ANGIOSPERM DIVERSITY OF MARIHAN REGION OF MIRZAPUR DISTRICT UP Dr. Sarvajeet Singh
Dept. of Botany, Assistant Professor, Rajkiya Mahila Mahavidhyala, Shahganj, Jaunpur UP
Abstract- The present study provides a taxonomic account of the angiosperms of Marihanregion of Mirzapur District in Uttar Pradesh. The region, which comes under the Vindhyan region of Uttar Pradesh, is one of the richest areas in the state as far as plant diversity is concerned. It is spread over about 156466.26ha of geographical area, which constitutes about 36% forest cover on highly undulated land. The extensive survey of the area conducted during 2011-2016, critical examination of previous collections housed at various herbaria, and review of published literature have resulted in a total of 705 species belonging to 459 genera under 110 families. Out of these, 541 species (76.73%) under 354 genera (77.12%) and 89 families (80.90%) belong to dicots, and 164 species (23.26%) under 105 genera (23.26%) and 21 families (19.09%) to monocots. The present enumeration of the species also includes about 78 species cultivated in the area for various purposes. Fabaceae (110 spp.) comprises of the maximum number of species, followed by Poaceae (89 spp.), Asteraceae (38 spp.), Cyperaceae (33 spp.), and Malvaceae (33 spp.). Some of the largest genera in the area are Cyperus (14 spp.), Ipomoea (9 spp.), Solanum (9 spp.), Ficus (9 spp.), Crotalaria (7 spp.), Desmodium (7 spp.), Bauhinia (6 spp.), Hibiscus (6 spp.), Fimbristylis (6 spp.), Acacia (5 spp. ), etc. The entire forest is chiefly dominated by trees such as Acacia catechu (L.f.) Willd., Boswellia serrata Roxb. ex Colebr., Butea monosperma (Lam.) Taub., Hardwickia binata Roxb., and Shorea robusta Gaertn.
Keywords: Floristic diversity, Vindhyan region, Marihan region of Mirzapur District, Uttar
Pradesh.
1 INTRODUCTION
The floristic account provides lots of information of plant wealth of an area, which may be beneficial for university students, botanists, researchers, NGOs, naturalists, environmental engineers, forest managers, conservation biologists, and policy makers. Knowledge of floristic composition and structure of forest is also useful in identifying ecologically and economically important plants, analysing their diversities, and protecting threatened plants (Addo-Fordjour et al.
2009). In the past, many important floristic accounts have been published for several states, districts, and biogeographic regions of India and abroad. In addition to general flora of Uttar Pradesh (Duthie 1903-1929;
Kanjilal 1966; Rau 1969; Uniyal et al.
1999; Chaudhary et al. 2016), several works related to different districts, national parks, and sanctuaries have also been produced by many (Kanjilal 1933;
Srivastava 1938; Singh 1969; Srivastava 1976; Sharma & Pandey 1984; Sharma &
Dhakre 1995; Verma & Ranjan 1995;
Singh 1997; Saini2005; Singh & Khanuja 2006; Kumar et al. 2015). The floristic account of Marihan region of Mirzapur District of Uttar Pradesh, however, is still unexplored in spite of its rich diversity.
The production of state flora will be easier, once all the district floras are worked out systematically. Hence, there is a need for proper inventory and documentation of all plants available in Marihan region of Mirzapur District with their up to date taxonomic information.
Since the area is very rich in minerals and forest resources, many industries and factories of different kinds have been set up here. Therefore, the present study will also be very important for a comparative study of the plants of the area in future and also to examine the effects of industrial pollutants on them.
Although the district has not been thoroughly inventoried earlier, the reference on the plants of the study area is available in Bhattacharyya (1963, 1964) and Srivastava (1955) wherein the plants of erstwhile Mirzapur District have been dealt with. In these studies, about 460 species have been listed including about 130 species from the study area. In addition, a few scattered works chiefly pertaining to ethnobotanical and medicinal plants have been carried out in the area (Singh et al. 2002; Chaudhary 2010; Singh et al. 2010; Singh et al.
2012; Singh & Dubey 2012; Mishra et al.
