PR.OCEEDINQS
OF
THE
WASHINGTON ACADEMY OF SCIENCES
Vol.
IV., pp. 49-86.[Plates
i-iii.]March
27, 1902.PAPERS FROM THE HOPKINS STANFORD GALA- PAGOS EXPEDITION, 1898-1899.
VII.
ENTOMOLOGICAL RESULTS
(6).ARACHNIDA.
By Nathan Banks.
With
FieldNotes
byRobert
E. Snodgrass.CONTENTS
Part I, byNathan Banks:
Extent and Character of the Collection 50
Affinities of the Fauna 51
Geographic Distribution 54
Technical Descriptions:
Araneida Filistatidae 55
Scytodidse 55
Pholcidae
....
56Djsderidae . 57
Drassidae 57
Clubionidie 58
Agalenidse 58
Dictjnidse 59
Theridiidae 59
Epeiridse 60
Thomisiidte 62
Sparassidae 63
Ctenidae 64
Lycosidae 64
Attidae 66
Arthrogastra . . Phrynida 67
Scorpionida 68
Pseudoscorpionida 68
Solpugida 69
Acarina Ixodidae 70
Tyroglyphidse 70
Part II,byR. E. Snodgrass:
FieldNotes 71
Proc. Wash.Acad.Sci.,March, 1902. (49)
50 BANKS
PART
I.EXTENT AND CHARACTER OF THE COLLECTION.
About
sixhundred and
fiftyArachnids were
collectedby
this expedition,making
thecollectionby
far the largestever brought from
theGalapagos
Islands.The specimens belong
to forty- eight specieswhich
fall in the three greatgroups
ofArachnida
as follows:Araneida,
thirty-nine species;Arthrogastra,
six species;Acarina,
three species.The
thirty-nine spiders represent fifteen families, ofwhich
the EpeiridcEhave
nine species, the Attidee five,fhe Theridiidae four, the Sparassidaeand Lycosidee
three each,while
the other families arerepresented by
butone
ortwo
species each.Twenty-five
species areconsidered new, and
arehere
firstdescribed
;two
are classifiedonly
generically;twenty-one
areknown
species, ofwhich
four are peculiartotheseislands.The
collection contains,
with two
exceptions,specimens
of allspeciesever taken on
these islands.The exceptions
areSegestria
ceqtiatoria ofMarx's manuscript, and
hisyoung Agalena.
The
first spiderknown from
these islands—
Gasteracantha insulana — was described by Thorell
in 1859. ^^^'^'^^been taken
by
allsubsequent
collectors. In1877
Butlerpublished
in theProceedings
of theZoological Society
ofLondon
a list of the species collectedby
the Petrel expedition. In1889 Dr. Marx pub-
lished alist(without descriptions)intheProceedings
of theU.
S.National Museum
of the species collectedby
theAlbatross expe-
dition.These
liststogether include notmore than
thirteen spe- cies,though appearing under
sixteennames. They
are asfollows :Recorded
by Butler.Recorded
byMarx.
Androctonus americus
Linn. Segestria ceqtiatoria sp. nov.Lycosa indomita
Nicolet.Agalena
(immature).Theridiuin caroli7iu?n sp. nov. Loxosceles galapagoensis sp. nov.
Lathrodectes apicalissp. nov. Filistata oceanea sp. nov.
Gasteraca7ithainsulana Thorell. Lathrodectesapicalis Butler.
Epeira
cooksoniisp. nov. Lathrodectes scelio Thorell.Thomisoides
uUrifor7nissp. nov.Gasteracantha
insulaiia Thorell.Epeira
cooksonii Butler.Heteropoda
venatoria Linn.Alenemertts galapagoc7zsissp.nov.
Ce77truroides luctifer sp. nov.
Vejovisgalapagoe7isissp. nov.
ARACHNIDA
5Probably
allof thoserecorded by
Butler arein this collection.The Thei-idhun
I consider as ayoung Lathrodectes
; theLycosa
as
one
of the three species,but none agree with
Nicolet's figure;and
theAndroctomcs was probably one
of thetwo
scorpions.Of
thoserecorded by Marx
all buttwo
arerepresented
in this collection; theSegestria and
theAgalena. Dr. Marx never published
descriptions ofthenew
speciesmentioned. His
Filis- tataand Loxosceles agree with my
species of thesame genera
his
Menenierus
isPlexippus
-paykiilli; hisCentruroides
isCen- trurus ^rtnceps
; his Vejovis isHadruroides
hcnattis,and
hisLathrodectes
sceliowas probably
theyoung
ofL.
apicalts.Many
of the spiders collectedby
this expeditionand
not col- lectedby
otherswere taken
inDecember, January, February and March. Other
species will doubtlessbe found
inSeptem- ber and October,
so that the totalArachnidous fauna
of the islands willprobably reach one hundred
species.With one important exception
all the largegroups
ofArach- nida
are represented.There
isno Phalangid. The presence
of aSolpugid
isunexpected, and
itmust have been
a rare acci-dent
thatstranded one
of theseanimals
alive somany
milesfrom
themainland.
AFFINITIES OF THE FAUNA.
As
to the affinities of thespider-fauna
of these islands, itshould be
stated that anumber
of thenew
speciesbelong
togroups which have been
but little collectedon
themainland, and
thereforemay
yetbe found
tooccur elsewhere. This
is illustratedby
the fact thatone
speciesdescribed by
Butlerfrom
these islandsnow
turns out tobe
identicalwith
a speciescom- mon
inMexico and
thesouthwestern United
States.The
affinities of thefauna can
bestbe shown by
tabulation.I
have arranged them
in fivegroups.
Cosmopolitan.
No. of specimens.
Tege7iaria derha7ni
on
shipboard iCo77i77ion T/'opical Species.
Argiope
arge7ttata 64JIete7-opoda veiiatoria 6
Plexippus paykulli 2
Total 72
52 BANKS
Known from Western South America.
No. of specimens.
