Proceedings of the Biological Society of Washington

^ ^tB _{2 5 1930} ^

EIGHT NEW ASTERACEAE FROM ME

GUATEMALA, AND HISPANIOLA.

BY

^S. F.

BLAKE.

The new composites described

in this

paper have been found among

collections recently

received

at

the National Herbarium

or in older

material

hitherto

not

critically

examined.

Erigeron ortegae Blake,sp.nov.

Herbaceous, 30 cm. high

and

probably

much more

(thelower partnot seen), essentially glabrous throughout; stem branching, striate-angled, pale green, whitish-dotted;stemleavesalternate, 1.5 to 3cm.apart, ellip- tic-linearto linear-oblanceolate orlinear-lanceolate,3to 4.8cm.long, 5 to 8

mm.

wide, acute, sessile or graduallynarrowed intoashort petioliform base,obscurely repand, ratherfirm,dullgreenon bothsides,very narrowly pale-margined

and when young somewhat

spinulose-ciliolate, quintupli- nerved with translucent veins, sometimes obscurely pubescent above;

branch^{leavessimilar}but

much

smaller,usually oblanceolate

and

narrowed into a petiole-hke base, sometimes with a few teeth toward apex; heads hemispheric, about 1.5 cm.wide,numerous,looselypanicled,the flowering branches minutely bracteate,erect, the sparsely subulate-bracted

mono-

cephalous peduncles3 to 9 cm. long; disk7to 10

mm.

high, about 1 cm.

wide; involucre strongly graduate, about 4-seriate, 4 to 6

mm.

high, the phyllaries all appressed, the outermost ovate-oblong, the others oblong, 1.5 to 1.8

mm.

^wide,obtuseorrounded,obscurelysubherbaceous withsub- scariousciliatemargins, glabrousdorsally,3-nervedor nerveless;receptacle rounded,alveolate;raysabout18, 1-seriate,white,fertile, thetubehirsutu- lous, 2.5

mm.

long, the laminalinear-cuneate, tridenticulate, revolutein drying, 6-veined,6

mm.

long, 1,6

mm.

wide; disk flowersnumerous,^fertile, their corollaspale yellow, hirsutulous on tube

and

lower partof throat,5

mm.

long (tube 2

mm.,

throat funnelform, 2.2

mm.,

teeth 5, ovate, 0.8

mm.);

achenes of ray

and

disk similar, oblong, scarcely compressed, 5- nerved, glabrous,2.5to 2.8

mm.

long;pappusofabout 70

somewhat

unequal, stiffish,barbellate,whitishbristles3.5

mm.

long;anthertipslance-subulate;

stylebranchesshort,withshort, deltoid,obtuse, externallypapilloseappen- dages.

6—

Proc. Biol.Soc.Wash.;Vol.37, 1924. (55)

56 Proceedings

of^theBiologicalSociety of

Washington.

Type

ⁱⁿtheU.S.National Herbarium,no. 1,084,210, collectedatBalboa, Sinaloa,Mexico,Jan. 1923,

by

J.G. Ortega(no.4974).

Erigeron ortegaeisnotclosely relatedto

any

other specieswith whichI

am

acquainted. It is

somewhat

similar in appearance to the smoother formsofErigeron bonariensisL.,but verydifferent inthe characters^ofthe involucre

and

rays.

The

plant is

more

or less intermediatein character between^Aster

and

Erigeron, butis referredtoErigeronon thebasis ofits short bluntstyletips.

Guardiola stenodonta Blake,sp.nov.

Suffrutescent, oppositely branched, slender, glabrous throughoutexcept forthe

somewhat

pubescentpedicels

and

tips ofthephyllaries;upperinter- nodes ¹ to 2 cm. ^long; leaves opposite; petioles slender, margined only towardapex,1to1.5cm.long;leafblades lanceolateor elliptic-lanceolate, 5to8 cm.long, 1.5to 2.2cm.wideacrossthe basallobes, 8to12

mm.

wide near middle, acuminate toanacute apex, oftenfalcate, hastately bilobed justabove the acutely cuneatebase (thelobes linear-subulate, attenuate, oftenfalcate,wide-spreading, entireorsparsely toothed, 5to 12

mm.

long, about1

mm.

wideatbase),abruptlycontracted

and

cuneateabovethe lobes (there2to4

mm.

wide), acutelyserratewithabout 13to16pairs ofteeth 1

mm.

highorless, slightlypalergreenbeneath thanabove,weaklytripli-

nerved abovethebase, finely reticulatewithtranslucent veins

and

veinlets;

heads about 4 to8, clustered at tips ofbranches, about^1.2 cm. wide, on sparselypubescentpedicelsabout 8

mm.