2012). Recently, Kushwaha et al. (2016 a,
b) have thoroughly studied the family Cucurbitaceae of the area and have also presented a preliminary overview of angiospermic plants. Hence, the present work is the first study of its kind from the study area after its recognition as a region.
2 MATERIALS AND METHODS
In this study, 12 field tours in different seasons were conducted between 2011- 2016 to survey the area. The plant materials were collected either in flowering or fruiting or in both stages with detailed information such as habit, habitat, colour of flowers and fruits, shape and size of trees, nature of bark of trees, GPS information, conservation status, etc. The herbarium specimens were prepared following the standard procedure outlined by Lawrence (1951) and Jain & Rao (1977). The identification of species was done with the help of different floras and important taxonomic works (Hooker 1872-1897; Duthie 1903- 1929; Brandis 1906; Singh 1997) and also after matching with the authentic specimens housed at various Indian herbaria such as Botanical Survey of India, Allahabad (BSA), Botanical Survey of India, Dehra Dun (BSD), Botanical Survey of India, Kolkata (CAL), Forest Research Institute, Dehra Dun (DD), and National Botanical Research Institure, Lucknow (LWG). All specimens collected in the presentstudy have been deposited at LWG for future records. In the present treatment, each species was furnished with the correct recent name following websites such as The International Plant Names Index (IPNI), Germplasm Resources Information Network (GRIN), International Legume Database and Information Service (ILDIS), The Plantlist, Wikipedia, and Tropicos, along with habit,
distribution and reference to voucher specimens. The families, genera, and species within the genus have been arranged alphabetically in the list.
2.1 Study Area
The study was carried out in Madihan forest range in Mirzapur district of Uttar Pradesh (state) of India which lies between 25°0'8.83"N to 24°56'37.61"N and 82°37'31.91"E to 82°45'59.10"E.
The total study area analyzed is 156,466.26 ha and the forest area is 31,380.03 ha. The climate here is characterized by three distinct seasons, the summer season (March - June) is hot and dry when temperatures can reach as high as 480C; warm and humid rainy season (July-October) where rainfall varies from 90-100cm, cool dry winter (November – February) when temperatures can be as low as 60C (Singh 2007). The soil varies from loamy sand to clay. Variation in colour of soil is observed from red to yellow (Das et al.
2007). The terrain is undulating with
hillocks dominating. (Figure 1 –
location of the study area). The forests of
this region are best described as
tropical dry deciduous forest (Champion
and Seth 1968). Large areas are of 5A/C3
and 5B/C2 types of forests, i.e., Southern
and Northern Tropical Dry Deciduous
Forests respectively. The flora is
dominated by tree species such as
Shorea robusta, Boswellia serrata, Anogeissus latifolia, Terminalia arjuna, Lagerstroemia parviflora, Hardwickia binnata, Acacia catechu, Butea monosperma, Zizyphus glaberrima, Diopsyros melanoxylon. Pterocarpus marsupium, Holorrhena antidysentrica, Emblica officinalis, Synzygium cumini, Tectona grandis, Madhuca longifolia.3 RESULTS AND DISCUSSION
The whole of the area is covered with natural scrubby jungle and thick forests.
Almost the entire plateau exhibits a uniform horizontal stratification of rocks.
The top of the plateau is unsuitable for the growth of broad-leaved plants and, is represented by uniform scrubby dry vegetation. The broad-leaved plants are generally seen on the slopes of the plateau. Overall, the entire forest is mixed dry deciduous, but evergreen trees are frequently seen in ravines. The data collected fromfield and herbarium studies reveal that the entire area contains about 705 species under 459 genera and 110 families including 78 cultivated ones that have been planted in the area for different purposes (Table 1; Images 2-3). Out of these, 541 species (76.73%) under 354 genera (77.12%) and 89 families (80.90%) belong to dicots and 164 species (23.26%) under 105 genera(22.87%) and 21 families (19.09%) to monocots (Fig. 2).