Argiope
argentata64
Tmarus
stolzmanni 2Heteropoda
venatoria 6Plexippus
-paykulli 2Hadruroides
lunatus 19Total 93
Known from
CentralAmerica^ West
Indies.,and
Colombia.Dictyna
parietalis iTheridiufn mixtufn 20
Argyrodes jucundus
10Argiope
argentata64
Epeira
oaxensis 109Epeira
labyrinthea 87Epeira prompta
11Epeira
gregalis4
Cyclosa
caudata
65Argyroepeira
nigriventris 10Heteropoda
venatoria 6Plexippus
payktilli 2Marptusa
califortiica iCentrurus
princeps 5Total
^
Known
only fro7n the Galapagos.Sicaroides ultrifor?fiis
4
Loxosceles longipalpis 10
Filistatafasciata 2
Coryssocne?nis
(two
species) 3Ariadne
tarsalis 5Pcecilochroa bifasciata i
Prosthesima galapagoensis
iAysha
pacifica 3Theridium
sp iLathrodectes apicalis 27
Gasteracatztha insulana 27
Tetragnatha
galapagoensis 22Misumena
inclusa 2Oliosgalapagoensis 14
Selenops galapagoensis 6
Odo (two
species) 9Zjj/co5a (three species) 14
Philceuspacificus 5
Cyrba
insularis iARACHNIDA 53
Admestina
insularis iCharinus
insularis 8Amfnotrecha
solitaria iPseudoscorpions
(two
species) 8Afnblyotnma
pilosu?n49
Arg-as transversa i
Tyroglyphus
sp 3Total 227
It thus
appears
that,though only
fourteen species arecom-
mon
to theCentral American
region, as against thirtyfound only
in theGalapagos,
theCentral American
species are repre- sentedby nearly twice
asmany specimens. Moreover, some
of the speciesconfined
to theGalapagos
arevery
closely allied to speciesfrom
theCentral American
region.For example,
the Oliosgalapagoensis
isvery
close toO
.fascictilatusfrom Mexico and
thesouthwestern United
States.Excepting Charinus
(an Asiaticgenus),and Coryssocnemis
(aBrazilian genus), thegenera
of theGalapagos
speciesareallrepresented
intheCentral Amer-
ican region.
Again, when we compare
the distributionon
theGalapagos
Islands of thecommoner
specieswe
find that the speciesfrom
theCentral American region
are themost widely
distributed, as willbe seen by
thefollowing
tables :Speciesrepresented
by more
than ten specimens are distributed as followsConjinedto
Galapagos.
Amblyo77ima
: allfrom Albemarle
except onefrom
Narboro.Tetragnatha
: Albemarle, Narboro.Lathrodectes: Bindloe,
Tower,
Charles,Chatham.
Olios: Narboro, Albemarle,
Chatham.
Gasteraca7itha: James, Charles,
Chatham,
Narboro, Albemarle.Loxosceles: Albemarle,
Hood.
Known
fro7n CentralATnerican Region.
TheridiuTn: Albemarle, James, Narboro.
Argyroepeira
: Albemarle,James,
Narboro.Argiope:
Albemarle, James, Narboro, Charles,Hood,
Bindloe, Harrington, Indefatigable.Epeira
pro77ipta: Albemarle,James,
Narboro.Epeira
oaxensis: Albemarle,James,
Charles,Chatham, Hood,
Duncan,
Barrington, Indefatigable.54 BANKS
GEOGRAPHIC DISTRIBUTION.
TABLE OF KNOWN DISTRIBUTION OF SPECIES ON THE
DIF-FERENT ISLANDS OF THE ARCHIPELAGO.
ARACHNIDA
55Epeira
labyrinthea: Albemarle, James, Narboro, Charles, Chat-ham, Hood,
Barrington, Bindloe,Tower, Abingdon,
Indefatigable.Cydosa:
Albemarle, Narboro,Hood,
Indefatigable.Argyrodes
: Albemarle, Narboro.From
these considerations Iconclude
that theArachnidan fauna
of these islands ismore
closely related tothatof theCen-
tral
American region than
to that ofany
other portion of the globe.TECHNICAL DESCRIPTIONS.
Order ARANEIDA.
Family FILISTATID^.
FILISTATA FASCIATA
sp. nov.(PI. I, fig. 15.)
Length
5mm.
Cephalothorax dull yellowish: legs similar, with not very distinct dark
marks on
thefemora beyond
middle,and
baseand
tipof tibia,and on
tip of metatarsus;abdomen
darkbrown
above, withtwo
pale spots at baseand
fourcurved pale bands, the intermediatetwo
being barely interruptedon
the middleline,none
ofthem
reachingthesides, also an indistinctapical spot; venter rather paler than dorsum, espe- cially at base; sternum yellowish. Cephalothorax of usual shape (brokenand
positions of eyes not evident) ; palpi largeand heavy
legs rather shortand
hairy;abdomen
large; projecting considerablybeyond
the spinnerets.Two
specimens, one young,from Wenman
inDecember.
Easily recognizedby
itsbanded abdomen and
legs. It will fall in F.O.
P.Cambridge's genusFilistatoides; but I do not
deem
a divisionofFil- istata to be necessary.Family SCYTODID^.
SICAROIDES ULTRIFORMIS
Butler.Thomisoidesultriformis Butler, Proc. Zool, Sec. Lond., 1877, p. 'j'j.
Several specimens
from Hood
Island,May.
Describedfrom
the Galapagos.The
genus occurs in CentralAmerica and
Chili.LOXOSCELES LONGIPALPIS
sp. nov.(PI. I,fig. 13.)
Length
9 1 1mm.
Cephalothorax, legs
and
sternum pale yellowish, the legsmore
red- dishtoward
tip; the mandibles pale reddishbrown
; dorsal groove56 BANKS
and
anterior furrowsblackish; eyeson
blackspots;abdomen
grayish,more
yellowbelow
; in themale
the palpi are yellowishon femora and
patellae, reddishbeyond
; the eyes in groupsas usual, the S. E.inthe female fully their longest diameter
from
theM.
E., in themale
they are closer totheM.
E. Otherwisethe structureis similartoL.
rufescens, but the
male
palpus, especially the tibial joint, ismuch
longer than in that species.
Ten
specimensfrom Albemarle and Hood, May.
Family PHOLCID^.
CORYSSOCNEMIS CONICA
sp. nov.(PI. I,figs. I, 2,3.)
Length
4.5mm.
