bunoni (vu Zwaluwenbws, 192^) Tut Upo cinnamomcus (Boisduval, 1835) Tut Upo tismani Candeze, 1893 Tut
i^nui (Le Guillou, 1844) SI Endomychidae
EidoTeus
minutuf Sharp, IS8S Upo Trochoidcus
desjardinsi Gucrin-Mcscvillc, 1838 Upo SI EtDtylidae
Euxestus
basalis (Moischulsky, ISS9) Tot Upo MoTiDthftliis
samoensis (Heller, 1920) Upo Eucnemidae
Dirhagus
hopkinsi Fleuriamc. 192S Tot Upo Dromacolus
gifijudj FleutiaiK, 1928 Tut
HigticT Tsxon Family
GCDUS
Sr.erí«r>seriberV^ MITA OFTI TAU TUT AUN ROS SWA irPO MKIH APO WJll SAV SI AS WR EucDcmidae (cont)
Fornax
occanicus FlcntUux. 1925 Tut Upo lamoensis Fleuiiaux, i928 Upó unifonnis Fleuüuix, \925 Upo M&clódnis
abcmns Flcudamc, I92S Upo SI
"n. íp." Muona 1993 Upo Forraulacus
buxtoni Fleutiaux, 1928 Mua Tau Tut Upo Sav humeralis Fleutiaux, 1928 Tut Upo Sav
"n. sp.''MuoDa 1993 Upo Tacerus
upolucnsis Fleutiaux, 1928 Upo Histeridae
Carcinops
quattuordecünstriau Stephens, 1832 Upo Pachy lister
chinensis (Quensel, 1806) Upo SI WS Plaesius
javamis (Eríchson, 1834) Upo Platysoma
cavicauda Amm, 1927 Tut Upo urvillei Le Ouillcw, 1S44 Tut Upo Hydrophil idae
Dactylostcmum
abdominale (Fabricius, 1792) Tin Upo SI nibquadratum (Falnnaire, 1849) Tau Tut Upo SI EDoehnis
bi^SDí d'OrehymoQi, 1927 Sav púvulus (Kuwcrt. 1890) Upo
tñlus (Broun, 1880) Upo NoEDopagus
poÜtus dlDichymoDt, 1919 Tut SI Laemophloeidac
Laemophloeus
igDotus Kessel, 1921 Upo ovalÍ5 Grtnivelle, 1883 Tin Upo pusillus Scbonhcrr, 1S17 Upo Lucanidae
Aegus
hindenbruigi (Kñesche, 1921) SI
»ralei AiTOW, 1927 Upo tntuílensis Arrow, 1927 Tur
i^luensis AiTow, 1927 Upo WS Figulus
auñtus AjTow, 1927 Upo
samoanus Kriesche, 1921 SI WS Lycidac
Melaneros
atroviolaceus Fairmaiie, 1879 Upo Sav qaadratícollis Fainnaire, 1879 Tau Tut Upo Sav actiticollis Fainnaire, 1879 Ofii Tau Tm
muiri Blair, 1928 Tut Lyctidac
Lyctus
brunneus (Stephens, 1830) Tut
Higher Taxon Family
Genus
SpwiwPggril?cr.Yar MUA OFU TAU TUT AITN ROS RWA iim VíNO APQ >njTi SAV SI AS WS
Lyctidac (cont.) Minthca
nigicollis (Walker, 1858) T« SI M^lachüdae
Carpburoidcs
pectíoams (Shaip, 1885) SI Carp hums
flavipes Blair, 1928 Upo Mclandryidae
Biyandla
sainoensis Blair, 1928 Sav Mordcllidae
Dellamora
castanea(Boheinaii, 1858) Tau Upo Sav SI coQsîmilis (Blair, 1922) Tut Upo
samoensis (Blair, 1928) Tut Upo SI Mordella
novemguttala MoDtnnisei, 1833 Upo Mordcllistcna
buxToni Blair, 1928 Tut Upo kellaraî Blair, 1928 Tut Upo
nigrescens Blair, 1928 Tut Upo Sav swezeyi Blair, 1928 Tau Tm Upo
Mycetophagidae Litargus
vMtitus Sharp, 1879 Upo SI Propalticus
ocuUas Sharp, 1879 Upo SI Typhaca
stsKOKí (Linnaeus, 1758) Upo Nitidulidae
Carpophilus
dimidiaius (Fabricius, 1792) Tau Tul Upo SI hcmiptems (Linnaeus, 1758) Upo SI Epuraea
upolueosis Arrow, 1927 Tut Upo Haptoicus
brunneus Cillogly. 1982 Tut Upo leviganu Cillogly. 1982 Tot
luteotus (Erichscm. 1843) Tm Upo luzonensis Cillogly, 1982 Tut Upo
ocularis (Faiimairc. 1849) Tnt Sav Oedemcridac
Ananca
bicolor (Fainnaire, 1849) Tut SI decolor (Fairmairc, 1849) Tul Upo
kanack (Fainnaire, 1849) Tm Upo SI Sessiuia
livida (Fabricius, 1775) Tut Upo SI Thelyphassa
laterrtia (Fainnaire, 1849) Upo Pbalaciidae
"Gen. et sp." Arrow, 1927 Upo Platypodidac
Crossotarsus
exteroedeniatus (Fairmaire, 1850) Tin Upo SI Diapus
pusillimus Chapuis, 1866 Upo quinquespinatus Chapuis, 1866 Upo
Higher Taxon.