The family Fabaceae has the maximum number of species (52 Genera, 110 species), followed by Poaceae (63 Genera, 89 species), Asteraceae (32 Genera, 38 species), Malvaceae (15 Genera, 33 species), Cyperaceae (9 Genera, 33 species), Cucurbitaceae (11 Genera, 20 species), Acanthaceae (11 Genera, 17 species),Convolvulaceae (8 Genera, 17 species), Solanaceae (5 Genera, 16 species), and Rubiaceae (13 Genera, 15 species) (Fig. 3). Some of the dominant genera in the forest are Cyperus (14 spp.), Ipomoea (9 spp.), Solanum (9 spp.), Ficus (9 spp.), Crotalaria (7 spp.), Desmodium (7 spp.), Bauhinia (6 spp.), Hibiscus (6 spp.), Fimbristylis (6 spp.), Acacia (5 spp.), etc. Among all plants, 443 species(62.70%) are herbs, 163 species (23.12%) are trees, 75 species (10.21 %) are shrubs, and 29 species (3.97 %) are climbers (Fig. 4).
Marihan Forests
The opening of forests in Marihan
Figure 1 Study site: Marihan region of Mirzapur District Uttar Pradesh.
Source: India map from Google and Marihan region of Mirzapur map from:
https://www.google.co.in/
maps/place/Marihan region of Mirzapur,+Uttar+Pradesh.
According to a revised classification of
forest type in India by Champion & Seth
(1968), the forest type of Marihan region
of Mirzapur District is tropical dry
deciduous. The whole area of the region
has an interesting diversity of flora and
vegetation due to variable plains, slopes,
hills, and climate, represented by dry
deciduous vegetation, natural scrub
jungle, patches of grasses, and thick
forest. The whole area of the regsion is
covered with rich plant diversity with
trees at the top layer, shrubs at the
middle layer, and herbs, climbers, and
twiners at the ground level.
Family Taxa Growth form DICOT
Acanthaceae Adhatoda zeylanica Medik. Shrub
Andrographis echioides (L.) Nees Herb
Andrographis paniculata (Burm. f.) Wall. ex Nees Herb
Barleria cristata L. Shrub
Barleria prionitis L. Shrub
Dipteracanthus prostratus (Poir.) Nees Herb
Elytraria acaulis (L.f.) Lindau Herb
Hemigraphis hirta (Vahl) T.Anderson Herb Hygrophila auriculata (Schumach.) Heine Herb Hygrophila erecta (Burm.f.) Hochr. Herb Hygrophila polysperma (Roxb.) T.Anderson Herb
Justicia diffusa Willd. Herb
Justicia prostrata Schltdl. ex Nees Herb Justicia quinqueangularis K.D.Koenig ex Roxb. Herb Peristrophe bicalyculata (Retz.) Nees Herb
Ruellia suffruticosa Roxb. Shrub
Rungia pectinata (L.) Nees Herb
Aizoaceae Glinus oppositifolius (L.) Aug. DC. Herb
Trianthema portulacastrum L Herb
Ampelidaceae Ampelocissus latifolia (Roxb.) Planch. Climber
Amranthaceae Achyranthes aspera L. Herb
Aerva lanata (L.) Juss. Herb
Allmania nodiflora (L.) R.Br. ex Wight Herb
Alternanthera ficoidea (L.) Sm. Herb
Alternanthera paronychioides A.St.-Hil. Herb Alternanthera sessilis (L.) R.Br. ex DC. Herb
Amaranthus spinosus L. Herb
Amaranthus tricolor L. Herb
Amaranthus viridis L. Herb
Celosia argentea L. Herb
Digera muricata (L.) Mart. Herb
Nothosaerva brachiata (L.) Wight Herb
Anacardiaceae Buchanania cochinchinensis (Lour.) M.R. Almeida Tree Lannea coromandelica (Houtt.) Merr. Tree
Mangifera indica L. Tree
Semecarpus anacardium L.f. Tree
Annonaceae Annona squmosa L. Tree
Miliusa tomentosa (Roxb.) Sinclair Tree Polyalthia longifolia (Sonn.) Thw. Tree Polyalthia suberosa (Roxb.) Thwaites Tree
Apiaceae Anethum graveolens L. Herb
Centella asiatica (L.) Urban Herb
Coriandrum sativum L. Herb
Family Taxa Growth form
Apocynaceae Alstonia scholaris (L.) R. Br. Tree
Calotropis gigantea (L.) R. Br. Small tree Calotropis procera (Aiton) Dryand. Shrub
Carissa carandas L. Small tree
Cascabela thevetia (L.) Lippold Tree
Carissa spinarum L. Shrub
Catharanthus pusillus (Murray) G.Don Herb Holarrhena pubescens Wall. ex G.Don Tree Ichnocarpus frutescens (L.) W.T.Aiton Climber Oxystelma esculentum (L.f.) Sm. Climber Tabernaemontana divaricata (L.) R.Br. ex Roem.