Cephalothorax pale yellowish; with a
median brown
stripe, not quite reaching to hind border,but in frontextendingdown
to clypeus, broad in front, with parallel sides tobeyond
middlewhere
it is sud- denlyconstricted to one half the width; a curvedbrown
stripe each sidewhich
is connected tothecornea of themedian
stripe; eachgroup
of side eyesis situatedon
a jet black spot; legspale, faintlygreenish, with abrown
preapicalband on femora and
tibiaebeyond which
is aband
of purewhite.Abdomen
ratherbluish gray,indistinctlymarked
with darker patches. Cephalothorax broadand
flat, eye-region slightly elevated.The median
eyes smalland
above the anterior laterals.Legs
longand
very slender.Abdomen
elevated behind into a pointed cone; theepigynum
prominent.Two
specimensfrom Hood
inMay.
CORYSSOCNEMIS INSULARIS
sp. nov.(PI. I, fig. 6.)
Length
4mm.
Cephalothorax yellowish,black
around
eyes, a broadmedian brown
stripe reaches
from
eyes to near hind margin, suddenlynarrowed
be- fore dorsal groove, abrown median
stripeon
clypeus; mandibles pale reddishbrown;
legspale brownish, with a very distinct apicalwhiteband on
allfemora and
tibiae;abdomen
bluish gray, without marks, paler beneath. Structure as in C. conica, but the legs aremuch
stouter
and
longer than in that species,and
theabdomen
is not pro- longed above ina pointed cone,but broadlyrounded
behind, althoughit projectsconsiderablyoverthespinnerets; the
epigynum
is largeand
prominent, reddishbrown.
One
specimenfrom
Iguana Cove,Albemarle
Island; in June.57 Family DYSDERID^.
ARIADNE TARSALIS
sp. nov.(PI. I, fig. 9.)
Length
8mm.
Cephalothorax yellowish
brown,
scarcely darkerin eye-region, sidemargin
black; mandibles dark red-brown, almost black at tips;sternum
and
legs yellowish, metatarsiand
tarsi of anterior pairs darkbrown,
metatarsus I nearly blackish at tips;abdomen
dark gray above, slightly paler beneath; spinnerets pale. General structure as usual; tibiae Iand
II withfour pairs of stout spines beneath; meta-tarsi I
and
II with nine pairs of spines beneath; afew
spineson
tibiaand
metatarsusIII; no spineson
legIV
except small one under baseand
one at tip of metatarsus.A few
specimensfrom Culpepper
Island,December.
This
is not the Segestria ceguatoria ofMarx, which
is a genuine Segestria of usual appearance; herring-bonemark on
back,and banded
legs.Family DRASSID^.
PROSTHESIMA GALAPAGOENSIS
sp. nov.(PI. II, fig. 7.)
Length
5mm.
Cephalothorax uniform yellowish
brown,
margins black; mandiblesmore
red-brown; legsand
sternummore
yellowish;abdomen
grayishbrown,
paler beneath. Posterior eye-row slightly procurved, the P.M.
E. ovaland
touching at hind angles, about their short diameter from the equal P. S. E.; quadrangle ofM.
E. onceand
one-half higher than broad, broadest in front: anterior eye-row plainly pro- cui-ved,A. M.
E. rather largerthan P.M.
E.,lessthantheirdiameter apart,and
still closerto the equalA.
S. E.Sternum
but little longer than broad, broad in front, but littlewider
in middle, pointed be- hindbetween
the hind coxae.Legs
moderately slender, no spines under tibiae Iand
II, one at baseand
one near tip under metatarsi Iand
II,many on
hind pairs,some
above.Abdomen
oneand
three- fourths times as long as broad, but little depressed,and
notmuch widened
in the middle, truncate at base, pointedbehind.One
femalefrom Albemarle
taken at sea level nearIguanaCove
in June.PCECILOCHROA BIFASCIATA
sp. nov.(PI.I, fig. 4.)
Length
5.5mm.
Cephalothorax
and
legs shiningyellowishbrown
; the mandiblesand
sternummore
reddishbrown;
eyeson
blackspots;abdomen
pale58 BANKS
yellowish, a
narrow
scarcelyvisible basal black band, a broad sub- basal band, a still broader,middle
band,and two
small spots each side near tip, black; spinnerets black; venter pale. Posterior eye-row
distinctly recurved; the P.M.
E. round, fully their diameter apart,and
as farfrom
the equal P. S. E., anterior eye-i"ow procurved,some
distance infront of the posteriorrow, the quadrangle ofM.
E.beingnearly twice aslongasbroad;
A. M.
E. larger than P.M.
E., lessthantheir diameter apart,and
about one-half their diameterfrom
the smallerA.
S. E. Metatarsiand
tarsi Iand
II with scant scopulas;
three pairsof spines under metatarsusI
and
II; sternum truncate in front, scarcelywider
in middle, blunt behind;abdomen
depressed, themale
with a basal reddishhorny
plate extending nearlyto middle band.One male from Narboro
inJanuary.Taken from Termite
excava- tions in a piece of driedwood washed
ashore nearthe northeast point of theisland.Family CLUBIONID^.
AYSHA PACIFICA
sp-. nov.(PI.I, fig.II.)
Length
$ 4.5mm.
; 9 5.5mm.
Cephalothorax pale yellowish
brown
; the dorsal grooveand
an ir-regular stripe each side not reaching hind margin,
brown;
eyeson
black spots; mandibles dark red-brown; legs pale yellowish, hindtibiae distinctly
banded
with blackish near tip;abdomen
pale,marked
with blackish, a large elongate spot each side near base, followedby
a series of smalland
irregular spots, blackish (in themale
thesemarks
nearly cover thedorsum)
; venterand
sternum pale (inmale
with threemedian
spotson
venter). P.M.
E. rather larger than other eyes,A. M.
E. fully equal toA.
S. E.; clypeus verylow;
mandibles stout, but nearly vertical; legs of moderate length; fold of venter
much
nearerto base thanto tip. Palpus of male small.Two
specimensfrom Albemarle
inJanuary,and
onefrom Chatham
in
May.
Family AGALENID^.
TEGENARIA DERHAMI
Scopoli.ScoPOLi, Entom. Carniolica, p. 400(1763).
One
specimenfrom
water cask (filled inTagus
Cove,Albemarle
Island) on schooner, in February.A
cosmopolitan species; thespecimen may
havecome from San
Francisco, and notnow
belong to the Galapagosfauna.59 Family DICTYNID^.
DICTYNA PARIETALIS
CamiDiidge.Cambridge, Biol. Cent. Amer., Arach. Araneida, i, p. 171 (1896).
One specimen from Albemarle
Island, inMarch.