Family GCDUJ
Sneci«D«cril»T.Year MTJA OFII TAU TTJT ATIN ROS SWA IJPO MNO APO T4UU SAV 53 AS WS Platypodidae (CODL)
Piatypus
tetracenu Beeson, 1929 Upo Prtistomidse
Prosiomis
SBinoensis Airow, 1927 Upo Rhipipbonda«
Miciopel ecotûmoides
fulvoserícans (Fuimoire, 1878) Tul Upo SI Salpingidoie
Inopeplus
metallescens Fainnairc, 1S81 Upo SI Scaphidiidae
Scaphisoma
tridematum (Lobl, 1973) SI Scaiabaeidae
Adoretus
versutus Harold, 1869 Upo SI WS Aphodius
lividus (Olivier, 1789) Upo Sav Ataenius
orbiculañs Sdunidt, 1913 Upo Coprií
iaccTtus prociduus Say, 1835 SI WS Oryctes
rbinoceros (Liimacus, 1758) Tut Upo S&v SI AS WS OxycAoRÍa
versicolor (Fabrícíus. 1775) Upo WS Phaeochjous
ooargiiiatus Castlenau, 1840 Upo Tnchiorrtiysa eanus
hirsudis (C)oun, 1901) Tut Upo Sav Scolytidae
Caiposinus
brcvior (Eggos. 1927) Upo granulifer Befsan, 1929 Tut Upo Coc«otryp«s
adveuaBlaiidfonl, 1894 Tut Upo SI caipopbagus (Homung, 1S42) Upo
cypeñ (Beeson, 1929) Upo SI médius ÇEggfSi, 1929) SI myiisticae (Roeplce, 1919) Tut Upo SI tapatapaoensis (SchetU. 19SI) Upo
tumiloisis (Beeson. 1929) Tut Upo SI vulgaiis (Eggas, 1923) Tut Upo SI Cryphalamorphus
samomus Browne, 1977 Upo Cryphalus
basihÎTtus Bceson. 1929 Upo molUi Schedl. 1955 Tul Upo
"n, 3p." Beeson. 1929 Tut
pexus Schedl 1979 Tut Upo WS samoensis Beeson, 1929 Tut Upo
silvaous Schedl. 1951 Tut Upo SI sylvicola CPeridns, 1900) Tut Upo WS upolucnñs Schedl, 1951 Tut Upo SI Cynogenius
brcvior (Eggen. 1923) Upo SI gnnulîfer (Beeson, 1927) SI
•fUA-Mnuia. OFt^^fii, TAl>*Tsu, TUTTuuila, AUN^Aianu, ROS^-Rose, SWA^^wiin^ UPCMJpolu. MNOManooo, APO-Apolbiia. NUl>>NiiiilEle, AV=Ssvwj, SI*SaiDinn lilnds, AS=Aneric>D Ssnoa. WS=Wesitin Ssnoi
Higber TtxoD Family
Genus
Species Descnber. Year- MIJA QFU TAU TIJT AIJN ROS SWA T7PO MNO APO MTTJ SAV SI Scolytídw (oint.)