long; diskthick-cylindric, 1.2to 1.5 cm. high,3 to5

mm.

thick; involucre 2-seriate, 8 to9

mm.

high, the outer phyllaries 4, subequal, oval-oblong, obtuse, about 4

mm.

wide, scarious-margined, not at all carinate, many-nerved, sparsely pubescent towardapex, the inner (subtending the rays)

membranous,

similar tothe pales;raysabout3, apparently white,^fertile, thelaminaoval, tridenticu- late,5.5

mm.

long,3.2

mm.

wide; disk flowersabout13, sterile,their corollas whitish orperhaps cream-color, glabrous, becoming 11.5

mm.

long (tube slender,8

mm.,

throatcampanulate,1

mm.,

teeth oblong,acutish, 2.5

mm.)

;

pales

membranous,

obtuse,ciUolateabove, few-nerved,about11

mm.

long;

ray achenes oblong, thickened,

somewhat

obcompressed, striate, sparsely pubescent,blackish,5

mm.

long, 2.2

mm.

wide,epappose;anthersgreenish, thefilaments denselypilose;style (ofhermaphroditeflowers) 2-parted,the brancheslinear-subulate,acuminate, hispidulous,withoutstigmaticlines.

Type

intheU.S.NationalHerbarium,no. 1,084,211, collectedatBalboa, Sinaloa, Mexico,Jan. 1923,

by

^J.G. Ortega(no.4986).

Relatedto Guardiolamexicana

Humb. &

Bonpl.

and

G. arguta(A.Gray) Robinson, but easily distinguished

by

the leaves, whicharehastate with extremely long

and

narrow basal lobes

and

are abruptly contractedjust abovethelobes.

Montanoa

pteropodaBlake,sp.nov.

Suffrutescent;stemstout,erectish-branchedabove,subterete, striatulate, sordidly pilose-subtomentose with mostly appressed hairs, glabrescent;

leaves opposite exceptⁱⁿtheinflorescence; petioles 1to3cm.long,broadly

lobeacuminate,crenate-serrate,thelateral acute,crenate-serrate

and

usuallyweakly2 or 3-lobedonthe outerside,theblade papery,deepgreen above, minutely

and

rather softly hispidulous-puberulous with incurved hairs, inage

somewhat

harsh, beneathratherdensely griseous-puberulous (the hairsalong the veins rufidulous)

and somewhat

gland-dotted, tripli-

nervedatbaseofblade

and

densely prominulous-reticulate beneath; leaves oftheinflorescenceovate,unlobed,2.5 to 4.5cm.long;headsrather

numer-

ous, cymose-panicled, 2.2 to 2.5 cm. wide, on sordidly pilose-tomentose straightor atlengthhooked peduncles1.5to 3cm.long; diskinflowerabout 7

mm.

high, 9

mm.

thick, in fruit 1.7to 2 cm. thick; outer phyllaries5, oblong-ovate, 4

mm.

long, obtusish, callous-apiculate, sordid-pilosulous;

rays 10, white, pistillate, thelaminaelliptic, bidentate, 8to 10

mm.

long;

disk flowers numerous, their corollas white, puberulous with short few- celled hairs, 3.1

mm.

long (tube0.8

mm.,

throatbroadly campanulate, 1.5

mm.,

teeth 0.8

mm.);

pales at maturity thin

and much

inflated, wedge- obovateinlateralview, 1 cm. long, 5

mm.

wide

(when

folded),

somewhat

repand atthe broad truncate apex, abruptly spinose-mucronate (mucro straight,0.5

mm.

long),stipitate-glandularontheupperhalf

and

piloseon thecosta;achenesobovoid,

somewhat

compressed,blackish,2.8

mm.

long, 1.8

mm.

wide, 1-ridgedonthesides,glabrous,epappose.

Type

ⁱⁿtheU.S.NationalHerbarium,no. 252,752,collectedbetween

San

Martin

and Todos

Santos, Guatemala, altitude 2,135-2,590 meters, 25 Dec. 1895,

by

E.

W.

Nelson (no. 3616).

A

species ofthesuhgeimsAcanthocarpha,nearest

M

^.purpurascensRobins.

&

Greenm., but readily distinguished

by

^its three-lobed leaves with the petiolesbroadly wingedtothevery base.

Hymenostephium

superaxillare Blake, sp.nov.