Apparently
thesame
as thiscommon Mexican
species.Family THERIDIIDiE.
THERIDIUM MIXTUM Cambridge.
Cambridge, Biol. Cent. Amer., Arach. Araneida, i, p. 206 (1898).
Twenty
specimensfrom Albemarle
inMarch
;from James
in April;and from Narboro
inApril. Describedfrom Guatemala
; also occurs in Texas, and probably in northernSouth America. Both
paleand
dark coloredforms were
takenon same
day and atsame
place.THERIDIUM
sp.A
small,pale species,with a largegloboseabdomen. The
cephalo-thorax with abroad black stripe, broaderbehind than in front; abdo-
men
with a large black basal spear mai'k, on each side ofwhich
isa whitemark and
awhite dot; leg I with reddish at tip offemur and
tibia, also
on
tip of tibia IV.One
femalefrom Albemarle
in January.Near
to T. lijnaense.Found
in a curled leaf, containing also a cocoonand
adult ofAysha
pacijica.
LATHRODECTES APICALIS
Butler.Butler, Proc. Zool. See. Lond., p. 75 (1877).
Twenty-seven
specimensfrom
Bindloe,June; Tower, June;
Chat-ham, May, and
Charles,May.
Describedfrom
the Galapagos. It isvery
much
likeourcommon L.
itiactansKoch. Does
not appear to have been foundon
Albemarle, though collectingwas
extensive there at theproper time.Theridium
caroliniDU Butl. ibid, is, I think, without doubt, ayoung specimen
of Lathrodectes.ARGYRODES JUCUNDUS Cambridge.
Cambridge, Proc. Zool. Soc. London, p. 326 (1880).
Keyserling,Die Spinn. Amer. Therid., p. 190 (1884).
Ten
specimensfrom
Albemarle, January,and
Narboro, January.Described
from
Brazil ; occurs also inMexico and
extreme southern California.6o BANKS Family EPEIRID^.
GASTERACANTHA INSULANA
Thorell.Thorell, Nya
Exotiska Epeirider; Ofv. K, Vet. Akad. Forh., p. 302 (1859).Twenty-seven
specimensfrom
Albemarle,January and June
;
Charles,
May
;Chatham, May
; James, April;and
Narboro, Janu- ary. Describedfrom
the Galapagos.Recorded by
both Butlerand Marx.
Itbelongsto agroup
of species verycommon
throughout thewarmer
parts ofAmerica. Those from Chatham
Island have thedorsum
mostly black, withtwo
rather small spotson
anterior half; in the other specimensthese spots covermost
of the anteriorpart of the dorsum.ARGIOPE ARGENTATA
(Fabricius).Araneaargentata Fabricius, Entom. System., 11, p. 414(1793).
Argiopes argentata C. Koch, Die Arach., v, p. 38 (1839).
Sixty -four specimens
from
Albemarle,May
; Charles,May
; James, April; Narboro, April;Hood, May
; Bindloe,June
; Harrington,May
;and
Indefatigable,May.
Very common
throughout tropicaland
subtropicalAmerica.
EPEIRA OAXENSIS
Keyserling.Keyserling, Sitzungsber. d. Isis, Dresden, p. 121 (1863).
Keyserling, DieSpinn. Amer., Epeiridae, p. 238 (1893).
E. cooksonii Butler, Proc. Zool. Soc. Lond., p. 76(1877).
E. vertebrata
McCooK,
Proc. Acad. Nat. Sci. Philad., p. 196 (1888).One hundred and
nine specimensfrom
Albemarle,March
; Charles,May
;Chatham, May
; James, April;Hood, May
; Indefatigable(many),
April; Harrington,May, and Duncan, May.
E.
cooksoniiwas
describedfrom
theGalapagos
;E.
vertebrata^from
California,and
later(McCook, Amer.
Spid., vol. Ill, p. 152), recordedfrom
the Galapagos.Marx
has called attention to the fact thatthetwo
areonespecies,and
in manuscriptalso refersthem
toE.
oaxensis^ with
which
I fully concur; this last specieswas
described, as itsnames
indicates,from
Mexico.EPEIRA LABYRINTHEA
Hentz.Hentz, Journ. Bost. Soc. Nat. Hist., v, p. 471 (1847).
Eighty-sevenspecimens
from
Albemarle, Charles,Chatham,
James,Hood (many),
Narboro,Tower,
Indefatigable, Harrington, Bindloeand Abingdon
Islands, in February,March,
April,May and
June.Widely
distributed throughoutNorth and some
parts of SouthAmer-
ica. Dr.
Marx
had, in manuscript, proposed to consider this a dis-tinct species, but
many
of the specimens agree perfectly with speci-mens from
the United States,and
the variations are not extreme.EPEIRA GREGALIS Cambridge.
Cambridge, Biol. Cent. Amer., Arach. Araneida, i, p. 22 (1889).
Keyserling, Die Spinn. Amer,, Vol, iv, Epeiridas, p. 177 (1892).
Four
specimensfrom Narboro
Island in April.Described
from Panama
; recordedby
Keyserlingfrom
Brazil.EPEIRA PROMPTA
Hentz.Hentz, Journ. Bost. Soc, Nat, Hist., v, p. 472 (1847).
E.parvula Keys., Beschr. n. Orbitel., p. 131 (1864).
Eleven
specimensfrom
Albemarle, Narboro, andJames
islands, in February,March, and
April. Describedfrom
the UnitedStates; oc- curs quitecommonly
inMexico and
CentralAmerica. An
extremely variable species, but these specimensare of an ordinary form.CYCLOSA CAUDATA
Hentz.Hentz, Journ. Bost, Soc. Nat. Hist., vi, p. 23 (1850),
Sixty-five specimens
from Albemarle, Hood, Narboro and
In- defatigable islands, inJanuary, February,March,
Apriland May. A
species widely spread over the United States,
Mexico and
CentralAmerica.
ARGYROEPEIRA NIGRIVENTRIS
Keyserling.Keyserling,
Neue
Spinn. a. Amerikas, i, p. 316(1879).A.fragilis Cambridge, Biol. Cent. Amer., Arach. Araneida, I, p. 6 (1889).