CyiTOBcnius
samoanus (Eggeis, 1928} Tut Upo SI Eccoptopicnu
eccoptûptenis(Schedl, 1931) Tut Upo SI Eidophetus
samoanus SchedI, 1972 Upo SI Eriayphalus
lylvicala (Pcitins. 1900) SI Ficicis
paciiicus (BecioD. 1929) S«v SI parcani! (Cbapuis, 1869) Upo SI HylesÎDUs
samoaniuScbedL, 1931 Tut Upo SI subcostanis Eggen, 1923 Tut Upo SI Hypoayphalus
afiamaius Schcdl, 1951 Upo SI basihirnu Beeson, 1929 TW Upo SI
¡milans Schcdl, 1931 Tm Upo SI mangiferae (Sicbbing, 1914) Upo SI polyneuae Schedl. 1979 Tut
tutuiUessis Schedl, 19SI Tot SI Hypothenemiu
arcccae (Homung, 1842) Tut Upo birmaiius (Eichboft 1878) Tut Upo cassavBcnsis Schedl, 1938 Tat Upo
cunipennis (Schedl, 1930) SI enidicus (Wcstwood, 1833) Tut Upo
muttipunaatus (Schedl, 1939) SI Ozodendron
grandis (Beeson, 1929) Upo Poccilips
cypcri (Beeson. 1929) Upo longicoUis Eggers. 1927 Tul
myrisiicae Rocplce. 1919 Tut Upo sciosus [BeesoD. 1929) Tut
vulgoris (Eggos. 1923) Tu Upo Ptilopodinï
maïquesDUS Beeson, 1933 Tut Upo ^ zinimemani Schedl, 1951 SI Scolytomimus
maculatus Beeson, 1929 Ttit Upo Xyleborinuï
Bitelineatus (BeesoD, 1929) TUt Upo SI XyleboTus
tSisisEichho^ 1868 Tut Upo baculnm Beeson, 1929 Upo
buxtoni Betson, 1929 Upo S«v SI CQQsulam Schedl. 1953 SI destniens Blaodfoid, 1896 Upo Sav SI fallaxoides Scbedl, 1953 Upo SI Cemigiceus (Fabricius, 1801) TU Upo Sav SI longicollis Browne, 1977 Upo
mêlas Eggas, 1927 Tut SI mustus Schfîdl, 1972 Upo SI peicxiguus Schedl, 1971 Upo
pcrfbnas (Wollaston, 1S57) Tot Upo Sav SI piccus (Motschulsky, 1863) Tot Upo Sav SI rameus ScbMS. 1940 Tvi Upo
similis Fcirarí, ]S£7 Upo SI
[igher Taxon Family
Genus
SwcicsPcscriKr.Ygar MIJA OKJ TAU TUT AUN ROS SWA UPO MNO APO NUU SAV ST AS WS Scolytidae (conL)
Xylebonis
swezeyi Beeson, 1929 Tut Upo
tapatapaoensts Schedl, 1951 Upo SI lorquanis Eichhoff, 1868 Tut Upo Sav unimodus Beeson, 1929 Tut SI wilderi Beeson, 1929 Tui Upo
Xylosandnis
compactus (Eicfahofi; 1875) Upo
crassiusculus (Motschulsky, 1866) SI morigai]S(BlandfoTd:, 1894) Tut Upo SI Silvauidae
Cryptaniorpha
desjardinsi (Guerin-Mcnevillc, 1844) Tut Upo Sav Monaous
concinnulus (Walker, 1858) Upo
biapicalis AITDW, 1927 Tut Upo
crudgcr (Watertiousc, 1876) Tut
iainnaiiciGrouvelle, 1913 S[
obscuras Airow, 1927 Upo Sav paUidipennis (Blackburn, 1885) Upo
Shoguna
tennitifcffinis (Fairmairc, 1883) Upo Silvaous
unidentatus (Fabricius, 1792) Upo Siapbylinidac
Aleochara
nigra Ktaaiz, 1859 Upo Br^othmusa
samoensis Pace, 1984 Upo Cañus
nauticus (Faiimaiie, 1849) Upo CocDonica
buxtOQJ Cameron, 1927 Upo Coproponis
formosae Bemhauer, 1922 Tut Upo Eulissus
holomelas Perroud, 1864 Upo SI Gnypcta
vaii^aia Bcrnbauer, 1902 SI Gyrophaena
albidicomis Bemhauer, 1913 Upo Holocor^nus
Itmgiceps Cameron, 1927 Upo Homalota
variivcmris Kraalz, 1859 Tut Upo Leptacinus
pallidus Cameron, 1927 Tut Lispinus
castancus Fauvcl, 1878 Tut Upo impressicollis Motschulsky, 1857 Upo Lithocharïs
vilis Kiaatz, 1859 Upo Mcdon
tutuilanus Cameron, 1927 Tut Oligota
chrysopyga Kraaiz, 1859 Upo semirafe Cameron, 1927 Upo
fUA-MBDua, OFU-Ofo. TAU-Tau. TTJT-Tuniila, AUN^Aimuu. ROS=Rose. SWA=SwBÍns, UPO=Upolu. MNO-Maoono, AFOApolima. NUU-Nmi AV=Savaü, SI^^amoBii Islands, AS^American Samoa, WS=Wcsteiii Saiioa
Kigber T&XOD
Funity Genus
Species DescriheT. Year MllA OFIJ TAP TTIT AIJN RQS SWA TIPO MTJO APO Nim SAV SI AS WS Staphyliiùdse (com.)