& Schult Tree
Wrightia arborea (Dennst.) Mabberley Small tree
Wrightia tinctoria R.Br. Tree
Plumeria rubra L. Tree
Plumeria alba L. Small tree
Aristolochiaceae Aristolochia indica L. Herb
Asclepiadaceae Ceropegia hirsuta Wight & Arn. Climber Cryptolepis dubia (Burm.f.) M.R.Almeida Climber
Gymnema rivulare Schltr. Climber
Hemidesmus indicus (L.) R. Br. ex Schult. Herb Marsdenia tenacissima (Roxb.) Moon Woody climber Telosma pallida (Roxb.) W. G. Craib Shrub
Family Taxa Growth form
Asteraceae Acanthospermum hispidum DC. Herb
Acilepis aspera (Buch.-Ham.) H.Rob. Herb
Acmella calva (DC.) Jansen Herb
Ageratum conyzoides (L.) L. Herb
Artemisia nilagirica (C.B.Clarke) Pamp. Herb
Bidens pilosa L. Herb
Blainvillea acmella (L.) Philipson Herb
Blumea lacera DC. Herb
Blumea axillaris (Lam.) DC. Herb
Blumea fistulosa (Roxb.) Kurz Herb
Blumea laciniata (Wall. ex Roxb.) DC. Herb
Blumea oxyodonta DC. Herb
Caesulia axillaris Roxb. Herb
Carthamus oxyacantha M.Bieb. Herb
Chrysanthellum indicum DC. Herb
Conyza leucantha (D.Don) Ludlow & P.H.Raven Herb Cyanthillium cinereum (L.) H.Rob. Herb Cyathocline purpurea (Buch.-Ham. ex D.Don)
Kuntze Herb
Echinops echinatus Roxb. Herb
Eclipta prostrata L. Herb
Elephantopus scaber L. Herb
Emiliasonchifolia (L.) DC. ex DC. Herb
Gnaphalium polycaulon Pers. Herb
Family Taxa Growth form
Gnaphalium indicum L. Herb
Grangea maderaspatana (L.) Poir. Herb
Helichrysum indicum (L.) Grierson Herb
Ixeris polycephala Cass. Herb
Launaea acaulis (Roxb.) Babc. ex Kerr. Herb Launaea procumbens (Roxb.) Ramaya & Rajgopal Herb
Parthenium hysterophorus L. Herb
Pentanema indicum (L.) Ling Herb
Pluchea indica (L.) Less. Herb
Pulicaria crispa Sch.Bip. Herb
Sonchus asper (L.) Hill Herb
Sphaeranthus indicus L. Herb
Tagetes erecta L. Herb
Tridax procumbens (L.) L. Herb
Xanthium strumarium L. Herb
Basellaceae Basella alba L. Climber
Bignoniaceae Handroanthus impetiginosus (Mart. ex DC.)