A. vohipis Keyserling, Die Spinn, Amer.,Vol. iv, Epeiridae, p. 356 (1893).
Ten
specimensfrom
Albemarle, Narboro,and James
islands inFebruary and
April. Describedfrom New Granada (Colombia)
; re- cordedby Cambridge from
Guatemala,and by
Keyserlingfrom
Brazil.One specimen comes from Cocos
Island.TETRAGNATHA GALAPAGOENSIS
sp. nov.(PI. I, fig. 10.)
Length
9 10mm., $
7mm.
Cephalothorax, mandibles and legs pale yellowish, eyes
on
black spots, fangof mandibles dark red-brown, tips of legs dark;abdomen brownish
yellow, with a pale irregular stripe around baseand on
sides, indistinct in themale,venterin the female witha
median brown
stripe.
Eyes
with S. E. aswidely separated asM.
E., andA.
S. E.rather smaller than the others.
Mandibles
in both sexes as long as62 BANKS
the cephalothorax, deflected at angle of 45° , slightlydivergent, sub- cylindrical,seen
from
thesidescarcelyconvexabove. Inthemale
thereis,
on
the under side, arow
of about twelve teeth at nearly equal dis- tances apart, those at thetip stouter but notmuch
longer than thosetoward
base; above there are three large subequal teeth near tip,on
innerside four, a small one near base, close to itone of the others, these three being subequal in sizeand
verymuch
smaller thanthose at tip.The
fangreachesinan even curve almosttothebaseofmandibles,its apical third
on
innermargin
is microscopically denticulate. In the female'there is one tooth over base of fang,and
one near apical fourth,on
innermargin
;below
there is arow
of six to eight small teeth; the fang at base beneath is swollen in a blunt tooth.Legs
moderately slender,femur
I fully twice as long as cephalothorax.Abdomen
abouttwice the length of the cephalothorax, inmale
nearly cylindrical, in female swollen at base above.The
tibiaand
patella ofmale
palpusare subequal inlength,femur
very long.Twenty-two
specimensfrom Albemarle and Narboro
islands inFebruary and March.
Differsfrom
all described species in the armatureof themale
mandibles.Family THOMISIID.E.
TMARUS vSTOLZMANNI
Keyserling.(PI.II, fig. 5.)
Keyserling, DieSpinn. Amer., Laterigradse, p. 138 (1880).
Two
specimensfrom Albemarle
Island inMarch.
Describedfrom
Peru. It agrees with the description and figure, except that there isa broad
brown
stripeon
theventer of these specimens as usual in the genus.The
agreement of themale
palpusmakes
the identification certain.MISUMENA INCLUSA
sp. nov.Length
6mm.
(P'-'^ ^S- 12.)Cephalothorax pale yellowish, side of caput rather darker; eye region surrounded
by
a white line leaving only an opening behindbe-tween
the P.M.
E.; a faint whitemedian
lineon
caput; legs pale yellowish,unmarked,
spines black; sternum yellowish;abdomen
pale grayish above
and
below. Qtiadradrangle ofM.
E. slightly broader above, about as high asbroad above; eyesof posteriorrow
subequal in size;A.
S. E. larger thanA. M.
E.which
are about equal to P.M.
E., eyes of anteriorrow
at subequal distances; tibiae I and II scarcely as long as cephalothorax, beneath withtwo rows
of63
four spines; metatarsi Iand
II with five pairs beneath;none on
thefemora
;abdomen
convex, broadestbeyond
the middle.The male
has abrown
stripeon
eachupper
side of the cephalo- thorax; eye region as in female; legs Iand
II with the greater part of patella:, apical third of tibiae, apical two-thirds of metatarsi,and
apical half of tarsi,red-brown
; a red-brown stripe on each side ofabdomen, and
a series ofmedian marks
above.One
pairfrom Tagus
Cove,Albemarle
Island, inMarch.
Family SPARASSID^.
HETEROPODA VENATORIA
Linn.LiNNE, Syst. Nat., Ed. x, p. 1037(1758).
Aranearegia Fabricius, Syst. Ent., 11, p. 408 (1793).
Six specimens
from
Charlesand Chatham,
inMay. A common
and
widely distributed tropical spider; itsnorthward
range extends through the extreme southern partsof the United States.SELENOPS GALAPAGOENSIS
sp. nov.(PI. I, fig.8.)
Length
12mm.
Cephalothorax reddish; side-margins, dorsal groove
and around
eyes,black; mandibles red-brown; legspale, metatarsi black,
femora and
patelljEmarked
above with blackish bands, thoseon
tibiae are often darker, confluent,and
extend around the joint; sternum paler, apparently slightly bifid at tip;abdomen
palebrown
above, paler be- low, above with blackishmarkings more
or lessplain, a basal spear-mark and
side spots.The epigynum
differsmuch from
S. aissaWalck, and
S. spixi Perty.Six specimens
from Chatham and Albemarle
in Febi'uary.OLIOS GALAPAGOENSIS
sp. nov.(PI. I,fig. 7.)
Length
16mm.
Cephalothoraxyellowish, darkest in front; mandibles black; legs yellowish, metatarsi
and
tarsibrown;
sternum pale yellowish;max-
illae
and
lips, except tips, darkbrown
;abdomen brownish
above, with a basal darker spear-mark,margined
with pale spots; venter pale, with amedian brown
stripe.Extremely
similar to O. fascicu- latusofMexico and
the southwestern United States. Differs in the rather smaller size,and
proportionatelylonger legs,most
noticeablewhen
coinparing legIIwhere
the tibiapluspatella of O.fasclculatusis only equal to the tibia of O.galapagoe^isis.
64 BANKS
Fourteen specimens
from Chatham, Albemarle and Narboro
inJanuary and March.
Family CTENID^.
ODO INSULARIS
sp. nov.(PI. I,fig. 14; PI.n,fig. 13.)
Length 917 mm., S
13mm.
Cephalothorax yellowish
brown,
with a broad black stripeon
each side, leavingan
equally broadmedian
area, side-margins black, eye- regionblack; mandiblesblack,red-brown on
tips; legsyellow-brown, the tibiae with blackbands
atbaseand beyond
middle; palpiblackishon
lastjoint;sternum
red-brown, blackishon
sides;abdomen
black- ish above,withan irregular pale centralarea,brokenintospotsbehind;
venterpale, with a
few
scattered dark dots.Eyes
as figured for O.agilis; mandibles long; lip broader than long; sternum broad; legs long
and
stout, three pairs of spines under tibiae Iand
II,two under
these metatarsi, several spines aboveon
tibiae.Legs
ofmale
very slender.A few
specimensfrom Tagus
Cove,Albemarle
Island, inFebruary and March.