Osoríus
smnoemis Bemhaucr, 1922 Tut Upo Sav Oxytclui
"sp. a." Cameron, 1927 Sav Palsminus
samoensis Cameron, 1977 Sav
"sp. n." Cameron, 1927 Upo Priochims
fofticomis (Fauvel, 1878) Upo Sav SI frcminvillci Le GuiUon, 1841 Upo S«v samoensis (Blanchard, 1853) Upo Sav SI Tactayusa
insulana (Faiimain, 1849) Tut Upo Thyrco cepfa alus
taiticDsis (Boheman, ISSS] Tot Upo Trogopblocus
siaiaeiisis Fauvel, 18S6 Upo Tencbrioflítlae
Alphiiobius
diaperinus (Panzer, 1796) Upo laevjgaois (Fabnciiu, 1781) Tot Upo piceus Olivier, 1792 Upo AmarygiBUî
tubcrea]igeifaiimaire, 1849 Tut Upo Sav SI Apteromerus
convexus (Faiimaire, 1849) Tau Tut Upo Bndymcnia
amiconnn (Fainnaire, I84S) Tau Tut Upo SI craMtui (Pascoe, 1870) Tut Upo SI granaiicollis Faiimaire, 1883 Upo SI lobicoUis Gebien. 1920 Tut Upo SI pectinatiis Kaszab, 1933 Upo
planicollis planicollis (Fainnaiie, 1S49) Tan Tut Upo Sav SI planicoUis samoana Kaszab, 19SS Tut Upo Sav Chariotheca
samoensis Blair, 1928 Sav SI sulcipomis Blair, 192S Upo Sav SI Dioedus
swalei (Blair, 1928) Upo SI (hiathocenis
comutua (Fabricîus. 1798) Upo SI Latbeticus
orizae Watcriiouse, 1880 SI Leiochrodcs
sumralis (Westwood, 1SS3) SI Lyphia
ínstríflia Pic, 1924 Tal SI Matendris
acnea Hug-Riitenberg, 1878 Tut Upo SI blsirí Kaszsb. 19SS Upo SI Menimus
samoenns Blair, 192& Upo SI Mesomorphus
villiger (Blanchard. 1853) Tut Upo Sav SI Palorinus
bumcralis (Gebieo, 1914) Tul SI PaJorm
aunrímis Champion, 1896 Upo SI i^luensis Blair, 1928 Upo
MUA-Manna. OFU-Ofij. TAl>'Ta^, TUT^TnOiila, AUN-Aiiomi. kOS^tou, SWA«Swiiní, ÜPOUpolu. MNOManono, APO^ApoliniB. NUU^Nuniíle.
SAV-SBVü. S=5amo«D Ulmds, AS-Amerícan sános, WS-Wenem Stnoa
ligber Taxon Family
OCDUS
Sneci« Dc^-h« Year MTIA OFTJ TAU TTrT ATTN ROS SWA UPO MNO APO NUU SAV SI AS WS Tenebríonidae (cent.)
Palonis
znnmennani Kaszab, 1955 Upo Parahyocis
cheoipioni (Fauvel, 1904) Upo SI Platolcncs
samoeosis (HAag-Ruteoberg, t$7$) Ten Tut Upo Sav SI Promechis
sulcjgcn (Boisduval, 183S) SI Rhipidaudnis
eioides clypeims K^szab, 1953 Tut Upo SI Sciopbagus
pandanicola (Boisduv&l, L83S) Tan Tui Up« SI Siroagyliuni
swczeyi Kflszab, 19SS Upo Szekessya
hypopbloeoidcs Kaszsb, 1955 Upo SI Thesilea
puncticeps Funoaire, 1881 Tut Upo Sav SI Tribolium
castaneum (Herbst. 1797) Tut Upo SI Uloma
cavicoUis Fainnaire, 1$49 Mm Tut Upo SI maddisoni Kasiab, 1985 Tut Upo
Throscidae Diapetes
samoensis Blaii, 1929 Upo Trogositidae
Tcncbroidcs
mauritanicus (Linnaeus, 175$) Upo SI njordax Sharp. 1891 Upo
:OLLEMBOLA Entomobryidae
Creiaastocqihalus
cdebouii Schaficr, 1898 Upo H«teromurus
letracsnúius Bomcr, 1906 SI Lcpidocynus
médius Schaffn. 1898 Upo Hy pogastTurí dse
Xcnylla
ociooculaia CsrpcntcT, 1928 Upo Onychiuridae
OnychiujTJî
fimetañm (Linnaeus, 1767) Upo lERMAPTERA
Carcinoptiarídae Euborellia
aimuLipes (Lucas. 1847) Upo SKV
pleteja CDiaaa, 1863) Upo Cbelisocbidac