Mattos Tree
Fernandoa adenophylla (Wall. ex G. Don) Steenis Tree
Jacaranda mimosifolia D. Don Tree
Kigelia africana (Lam.) Benth. Tree
Tabebuia rosea DC. Tree
Tecoma stans (L.) Juss. ex Kunth Tree
Bombacaceae Bombax ceiba L. Tree
Boraginaceae Cordia dichotoma G. Forst. Tree
Ehretia laevis Roxb. Tree
Heliotropium indicum L. Herb
Heliotropium ovalifolium Forssk. Herb
Heliotropium supinum L. Herb
Trichodesma indicum (L.) Lehm. Herb
Brassicaceae Brassica nigra (L.) K.Koch Shrub
Brassica rapa L. subsp. campestris (L.) Clapham Herb
Eruca vesicari (L.) Cav. Herb
Raphanus sativus L. Herb
Burseraceae Boswellia serrata Roxb. ex Colebr. Tree Caryophyllaceae Polycarpaea corymbosa (L.) Lam. Herb
Polycarpon prostratum (Forssk.) Asch. &
Schweinf. Herb
Spergula arvensis L. Herb
Vaccaria hispanica (Mill.) Rauschert Herb
Cactaceae Opuntia dillenii (Ker Gawl.) Haw. Shrub
Opuntia cochenillifera (L.) Mill. Shrub
Opuntia stricta (Haw.) Haw. Shrub
Family Taxa Growth form
Campanulaceae Lobelia alsinoides Lam. Herb
Cannabaceae Cannabis sativa L. Shrub
Celtis australs L. Tree
Celtis tetrandra Roxb. Tree
Family Taxa Growth form
Trema orientalis (L.) Blume Tree
Capparidaceae Capparis zeylanica L. Shrub
Casuarinaceae Casuarina equisetifolia L. Tree
Celastraceae Cassine glauca (Rottb.) Kuntze Tree
Celastrus paniculatus Willd. Shrub
Ceratophyllaceae Ceratophyllum demersum L. Herb
Chenopodiaceae Chenopodium murale L. Herb
Chenopodium album L. Herb
Dysphania ambrosioides (L.) Mosyakin &
Clemants Herb
Cleomaceae Cleome gynandra L. Herb
Cleome viscosa L. Herb
Combretaceae Anogeissus acuminata (Roxb. ex DC.) Wall. ex
Guillem. & Perr. Tree
Anogeissus latifolia (Roxb. ex DC.) Wall. ex
Guillem. & Perr. Tree
Terminalia elliptica Willd Tree
Terminalia arjuna (Roxb. ex DC.) Wight & Arn. Tree Terminalia bellirica (Gaertn.) Roxb. Tree
Convolvulaceae Convolvulus arvensis L. Herb
Cuscuta reflexa Roxb. Climber
Erycibe paniculata Roxb. Woody climber
Evolvulus alsinoides (L.) L. Herb
Evolvulus nummularius (L.) L. Herb
Ipomoea aquatica Forssk. Herb
Ipomoea batatas (L.) Lam. Climber
Ipomoea cairica (L.) Sweet Climber
Ipomoea carnea Jacq. Shrub
Ipomoea dichroa Choisy Climber
Ipomoea nil (L.) Roth Climber
Ipomoea obscura (L.) Ker Gawl. Herb
Ipomoea pes-tigridis L. Herb
Ipomoea quamoclit L. Herb
Merremia hederacea (Burm. f.) Hallier f. Herb Operculina turpethum (L.) Silva Manso Climber
Porana paniculata Roxb. Climber
Cornaceae Alangium salviifolium (L.f.) Wangerin Tree Cucurbitaceae Benincasa hispida (Thunb.) Cogn. Climber
Cayaponia laciniosa (L.) C.Jeffrey Climber Citrullus colocynthis (L.) Schrad. Herb Citrullus lanatus (Thunb.) Matsumara & Nakai Herb
Coccinia grandis (L.) Voigt Herb
Cucumis sativus L. Herb
Cucumis melo L. Herb
Cucumis melo L. subsp. agrestis (Naudin)
Pangalo Herb
Cucurbita maxima Duch. Herb
Diplocyclos palmatus (L.) C. Jeffrey Herb Lagenaria siceraria (Molina) Standl. Herb
Family Taxa Growth form
Luff a hermaphrodita Singh & Bandhari Herb
Luffa acutangula (L.) Roxb. Herb
Luffa cylindrica (L.) M.Roem. Herb
Momordica charantia L. Herb
Momordica dioica Roxb. ex Willd. Herb
Mukia maderaspatana (L.) M.Roem. Herb
Trichosanthes cucumerina L. Climber
Trichosanthes cucumerina L. var. anguina (L.)