Herbaceous atleast above, opposite-branched, thebranches divergent, superaxillary(1to4

mm.

abovetheaxils);stemslender, striatulate, some-

what

^flattenedatthe nodes, glaucescent, sparsely tuberculatenearthenodes withshortincurved1-celledconichairs,these usually deciduous; internodes 8to12cm.long;petiolesslender,naked,similarlytuberculate, 5to15

mm^

long; blades ovate, or theuppermostlance-ovate, 8to 13cm. long, 2.5to 6cm.wide, falcate-acuminate,atbasenarrowlyorbroadlycuneate,serrate nearly throughout (teeth about 18 pairs, depressed, mucronulate), thin- papery,above deepgreen,evenlybut notdensely scabrous-tuberculatewith incurvedconichairswithlepidotebases, beneath

somewhat

lightergreen, evenly but not densely tuberculate-hispidulouson veins

and

veinlets

and

sometimes minutely ^so on surface, triplinerved from near the base

and

weaklyprominulous-reticulatebeneath; heads2.5to4cm.wide,solitaryat apex^ofstem

and

intheuppermostaxils, onslenderglabrous peduncles2.5 to 8 cm. long; disk hemispheric, 8 to 10

mm.

high, 8 to 13

mm,

thick;

involucre 3-seriate,

somewhat

graduate, 5 to 6

mm.

high, the phyllaries ovate(1.8 to 2.5

mm.

wide),acute oracuminate,withindurate,ribbed

and

58 Proceedings

of^theBiological Society of

Washington.

vittatebase

and

subequal, usually spreading,herbaceousapex,tuberculate- hispidulous-ciliolate,onbackglabrousorsparsely tuberculate-hispidulous

;

receptacle convex; rays 12, neutral, golden-yellow, the lamina elliptic- oblong, bidenticulate, about 15

mm.

long, 4.5

mm.

wide, 13-nerved; disk corollasvery numerous, yellow, slender-cylindric,hispidulouson tube

and

lowerhalf ofthroat, 5

mm.

long (tube0.8

mm.,

throat3.5

mm.,

teeth0.7

mm.);

palesacuminate, many-striate,

somewhat

hispidulous,about6

mm.

long; achenes obovate-oblong, compressed, glabrous, epappose, 2.5

mm.

long.

Type

inthe U. S. National Herbarium, no, 1,083,277, collected at

La

Bajada, Tamazula, Durango, Mexico,altitude 300-600meters,Nov. 1921,

by

^J.G. Ortega(no.4437).

Thisspecieshasthelargestheadsof

any known

Hymenostephium,

and

is further distinguished

by

^{itsphyUaries,}whicharebroaderthanin

any

other species

and

do not have the attenuate or very narrowly acuminate tips foundin practicallyalltheothers. Itisnearest to theepappose formofH.

cordatum (Hook.

&

Arn.) Blake.

The

conspicuously superaxillary branches from which the

name

is derived are

shown by

^allthe sixoreightspecimensofthe type collection examined,

and by

noothersinalarge series ofspecimensrepresentingthe other speciesofthe genus.

Verbesinapeninsularis Blake, sp.nov.

Suffrutescent orfruticose, 60 cm. high

and

more; stem rather slender, subterete,striatulate,wingless,white,ratherdensely hispidulous-pilosulous with mostly incurvedorspreadinghairswithsubtuberculate bases; lowest leaves (ofincomplete specimen examined) opposite, the others alternate;

petioles 5 to 10

mm.

long, cuneately winged to base

by

the decurrent lamina; blades triangular-ovate orsometimeslanceolate,thelarger9to11 cm.long,4.5to 5.8cm.wide,acuminate, cuneateatbase

and

decurrenton thepetiole, coarsely

and

irregularlydentate (theteethtriangular, callous- tipped)

and

usually

somewhat

hastate-lobedat base, firm-papery, above dull green, densely incurved-hispidulous with tuberculate-based mostly deciduoushairs, scabrous,beneathslightlypalergreen, ratherdensely

and somewhat

harshly hispidulous with antrorse tuberculate-based hairs

and somewhat

gland-dotted,triphnervedabout1cm.abovethebase

and

loosely prominulous-reticulate,thesecondaryveins few;heads2.5to 3cm.wide,in aterminal short-pedunculate

cyme

ofabout5and on1 or2-headedpedun- clesfromtheupperaxils,thepedicels1to8cm.long,pubescentlikethestem;

disk campanulate-hemispheric, 1.1 to 1.3 cm. high, about 1.5 cm. thick;

involucre2-seriate,subequal, 7to8

mm.

high,thephyllariesoblong-ovate or oval-oblong,subappressed, the outer herbaceous, obtuseor acutish, hispid- pilosewithglandular-tuberculate-basedhairs

and

ciliate,2to 3.5

mm.

wide, the inner thinner, subscarious, obtuse, lesspubescent ornearly glabrous;

raysabout10, pistillate,yellow,the tubepilose, 1.8

mm.

long, thelamina

linear-elliptic,9to12

mm.

long,about2.8

mm.

wide; diskflowersnumerous, their corollasyellow,piloseon tube

and

baseof throat, 6.7

mm.

long (tube

thefaces or 2-winged, thewings

ciliate;

awns

2,slender,unequal, 3to4

mm.

long.