Cbelisocbes
lilyanus Holdhaiu, 1908 Upo
mono (Fabriciua, 1775) Tut Upo Sav SI WS aigroniñu (Buir, 1902)
ForñculidM Elaunon
bipaninis OÜiby, 1891)
Higher Taxon Family
Genus
Sped«De^hEf Year MTIA OFTl TAi) TITT AITN ROS SWA TIPO MNO APO NTHI SAV 5;i Labiduridac
Labidun
riparia (Pallas, 1773) Upo Labiidae
Marava
arachidis (Ycrsin, 1860) TK Upo Sav
NuogasUr
>£uleatu£ (Bonnans, 1900) Tut Upo SI Ptielabtllä
curvicauda (Motscbulslcy, 1863) Tut Upo SI Spbingoiabis
bawùieosis (Barmans, 1882) Upo Sav 51 Spirolabia
pílieoiBÍs (Motschulsky, 1863) Upo SI DICTYOPTÎRA
Blaberidae Diploptera
punctao. (Escfaschohz, 1822) Tau Upo Sav Pyawscdus
indÍCTij (Fabricius, 1775) Tul Upo SI Blatteltidi£
Balta
DotulatzfStal, 1861) TU Upo $sv SI obtmoU (Bnumer, 189S) SI similis (Ssiume, 1869) TUt Upo
unicolor Cbopard, 1929 Upo Blandía
gennaoica (Liimaeus, 1767) Tm Upo SI sp. SI LobopUTclla
dimidistipcs (Bolivar, 1890) Upo LuppariA
lau (Chopaid, 1929) Sav Mcganarcia
ä£ci£rDns(Chopaitl, 1929) Tut Upo Sav BIsRídac
Cosmozosiena
bicolor (Saussure, 1864) SI Dorylaca
fiavidacta (Haan, 1842) Upo Sav rotoodaa (Bnuiner, 1865) Sav Melanozosteria
iiJiú]B(Bnmner, 1865) Tut Upo Sav SI Feri planeta
unerícana (Linnaeus. 1758) Upo SI australasíac (Fabricius, 1775) Tul Upo Sav SI bnnmea Bunocister, 1S3S SI Plicyzostcña
sóror (Bnumer, 1865) Upo SI Tdofiopteiyx
obscura Saiissuit, 1869 SI ThegsoDptejyx
bninnea Cbopxrd, 1929 Sav Euthyrrhaphidae
HoJocompsa
nitidula (Fabricius, 1781) Upo Panesihiidae
Ancaudellia
semüssima (Brunner, 1865) Upo Sav SI
Hitler Taxon Family
GCDQS
SpeciesDescriberVair MITA OFU TAIl TUT AIJN ROS SWA UPO MNO APO NTTIT SAV SI AS WS DIPLOPODA
Cambalopsidae Glyphiulus
granulatus (Gervais, 1S47) Tm SI Hypocambala
anguiaus (Attcms, 1900) Upo Pachybolidae
Leplogoniulus
naiesi (Pocock, 1893) Upo WS Tñgoniulus
Uanbridnus (Geistaeckcr, 1873) Upo WS Paradoxosomati dàe
Asiomoipha
Goarctaia (Saussure, 1859) Upo Sav WS Desmoxytes
daros (Chamberlbi. 1920) Upo SI Ortbomoipha
gnnosa AOems, 1920 SI Oxidus
giacüis (C. L. Kodi, 1847) SI Pseudospirobolellidae
Pseudospiroboleltus
bulbifenis (Ancms, )903) Upo WS DIPLURA
Japygidae gen.
sp. Upo ParBjapygidae
Pantjapyx
saraoaous Silvestrí. 1930 Tut DíPTERA
A^romyzidftc Agroinyza
sp- Tut Sav Ccrodontha
püiictt (Becker, 1$0Î) Upo WS Melanignniiyza
«Ibisquoma (MaUoch. 1927) Upo Sov bryani Sasakawa. 1963 Tm SI ip<nnoeavoTa Spencer, 1963 Upo
sojae (Zdmtner, 1900) TU Upo spenceri Sasakawa, 1963 Tut
spcrroboli Sasakaw«. 1963 Tiit Ophiomyia
COTniiia(dc Meijerr, 1910) Ofii Tm Swa Sav iBntanae (FroggaR. 1919) Upo
irfiaseoli (Tiyon. 1895) Upo SI PseudoDapomyza
dilatata Sasakawa. 1963 SI spiua (MaUoch, 1914) Tvt Swa Upo SI spÍDoa Spencer, 1973 AS Tropicomyia
momordicae (Sasakawa, 1963) Tut AnJsopodidae
Mesochria
buxtooiaiía Edwards, 1928 Upo Asílídae
Atonía ?