ODO GALAPAGOENSIS
sp. nov.Length
12mm.
(PI. 11,fig. 2.)Cephalothoraxclearyellowish,black side-margin; eye-region black,
and
a broad black stripeon
each side, leaving a broad areawhich narrows
behind head, then suddenlywidens
to surround the black-margined
dorsalgroove; mandiblesred-brown
; sternumyellow-brown
;legs yellow-brown, usually with apical
bands on
femora,and
basaland
preapical oneson
tibiae;dorsum
ofabdomen
mottled with grayand
black,below
pale, with afew
black dots.The
S. E. of pos- teriorrow
arenot sofarback
asusual,the S. E.of lowerrow
are rather oblong, thetwo rows
rather closertogetherthanusual; lipbroader than long; sternum broad; legs slender, buttwo
pairs of spinesunder
tibiae I
and
II,and two under
these metatarsi;no
spines aboveon
anteriortibiae, but severalon
posterior.A few
specimensfrom Hood
inMay, Tower
inJune,and Chatham
in
May.
Family LYCOSID^.
LYCOSA GALAPAGOENSIS
sp. nov.Length
12mm.
(^'^te 11, Fig.3.)Cephalothorax yellowish
brown,
witha palestripethrough themid-dle,
narrow on
eye-region,widened
behind, then suddenlyconstricted,then slightly
widened
again, then taperingto hindmargin;
the lateral margins irregularly pale; eye-region black; mandibles dark red-brown;
palpi yellow-brown, the last joint blackish; legs yellow-brown,
usuallyshowing
faintdarkerbands on femora and
tibiae,espe- cially of those of the hind pairs, the tarsi of anterior pairs darker;sternum
and
maxillae reddishbrown; abdomen
blackish above,witha spear-mark at base,margined
each sideby
pale; venter pale. Ante- rior eye-row nearly straight, theM.
E. about one-half their diameter apart, slightlyfartherfrom
thesubequal S. E., eyesof secondrow
less than diameter apart, eyes of thirdrow
equal to those of second; legs moderately stout,on
legI three pairsof spines vmdertibiae, the basal pairaslong aswidth of joint,two
pairsunder
metatarsus I.Several specimens
from
Iguana Cove,Albemarle
Island, inDecem-
berand June
;and Chatham,
inMay.
LYCOSA ALBEMARLENSIS
sp. nov.(Platen,Fig. 10.)
Length
12mm.
Cephalothorax
brown
; anarrow median
pale stripe starting in eye regionand
reaching hind margin, of nearlyevenwidth
throughout,and
a submarginal pale stripe each side; in thebrown
neartheme-
dianstripeisanindistinctoblongpale spot;eyeregionblack; mandibles lightyellow-brown; legsand
palpibrownish
yellow, darkeron
tips, rarelywith indistinctbands; sternumand
maxillaeyellowish;abdomen
black, a spear-mark atbase,
margined by
pale,which
behindisbroken
into spots; venter pale. Anterior eye-row slightly recurved,
M.
E.less than one half their diameter apart, plainly farther
from
the slightly smaller S. E.; eyesof secondrow
less than their diameter apart; legs of moderatesize, tibiaIwiththree pairsof spines beneath, the basal pairsmuch
longerthan width of tibia;metatarsus Iwithtwo
pairsoflong spineson
basal halfbeneath.Several specimens, all
from
Albemarle, in January.LYCOSA SNODGRASSI
sp. nov.(PlateII, Fig. i.)
Length
$ 16mm.,
$ 13mm.
Cephalothorax
brown,
with a ratherbroadpalemedian
stripe, start- ingas a linebetween
eyes of the secondrow
in front; dorsal groove, with a small indentation each side, a submarginal pale stripe each side quite close to the margin; eye regionand
mandibles black;legs
and
palpibrownish yellow, darkeronmetatarsiand
tarsi; sternumand
maxillae dark;abdomen
darkbrown
above, in female scarcelyProc.Wash.Acad. Sci., March, 1902.
66 BANKS
marked,
inmale
with a large basalpale area containinga dark spear-mark;
venter pale or lightbrown.
Anterioreye-row
slightly pro- curved, the AI. E. less than one-half their diameter apart,much
farther
from
the plainly smaller S.E.; eyesof secondrow
a little less than diameter apart; legs stout;on
tibia I three pairs of spines beneath, the basal pair (in the female)much
shorter than width of thetibia, inmale
longer;on
metatarsus Itwo
pairs of shoi't spines, inmale
long ones.A few
specimensfrom Albemarle
in June,Hood
inMay, and Chatham
inMay.
Family ATTID^.
PLEXIPPUS PAYKULLI Aud.
et Sav.AuDOUiN
etSavigny, Descr. d. I'Egypte, xxii, p. 172 (1827).A//USligo
Walckenaer,
Ins. Apteres, i, p. 426 (1837).Two
specimens, onefrom Tagus
Cove, Albemarle,March,
the otherfrom James
inApril.A common
tropical spider, extendingnorthward
into the southern parts ofthe UnitedStates.MARPTUSA CALIFORNICA Peckham.
Peckham,
Attidaeof N.Amer., p. 81 (1888).One male from
Albemarle, inMarch. Does
not appear to differfrom
typical Californian specimens.Known
alsofrom Mexico.
CYRBA INSULARIS
sp. nov.Length
6.7mm.
Cephalothoraxblack,
abdomen
dullblackaboveand
below,no
trace of thewhite lines seen in C. tce?tiola^ butventer with arow
of pale dots each side; legsbrownish
yellow, the first pair darkest,thoughnone
are blackon any
of the joints; palpi pale; sternum reddishbrown.
Cephalothorax flat, similar to C. tceniola^ as is the structure throughout.Epigynum shows
a pale cavity nearly asbroad as long, traversedby two
red parallel lines connected behind.One specimen from Tagus
Cove,Albemarle
Island, in January.ADMESTINA INSULARIS
sp. nov.(PlateII, Fig. 4.)
Length
3.8mm.