Type

ⁱⁿtheU.S.National Herbarium,no.565,313,collected32kilometers eastof

San

Ignacio, BajaCalifornia,Mexico, altitude365-610meters, 19 Oct. 1905,

by

E.

W.

Nelson

and

E. A.

Goldman

(no. 7227).

A member

ofthe sectionSonoricola,relatedtoV.leptochaetaA.

Gray and

V. erosa T. S. Brandeg.

The

^latter has

much

narrower phyllaries

and

chieflyopposite leaveswhicharedensely antrorse-hirsute orhirsute-pilose beneath.

The

former, a

Chihuahuan

species

known

to

me

only

from

description,issaidtohaveoppositeserrate leaves, narrowly oblongwide- spreading outerphyllaries,

and

smallrays.

Perityle

macromeres

Blake, sp.nov.

Herbaceous, 60 cm. high

and

more,thebasenot seen;stemstout (8

mm.

thick below),

somewhat

branched, striate, sparsely villousnear thenodes

and

verysparselybetween

them

withseveral-celled hairs,obscurely glandu- lar;lower internodes 14cm.long;lower leaves (2or

more

pairs)opposite, the others mostly alternate; petioles 1 to 3 cm. long; blades ovate-sub- orbicular in outline, 2 to4 cm. long, 2 to 5.5 cm. wide, 3-nerved, deeply 3-lobed, thelaterallobeswith2oblonglobesonthe lowerside,the terminal lobe rather deeply 3 or 5-lobed, the primary lobes all lacinate-lobed or toothed withacute or acutish teeth

and

roundedsinuses, thinnish,sparsely villous chieflyonthe veins

and

margin; headsseveral,about2.5cm.wide, solitaryondenselystipitate-glandular,sparselyvillous,

naked

or bracteate peduncles 6 to 12cm.long;diskrounded,9to10

mm.

high, 1.3 to 1.8cm.

wide; involucresub-2-seriate, equal,5

mm.

high,thephyllaries lanceolate, acuminate, 2-ribbed, villous

and

stipitate-glandular; rays 13, fertile,

apparentlywhite, the lamina oblong, bluntly 3-toothed, 8 to9

mm.

long, 2.5 to 3

mm.

wide; disk flowers verynumerous, their corollasapparently whitish, 4-toothed, densely stipitate-glandularontube, sparselysoonlimb, pilosulousonteethwithseveral-celled hairs, 2.8 to 3.2

mm.

long (tube1.3to 1.6

mm.,

throat funnelform-campanulate, 1 to 1.3

mm.,

teeth 0.3 to 0.5

mm.);

achenes linear-obovate, 3

mm.

long, very flat, blackish, densely ciliateonthenarrowcallousmargins, glabrousonthefaces;

pappus awn

1, setiform, weak, retrorse-hispid abovethe middle, 2.2

mm.

long;squamel- laceouscoronaminute,cihate; stylebranches withrathershort, lanceolate, hispidulous tips.

Type

intheU. S. NationalHerbarium,no.565,548,collectedat

La

Paz, Baja California,altitude3 meters, 3 Feb. 1906,

by

E.

W.

Nelson

and

E.A,

Goldman

(no.7483).

RelatedtoPerityle crassifoliaT.S. Brandeg.

and

P. robustaRydb.,

and

agreeingwith

them

inthesingle,weak,retrorselyhispidpappus

awn and

the clavellate cilia ofthe achene, butdifferinginits

much

larger

and

thinner leaves

and

longer peduncles, as well asinits

much

scantierpubescence

and

60 Proceedings

of^theBiologicalSociety of

Washington.

glandularity. P.crassifoliaisverydenselyviscid-villous,almostarachnoid,

and

P. rohusta, whichisdoubtfully distinctfrom P.crassifolia, isdensely stipitate-glandularthroughout

and

denselyvillous tomiddle,less densely soabove.

Perityletrichodonta Blake, sp. nov.

Suffruticulose,branched,

many-stemmed,

10cm.high,thestemsglabrate;

branches, peduncles,

and

leaves densely griseous-pilosulous with several- celled mostly spreading hairs; internodes of branches 3 to 5

mm.