sp. Upo
MUA^iÄÄina. OFU-Ofti, TAIHÍBJ, TUT-Tiiniila. AXm-Auonu. ROS-Rose, SWA-SwaiiB, UPOUpolu, MNOManoao, APO=Apolimí, NUÜ=Niintde;
SAV^Snraii, Sl-^amoan lilnd^ AS-Anuricm sñioa, WS^Wcaero Samen
Higher Taxon Fnnity
Oenus
SpeciesDe^brrYMT MIÍA OFIT TAU TITT A1TM RDS SWA tlFO MNQ APO NUU SAV SI AS W<:
Asilidae (coat.) LeptogostCT
Javaneosis ? de Meijcrc, 1914 Upo Sav Maira
aeaea(F&bricius, 1805) Mua Tul Upo Sav sodalis (Walker, 1838) SI varions Ricardo, 1929 T« Upo Sav Ommáous
vuitibinus (Ricardo, 1929) Sov Pnnnachus
graeffi SchinelB. 18M SI Calliphoridae
CalliphoFB
leucosticta Bczzi, 1927 Upo Sav SI Chiysomya
megacephala (Fabricius, 1794) Tut Upo
lufi&cies (Macquan, 1843) Upo SI
¡TI- SI Hemipyrelli&
ifaodocen BczZL. 1927 Ite Upo Sav Melinda
auriceps {Mallûch, 1930) SI biseiosa (Bczzi, 1927) Upo Sav SI xipbopborB(Bcm, 1927) Sav Cecidamyiiâae
Allabrêmia
upalui Bames, 1928 Upo Sav AAArete
pallida Edwards, 1928 Tul Anaretella
nitida (Edwanls, 1928) Upo Lestodiplosis
sp. Upo Liebeliota
biñircala Banes, 1928 Upo PonicoQíiyla
ouricuUta (Bames. 1928) Upo Cerno pogoQidac
AtríchopogoD
abrasus Edwaids, 1928 Sav aiTOScutcUaus Edwai^ 1928 Sav SI jacobsoDÍ (de Meijere, 1907) Tut Upo SI Bezia
sexspiíLosa Edvranls, 1928 Tau Tut Sav Bracbypogon
pengrinaiOT (Edwards, 1928) Sav Culicoides
bcUmi Wiith &. Amaud, 1969 Tut
infuiaaus ^4acfîe, 1933 Tut Upo mollis Edwards. 1928 Tut Upo polynesise Winh & Arnaud, 1969 Tut
saEXKasis Wirth & Amaud, 1969 Tui Upo Dasytaelea
carolinensis Tokunaga, 1941 Tni radialis Edwanls, 1928
sp. Tw Forcipomyia
boibonica Ciastrier, 1959 AS carolmensis (Tokunaga, 1940) AS cûinis lohannses, 1932 SI MUA'*linuH. OPUOfii. TAU-Tüi, TUT-Toùrik. AUN=Awmu, ROS-*««. SWA-^ïiins. UPO-Upolu. MNC^Maoono. AK^Apoiima. NUU-Nuiffile.
SAV>Savaii. Sl-^nwaa UlBds, AS*Amaicm Siiiioi. wS=Wa>en Samoa
Upo
Higher Taxon Family
GCDUS
Sped« nesenher. Year \niA OFII TATI TUT ATTN ROS <!WA UPO MNO APO TJIÏÏI SAV SI AS WS Ceraiopogonidoc (cont.)
Forcipomyia
fuliginosa (Mdgen, IS18] Upo Sav SI giobulsris Edwards, 1928 Upo
ingnmi Carter, 1919 Upo Sav ST pecDDunguis (de Meiere, 1923) SI puDctipes Edwards, 1928 Upo SI jamocDsis (Edwutls. 1928) Sav SI saidcri Kieffcr, 1912 W$
iwezeyi Edwards, 1928 Tut Stilobezxia
samcaae Edwards, 1928 Upa Sav Chironomidac
Ablabesmyia
atrom»:ulaia Edwards, 1928 Upo
hilli Fieanoo, 1961 WS Dotabilis (Skuse, 1889) UpD
Chiionomus
samocnsis Edwards, 1928 Tot Upo SI spp. SI OûTODomus (Polypcdjlum)
Chinnomus (s. str.)
Upo Upo ClUDÍO
paciñcus Edwards, 192ti Tut Upo Culícoides
sp. Upo DicTotendipes
cmdidibflsis Edwards, 1924 Upo Polyp edilum
sp.
Pontomyia
natans Edwards. I92ti Upo Pseudosmittia
tnaoobianchia (Edwanls. 1928) Tanyístsus
halopbilae Edwards, 1926
hoplcúui Edwards, 1928 Upo upolucnsis Ashc, 1985 Upo Thalassoinya
maritima Wirtii, 1M7
pilipes Edwards, 1933 Tvt Chloropidae
Apotropioa
puipuiasce&s (Malloch. 1930) Upo Cadrrma
paluda (Loew, 1866) Tsu Tut Upo samoaensis Malloch. 1930
CbloTúps
minutula Malloch. 1930 Upo Diplotoxa
polira Malloch. 1930 TU Gamax
agUis (Malloch. 1930) bryaní (Malloch. ]930) MeiJCTtJla
inaequalis (Becker, 1911)
MU/t*temifl, OFU-Ofu. TAU=Tau. 71/1^1101118. AUN-Aunuu. ROS-Rose, SWA-Swams, UPO^ípolu. MNO=Maiiínio, APOApoKraa, NUU-NuuIde.
SAV-Ssvaii, Sl-Semoan Uands, AS=Aaaian sñnoa. WS-Wstera Samoa
Higher Tsxon Family
Genus
SneciMPe^heiYMr MTIA OF11 TMJ TUT AUN Tt• SWA UPO \ÍNO APO Wni SAV SI AS W<i Clusiídoe
HeteromcríngÍB
polynesicnsis Sasakawa, 1966 Tin Culicidáe
Aedes
aegypti (Linnaeus, 1762) Upo SI WS kochi (Donitz. 1901) Upo
Doaumus (Theobald, 1903) Upo
Dceanicuâ Beltis. 1962 AS WS polynesicMis Martes, 1951 Upo S( AS WS pscudoscuteüaris CTIicobald, 1910) Upo SI samoanus (Gnmberg, 1913) SI AS WS scmeUaris (Walker, 1858) SI sp. Tut
tutuiloe Ramalingam & Belkin, 196S AS WS upoleniis Maiks, 19S7 AS WS vexans (Meigen. 1830) Upo Sav SI Aedes (Aedimoiphus)
sp. SI Aedes (Fialaya)
sp. SI Aedes (Stegomyia)
sp. SI CoquillCTtvdia
fijiensis (Belkins. 1962) WS?