Black, with black, white
and
yellow hairs, thewhiteand
yellow slightly scale-like; cephalothorax with mostly black hair, anarrow
white side-margin;abdomen
aboveand below
sparsely, butregularly clothed with white hairsand
afew
yellow oneson
middle of sidesabove, not
enough
toform markings
; legs black, the coxae, patellaeand
aband on
tibiaeand
tarsiof hind pairs pale,and
here clothedwith white hair;some
whitehair aboveon
basal jointsof palpi. Cephalo- thoraxlong,narrow and
flat, thoracic part scarcelywidened
; anterioreye-row
slightly up-curved, the eyes of secondrow
rather nearer to thoseof the thirdrow
than to lateralsof first row.Legs
short, I the stoutest, especially the tibia,which
is sparsely clothed with long fine blackhairs; legs without spines, excepttwo on
anterior metatarsiand some weak
ones at tips of tibiaa IIIand IV; some
stiff bristles aboveon
the femora.Lip much
longer than broad; anterior coxae sepa- rated by width of lip, hind coxae contiguous,abdomen
long, slender, depressed; spinnerets distinct.One male from Mangrove
Point,Narboro
Island, in April.Appears
to beas close toAdmestina
as toany genus
yet described.PHIL^US PACIFICUS
sp. nov.Length 4 mm.
Cephalothorax reddish
brown,
darkest in eye region, inmale
thereis a white stripe eachside of eye region, but no trace of white
median
spot; legsand
mandibles yellowishbrown,
leg I quitedark,hind pairs often pale;abdomen margined
with a whitish stripe; in female pale above, with four pairs of black spots; inmale
shiningbrown,
with three pairsof white dots. Mandibles ofmale
elongateand
porrect.Similar instructure to
P.
militarise but smaller.Five specimens
from Albemarle and Narboro
inJanuary and
April.Order ARTHROG ASTRA.
PHRYNIDA.
CHARINUS INSULARIS
sp. nov.(PI. II,fig. 8.)
Length
9mm.
Cephalothorax, mandibles
and
palpired-brown
; legsmore
yellow- ish;abdomen
lightbrown
above; coxaeand
venter paler.Cepha-
lothorax very broadand
short; palpi of usual length;femur
in frontabove
with three short spines,below
three longer ones, the basalone
the longest; the tibia has above in front five spines, the first near the middle being short, the next three increasing in length,and
the fifth shorter than the second;below
there aretwo
spines, the apical the longer; thehand
hason
inner sidetwo
spines, the apicalmuch
the longer,and on
outersideneartiponespine; legs short, hindtibiaefour-68 BANKS
jointed,the basaljointaslongasthethree others; metatarsus aslongas
first jointof tibia; tarsus tippedwith a pulvillus;
sternum
is rather small.Eight specimens
from
Albemarle, Narboro,Chatham, and Hood
in
March and May.
This genus is found only in Australia, in islands of thePacific,
and
insouthern Asia.
SCORPIONIDA.
HADRUROIDES LUNATUS Koch.
Koch, Verb. zool. hot. Ges., Wien, p. 235 (1867.)
H. maculatus
Thorell,
Atti Soc. Ital., xix, p. 186 (1877).H. charasus KARSCH.'Mitt. Miinch. Ent. Ver., iii, p. 135 (1879).
Nineteen specimens
from
Albemarle,James,
Barringtonand Nar-
boro, inMarch,
Apriland May.
Distributed alongthe coast regionof western South
America.
CENTRURUS PRINCEPS
Karsch.Karsch, Mitt. Miinch. Entom. Ver., in, p. 121 (1879).
Kraepelin, Mitt. Mus. Hamburg, viii, p. 139 (1891).
Six specimens
from Hood and Chatham
inMay.
Previouslyknown
onlyfrom
Hayti, but the specimens agree with this species in all important particulars.The hand
is hardly fuscous; thebody
is palebrownish,
sometimes
withtraces of transverse darkmarks;
onespecimen shows
the regionbetween
thesubmedian
ventral keels darker than elsewhere; there are ten full keelson
segments one to three;on
under side of fifthsegment
isamedian
keel, also a lateral keeland
asubmedian
keel,which
latter, however, is not complete but lost inthe granulationof the general surface; the tubercle under thestingis very small; the fingersshow
the lobeand
cavityas des- cribedby
Karsch.The
largestspecimen
is about44 mm.
long.PSEUDOSCORPIONIDA.
ATEMNUS INSULARIS
sp. nov.(PI. II, fig. II.)
Length
3.7mm.
Cephalothorax
and abdominal
scutoe yellowishbrown,
darkeron
anterior half of cephalothorax; palpi reddishbrown,
darkeston
fingers; legs pale. Cephalothorax
rounded
in front, onceand
one- third as longasbroad: at anterior third there is ashort transverse de- pression;on
anteriormargin
aretwo
white eye-spots.Mandibles
ARACHNIDA 69
small; lower finger with a minute styletneartip. Palpi large, stout;femur
broadest near base, convexon
inner side near base; tibia about equalto femur,convex
each side;hand
stout, tapering each side to thefingers,which
aremuch
shoiter than the hand.The
trochantinsshow
plainly in all legs.Abdomen
elongate, nearly three times as longas cephalothorax, the scutse entire;body and appendages
spar- ingly clothed with simple hairs.Seventeen specimens
from Albemarle
inJanuary and
February.CHELANOPS NIGRIMANUS
sp. nov.(PI. II, fig.6.)
Length
2.9mm.
Cephalothorax nearly black; palpi dark red-brown, black
on
hand, nearly soon
base offemur and
middle of tibia; legs pale greenishbrown
;abdominal
scutae reddishbrown.
Cephalothoraxmuch
taper-ing in front, with aneye-spot
on
each side; palpi moderately longfemur
subcylindric, aboutas longascephalothorax isbroadin middle;tibia fully as long as femur, long pedicellate, inner
margin
slightly concave before tip;hand
nearly twice as broad as femur,rounded
at base, tapering each side to fingers,which
ai'e plainly,though
not greatly, shorter than the hand, and slightly curved;on
the trochanter,femur and
tibia the hairs are shortand
thick, but not plainly clavate similar hairsborder theabdominal scutge.One specimen from Albemarle
inMarch.
SOLPUGIDA.
AMMOTRECHA SOLITARIA
sp. nov.(PI-i, fig. 5-)
Length, without mandibles, 10
mm.