^long;

leaves opposite essentiallythroughout;petioles2to3

mm.

long, griseous- pilosulous; leafblades ovateorrhombic-ovate, 3to5

mm.

long, 2.5to5

mm.

wide, obtuse, at basecuneatetosubtruncate,entireorhastately1-toothed on each side at base, thickish, densely impressed-punctate especially be- neath; heads7

mm.

^wide, solitaryat apexofstem

and

in theuppermost

axils, onpeduncles 5to7

mm.

long;disksubglobose-campanulate, 5

mm.

high, 5 to 6

mm.

thick; involucre sub-2-seriate, equal, 4

mm.

high, the phyllaries about 24, linear, acute, double-ribbed, rather densely griseous- pilosulousontheirexposedsurface;rays8, fertile,thelaminawhite, 1.5to 1.8

mm.

long,about1.5

mm.

wide,bluntly 3-toothed,gland-dottedon back;

disk flowers very numerous, their corollas white, stipitate-glandular throughout,

somewhat

hirsute-piloseontube,barbate-tuftedonteethout- sidenear apex, 3

mm.

long (tube 1

mm.,

throat funnelform-campanulate, 1.5

mm.,

teeth4,ovate, 0.5

mm.

long);achenesobovate-oblong, 1.8 to2.2

mm.

long, flat, blackish, ciliate on the whitish callous margins, sparsely hirsutulousonthefacesabove;

awns

2,setiform,minutelyhispidulous,equal or unequal, 1.5 to 2.2

mm.

long; squamellaelacerate,0.4

mm.

long; style branches withrathershort,subulate, hirsutuloustips.

Type

ⁱⁿtheU.S.NationalHerbarium,no.301,934,collected intheSierra

Madre

west of Bolanos,Jalisco, Mexico, 15-17 Sept. 1897,

by

^J.N.

Rose

(no.2978).

The

typesheet has been identifiedas Peritylejaliscana A. Gray, a re- lated species with

much

larger leaves, discoid heads, merely glandular corolla-lobes,

and much

shorter

awns and

squamellae. P. trichodonta is

nearer P. hofmeisteriaRydb.,afinelypuberulent plantwithpetiolesmostly

much

^{longerthantheblades,}

^and

^{withthe diskcorollasmerelyglandularon}

theteeth.

Anastraphia enneanthaBlake, sp.nov.

Shrub

orsmall^tree,leafy,the leavescloselycrowded onthe shortlateral branches;

young

branchlets angulate, closely ochroleucous- or cinereous- tomentose, the older ones^glabrate, gray-barked; leavesalternate; petioles cinereous-tomentose, 2 ^to 3

mm.

long; leaf blades cuneate-obovate or obovate, 2to 3.5cm. long, 1 to2 cm. wide (excludingtheteeth),acute to subtruncate, spinose-tipped, acutely cuneate at base, spinose-dentate (teeth 3to 5pairs, 3 to6

mm.

long including thebrownishspines), cori- aceous, above deep ^{green, glabrous} from the first,

somewhat

shining, finely reticulate, beneath closely cinereous- or ochroleucous-tomentose,

somewhat

reticulate, the chief lateral veins about 4 ^pairs, prominulous

outermost

somewhat

tomentose, belowthe apex, theoutermost verysmall,deltoid-ovate,themiddletriangular,theinnermostlance-linear, allacuminate, scarcelymucronate;corollastubular,

somewhat

hispidulous

and

glandular, at maturity 16.5

mm.

long (tube 4.5

mm.,

throat 2

mm.,

teeth 10

mm.,

revolute at apex); achenesdenselyshort-pilose, 5

mm.

long;

pappus brownish-tinged, 1.5cm. long, the slenderbristlesstiff,spreading,

somewhat

paleaceous-connate at base; anthers

and

style long-exserted;

stylebranches veryshort.

Type

intheU.S.National Herbarium,no. 1,079,231, collectedatLajana,

Samand

Peninsula, Santo

Domingo,

altitudeabout100 meters, 30April-2

May

1922,

by W.

L.Abbott(no.2287).

Among

the species of Anastraphia hitherto

known

from the island of Hispaniola theclosest ally of this speciesisapparentlyA.oliganthaUrban, whichisreadilyseparated

by

^its5-flowered heads. A.enneanthaseemsto be distinctfrom

any

of the

numerous Cuban and Bahaman

^{species, all}of which, aswell asthespecies ofHispaniola, have beenlisted

and

keyed

by

Britton.i

iBull.TorreyClub42:379-387.1915.