samïwcQsis Siose, 1966 AS WS Culex
annolirostris Stuse, 1889 Tut Upo Sav SI quinquefascistus Say, 1823 Tut Upo Sav SI W5
£amiMDSi5i5 (Theobald, 1914) Upo SI sitieos Wiedemann, 1828 TiM Upo
Culex (Coloi)
^. SI Mansoak
sp. $1 Toxortiynchites
amtMiDcssis (Doleschall, 1SS7) AS W5 brevipalpis Theobald, 1901 SI sp. Tm
splendens CWiedemami, 1819) Tttt Tríptcroides
puipuratus (Edwards, 1921) AS?
Dolichopodidae Amblypsilopus
SOTO-(Parent, 1934) SI Cfaiysosama
uBDpliaaim Becker, 1922 Swa Upo AS WS Enile Lamb, 1929 Tau Tut Upo
" ). J9» TM Tut Upo Sav
inriatiiTii Lamb, 1929 Upo Sav lactdmicms Becker, 1923 Upo Mno Sav
leucopogon (Wicdemann, 1824) WS parvicumllatum Lamb, 1929 Upo Sav patelliferum (Thamson, 1869) SI viduum haab, 1929 Upo
Chrysotus
spp. SI Ciypiophleps
aotoaaais Bickcl, 1996 Upo
MUA=Mantt4. OFUK*i. TAU=Taii, TV>Tunùla. AUN-Aumai, ROS-Rose, SWA=S\wiBí, UPOUpoIu. MNO=Míoono. APO-Apolima. NUU=>íwnrle.
SAV^avBJ, SI=SamM)) ülmiOi. AS-American Samoa, WS-Wesnn Saoooa
Higher TaxMi Family
Genus
Soedes De«:riber. V^ MUA OFTI TAU TUT ATTN RQS SWA IJPO MNO AVCi NIHT SAV ST AS WS Dolicbopodidae <COUL)
Cynaiopus
spinosus (Parent. 1934) SI Helixocems
meadosum Lamb, 1929 Tot Upo Medciera
griscsccDS de Mcijerc, 1916 Upo
saJomonis (Parent, 1941) AS Syrnpycnus
sp. SI Drosophilidae
Dettopsomyia
fbnnosa Lamb, 1914 Tut
sp. SI Drosopbila
albiÊQiiUta Malice 1934
•nanassae Dolcschall, 1858 Tut hipectînau Duda. 1923 Tut birani Mallocfa, 1934 Tau Tul convexa MaUoch. 1934
excepa Mallocfa, 1934 Tut ílavipcs (Hamsoo. 1954)
ñiscovñtata Hanism. 1954 Tut bypopygialis Mallocb, 1934
innocua (Malloch, 1934) Tui kikkawai Burla, 1954
manonoensis Harrison, 1954 Tut marjoryae Hamsoo, 1954
mdaDogBStcr MdgeD, 1S30 Tnl mcrcamruni PaiicTSOD EL Wheeler, 1942 Tut mimtium de Meijerc, 1916
nannaLamb, 1914 Tut nasutoides OkadM, 1964 Tut nigrifrons Malloch, 1934
novida Whecler ft Takada. 1964 Tni nublada Wbeeler & Xambysellis, 1966 Tm paludosa Bock & Wbeeler, 1972 Tm pleuTovittaia Harrison, 1954
polychaeia PaOersoD & Wheels, 1942 Tat samoaensis Hamson, 1954 Tut
semioigra Duda, 1926 Upo
«mulaiis Sturtevant, 1919 Tut nramincipes (Malloch, 1934) Tul niblincata Duda, 1926
Rilñuigaster (Dnda, 1923)
unicolonta Wheeler, 1959 Tot Upo upoluae Malloch, 1934 Upv Drosophila (Drosopbila)
spp.
Drosophila (Hinodiosophila) spp.
Drosophila (Scaptodrosopbila) spp.