Cephalothorax uniform dull
brown
; mandiblesbrown
above; fin- gersred-brown
; legsand
palpi pale yellowish, the femora, tibiaeand
base of metatarsi of hind pairs blackishbrown
;abdomen uniform
dull
brown
above, palerbeneath. Cephalothorax with amedian
fur- row, eye-tubercleslightlyelevated; eyes scarcelytheirdiameterapart palpi shortand slender, the tibia equal to femur, the metatarsus one- half the length of the tibia, the tarsus two-thirds the lengthof the metatarsus.The movable
fingerof themandibleshastwo
largeteeth,and
one small tooth at the base of the first large one; theupper
finger hasthree small teeth, then a large one with a small oneon
itsouter base, then
two
rather small teeth; there is a small ridge aboveon
the fingrer.yo BANKS
One specimen from Iguana
Cove,Albemarle
Island, inDecember.
Order A CARINA.
Family IXODID.^.
AMBLYOMMA PILOSUM Neumann.
Neujuann,
Mem.
Soc. Zool. France, p. 209 (1899).Many
specimensfrom Albemarle
inMarch
; onefrom Narboro
in April. Describedfrom
theGalapagos
Islandsfrom
aspecimen
intheMarx
collection.ARGAS TRANSVERSA
sp. nov.(PI. II, fig. 9.)
Length
i.imm.; width
2.1mm.
Blackish, legs paler.
Body
flattened, nearly twice as broad as long;around
themargin
arerows
of tubercles, each bearing a shortstiff bristle, the
upper row
with the longestbristles; the surfaceaboveis traversed
by many
irregularlines, dividing the surface intoirregular flattened elevations; amedian and two
lateral depressionsand two
behind;below
granulateon
sides; behind the legs the surface isclothed with yellowish sharp denticles; legs short, scarcely visible
from
above, hairy; mouth-parts small.One specimen from Tagus
Cove,Albemarle
Island, in February.Easily recognized
by
shape of body.Family TYROGLYPHID^.
TYROGLYPHUS
sp.A few
specimensfrom Narboro
Island, inMarch. Not
sufficientmaterial for determination.
*J1
PART
II.field notes on species described
inpart
i.By Robert
E.Snodgrass.
SICAROIDES ULTRIFORMIS.
Rare;
secured at only one place in the archipelago, viz.,under
rockson
flat area at top of cliffswestofGardner
Bay,Hood
Island.The
soil at this placeand
theunweathered
undersides of the surface rocks are of a bright brick-red color. All the individuals of this spider seenwere
also of thissame
color.They
were, accordingly, inconspicuous so long as theyremained
quiet. Several other nor- mally colored species occurredat thesame
place.The
eggs are inclosed ina cup-shaped case,havinga flattopwith a projecting rim (Plate iii, fig. 9).A
nest secured iscomposed
of fine particles of earth held together in a delicatenetwork
of threads (Plate III, fig. 10), having, consequently, thesame
color as the rock towhich
itwas
attached, and,were
it not forits extraordinary shape,would
appear to be a part of it.The
nestwas
taken inApril. InSeptember numerous young
spidershad emerged
through a transverse slit-likeopening nearthe top, extendingso fararound thatthetopwas
almost separatedfrom
the rest like a lid.LOXOSCELES LONGIPALPIS.
Allthe
Hood
Island specimenswere
taken at thesame
localityas the last species.They
occurredin the crevices of rocks lyingon
the surface.Some were
withoutwebs, othershad
an irregularonespun
aboutthem
in the recesses of the rock.CORYSSOCNEMIS CONICA.
Found
onlyon Hood
Island, under rocks at top of cliffswest ofGardner
Bay, along with lasttwo
species. Spins an irregularweb on
theunder
surfaces of rocks. Exceedingly swift in itsmovements.
On
disturbing one it first darted out of itsweb,
then as quicklyback
again, and, clingingto the center, vibrated it so rapidly asto be almostinvisible.AYSHA PACIFICA.
Rare
in the archipelago.The Albemarle
specimenswere
both taken in the Turtle Pointmangrove swamp. One was
secured in a netby
beating the leaves of the trees, the otherwas
taken with its72 SNODGRASS
cocoon withinthe cavity of a dead pendent
mangrove
leaf havingthetwo
edges rolled together (Plate in, fig. 2).The Chatham
speci-men was
takenfrom
an exposedcocoon
containing the adultand numerous young
ones.DICTYNA PARIETALIS.
Only
onespecimen
seen.Taken
atTagus
Cove,Albemarle
Island,from
irregularnetwork
of straight threadsspun
about theend
ofan
acacia twig.THERIDIUM MIXTUM.
Found
only on Albemarle,Narboro and James
Islands.On
Albe- marle the specieswas
found onlyon
the dryer parts ofthe island. Itwas abundant
in the dry, brushy region surroundingTagus
Cove, butnone were
found at IguanaCove
orinthemangrove swamps. Of
thetwo Narboro
specimens one isfrom
the southeast side of the island atanelevationof about 1,500 feet,the otheris
from
themangrove swamp
at
Mangrove
Point.The James
specimenswere
taken atJames
Bay.The webs
constructedby
this specieson
theGalapagos
Islands gen- erally consist of a largeand
irregular netsvork of threadsspun
across one another in all directions, most frequently attached to the side of the trunk of aPalo Santo tree (theonly tree of the islands) or sus-pended between
the trunks oftwo
neighboringtrees.One was
found in the hollow of adead Palo Santo stump. Often theweb
has theform
of a flat sheet.Two
specimens takenatTagus Cove had
appar- entlyappropriated desertedwebs
ofEpeira
labyrinthea^ since theywere
foundinwebs
consisting inpartof anetwork
likethatspun
nor- mallyby
theirown
species, but having connected with it also a reg- ularorb.The
spider remains in the center of theweb
concealed beneath a shelterformed
of bits of foreign material generally takenfrom
theimmediate
vicinity,and
resemblinganaccidental accumulationof such matter.For
example, hiding places ofwebs
in acacia busheswere
generallymade
of bitsof acacia leaves; the hiding place of one sus-pended
inthehollowofadeadstump was made
of piecesofdeadwood,
while hiding places ofwebs
on the sides of trees frequently had bits of bark in them.The
onespecimen from James
Islandhad
its hiding placeformed
of a small sheet of closely-spun silk in the center of theweb,
togetherwith afew
bits of dead twigsand
a driedleaf. In one case the spider had for its nest a curled leaf bent into a U-shape, sus-pended
at the center of theweb
with thearms
hangingdownward
(Plate III, fig. 6).