Drosophila (Sopbophora) Leucopbenga
^.
samokouis Hanison, 1954
Upo
Upo kin« Apo Sav Upo
Upo Mno Apo Sav Upo
Upo Upo Upo Moo Upo Moo
Upo Sav
Upo Upo Mno
Mno Upo Upo
Upo Mao Sav
Upo Sav
Upo Mno Sav
Up«
Upo Sav
Upo
Higher Taxon Family
Genus
Sned«rte^-WV^ MtlA OF» TAU TUT AITN ROS SWA TIPO \ÍNO APO NTHI SAV SI AS WS Clusttdae
Hcnronefingift
polynesiensis Sasakswo, 1966 Tut Culicidae
Ac des
aegypti (Linnaeus, 1762) Upo SI WS koclii(I>crRÍtz. 1901) Upo
níxaumus (Theobald, 1903) Upo
occanicus Belkin. 1962 AS WS polynesiensis Mario, 1951 Upo SI AS WS puudosoncllam (Tlieobald, 1910] Upo SI samoenus (Gnniberg, 1913) SI AS WS scutcllaiis (Walker, 1858) SI
$p. Tul
tutuilae Ramalingam ¿t Belkin, 1965 AS WS upolensis Maiks, l957 AS WS vczaiu (Mägen. 1830) Upo Sav SI Aedes (Aedimoiphus)
sp. SI Aedes (Finlaya)
sp. SI Aedes (Siegomyia)
sp. SI Coquillettidia
»jienïis (Belkins. 1962) WS?
samoaensis Sume, 1966 AS WS Culex
annulirostrís Sfcuse. 1889 Tul Upo Sav SI
^uinquefascialus Say, 1823 Tut Upo Sav SI WS samoancnsis (TheobaH. 1914) Upo SI sitiens Wiedànaan, 1S28 Tut Upo
Guien (Culex)
sp. SI Mansonia
sp. SI Toxorhynehites
amboínensis (Dolescball. I8S7) AS WS brevipolpis Theobald. I9D1 SI
^. Tvt hiéndeos (Wiedsnaim, 1819) Tut Tripteroides
purpuiatus (Edwards. 1921) AS?
DoliehDpodidae Ambiypsilopus
SOTOT (Parent, 1934) SI Chrysosoma
complicatum Becker, 1922 Swa Upo AS WS coDsiniile Lamb, 1929 Tau Tut Upo
ftiiifaimj Lamb, 192» Tau Tin Upo Sav ÍDnanim Lamb. 1929 Upo Sav iBTtcimicans B«Aer, 1923 Upo Mno Sav Icucopogon (Wiedemann, 1824) WS parvicucullaium Lamb, 1929 Upo Sav pBtellifenun (TbomsoD. 1869) SI ñAuum Lamb, 1929 Upo
Chrysotus
spp, SI Crypiophleps
solonumis Bickcl. 1996 Upo
Higher Taxon Family
GCDUS
SMcies De«^-her, Vaar MUA OFII TAÜ TUT AtlN ROS SWA tlPO MNO APQ NUTJ SAV Sf AS WS Dûlichopodidae (com.)
Cymaiopus
spinosus (Parrat, 1934) SI Hclixocems
meadosum Lamb, 1929 Tut Upo Mcdeten
griscscecs de Meîjac, 1916 Upo
süooioms (Pami, 1941) AS SyiQpycnus
sp. SI DiosophilidoB
Dettopsomyia
formoaLamb, 1914 Tut
sp. SI Dtosophila
slbiñttütata Malloch, 1934 Upo SI
•nuniKipaf Dolcscball, 1858 Tut Upo Mno Apo Sav SI bipcctiiiata Duda, 1923 Tut Upo SI
biyani Malloch, 1934 Tau Tut Upo Mno Apo SDV SI
convexa Malioch. 1934 Upo SI excqna Malloch, 1934 Tut Upo SI ftovipcs {Harmon. 1954) Upo SI Aiscovittata Hanison, 1954 Tut Upo Mno SI hypopygialis Malloch. 1934 Upo Mno
innocua (Mallocb, 1934) Tut Upo SBV
kikkawaiBurU, 1954 Upo SI manonoensis Hairisoo, 1954 Tut Upo Mno
marjoryae Mamson, 1954 Mno SI mclanogastcr Mcigcn, 1830 Tut Upo SI mcKatonim Pancnou & Wheeler, 1942 Tut
monthim de Meijeic. 1916 Upo
DBsuta Lamb, 1914 Tot Upo Mno Sav SI nasutoides Okada, 1964 TUt Upo Sav SI nigrifrons Malloch, 1934 Upo Mno Sav SI Dovicia WheelcT & Takada, 1964 Tut
nublada Wbecler & Kunbysellis, 1966 Tut Upo SI pellidosa Bock & Wbccier, 1972 Tut Upo Sav SI pleuroTittaia Hanison, 1954 Upo SI polychaOa Palíereon & Wheeler, 1942 Tut
samoaensis Hamson, 1954 Tul MDO SI
seminigra Duda, 1916 Upo Sav SI sixDuIans Sturtevaní, 1919 Tul
smmincípes (Malloch, 1934) Tut Sav SI Wblineaia Dada, 1926 WS?
milñirigaster (Duda, 1923) AS unicolorata Wheeler, 1959 Tot Upo Sav upoluae Malloch. 1934 Upo
Drosophila (Drosophila)
spp. SI Drosophila (Hirtodioiophíla)
spp. SI Drosophila (Scaptodrosophila)
spp. SI Drosophila (Sophophora)
sp. SI Lcucopheaga
samoiensis Hanison. 19S4 Mno
sp. SI