Haines Climber
Trichosanthes dioica Roxb. Herb
Dipterocarpaceae Shorea robusta Gaertn. Tree
Ebenaceae Diospyros melanoxylon Roxb. Small tree
Diospyros montana Roxb. Small tree
Family Taxa Growth form Elatinaceae Bergia ammannioides Roxb. ex Roth Herb
Euphorbiaceae Acalypha ciliata Forssk. Herb
Baliospermum solanifolium (Burm.) Suresh Shrub Chrozophora rottleri (Geiseler) A.Juss. ex Spreng. Herb
Croton bonplandianus Baill. Herb
Euphorbia hirta L. Herb
Euphorbia hypericifolia L. Herb
Euphorbia prostrata Aiton Herb
Jatropha gossypiifolia L. Shrub
Mallotus nudiflorus (L.) Kulju & Welzen Tree Mallotus philippensis (Lam.) Mull.Arg. Tree Microstachys chamaelea (L.) Mull.Arg. Herb
Ricinus communis L. Shrub
Fabaceae-
Caesalpinioideae Bauhinia malabarica Roxb. Tree
Bauhinia purpurea L. Tree
Bauhinia racemosa Lam. Tree
Bauhinia semla Wunderlin Tree
Bauhinia vahlii Wight & Arn. Tree
Bauhinia variegata L. Tree
Cassia fistula L. Tree
Chamaecrista absus (L.) H.S.Irwin & Barneby Tree Chamaecrista mimosoides (L.) Greene Herb
Delonix regia (Hook.) Raf. Tree
Hardwickia binata Roxb. Tree
Parkinsonia aculeata L. Small tree
Peltophorum pterocarpum (DC.) K.Heyne Tree Piliostigma malabaricum (Roxb.) Benth. Tree
Senna alata (L.) Roxb. Shrub
Senna occidentalis (L.) Link Herb
Sennasiamea (Lam.) H.S.Irwin & Barneby Tree
Senna tora (L.) Roxb. Herb
Family Taxa Growth form
Fabaceae-
Mimosoideae Acacia auriculiformis Benth. Tree
Acacia catechu (L.f.) Willd. Tree
Acacia nilotica (L.) Delile Tree
Acacia pinnata Link Tree
Acacia torta (Roxb.) Craib Shrub
Albizia lebbeck (L.) Benth. Tree
Albiziaprocera (Roxb.) Benth. Tree
Albizia odoratissima (L.f.) Benth. Tree Dichrostachys cinerea (L.) Wight & Arn. Tree Leucaena leucocephala (Lam.) de Wit Tree
Mimosa himalayana Gamble Small tree
Mimosa rubicaulis Lam. Small tree
Pithecellobium dulce (Roxb.) Benth. Tree
Prosopis cineraria (L.) Druce Tree
Prosopis juliflora (Sw.) DC. Small tree Fabaceae-
Papilionoideae Abrus precatorius L. Climber
Aeschynomene indica L. Herb
Atylosia scarabaeoides (L.) Benth. Herb
Alhagi maurorum Medik. Shrub
Alysicarpus bupleurifolius (L.) DC. Herb
Alysicarpus glumaceus (Vahl) DC. Herb
Alysicarpus hamosus Edgew. Herb
Alysicarpus monilifer (L.) DC. Herb
Alysicarpus vaginalis (L.) DC. Herb
Butea monosperma (Lam.) Taub. Tree
Cajanus cajan (L.) Millsp. Shrub
Cajanus platycarpus (Benth.) Maesen Herb Cajanus scarabaeoides (L.) Thouars Herb
Cicer arietinum L. Herb
Clitoria ternatea L. Herb
Codariocalyx motorius (Houtt.) H.Ohashi Shrub
Crotalaria albida Roth Shrub
Crotalaria spectabilis Roth Herb
Family Taxa Growth form
Crotalaria juncea L. Shrub
Crotalaria medicaginea Lam. Herb
Crotalaria mysorensis Roth Herb
Crotalaria prostrata Willd. Herb
Crotalaria quinquefolia L. Herb
Daldergia sisso DC. Tree
Dalbergia lanceolaria L.f. Tree
Dalbergia latifolia Roxb. Tree
Desmodium oojenenensis (Roxb.) H. Ohasi Tree Desmodium dichotomum (Willd.) DC. Herb
Family Taxa Growth form
Desmodium polycarpum (Poir.) DC. Herb
Desmodium gangeticum (L.) DC. Herb
Desmodium heterocarpon (L.) DC. Shrub
Desmodium triflorum (L.) DC. Shrub
Desmodium velutinum (willd.) DC. Herb
Erythrina stricta Roxb. Tree
Erythrina suberosa Roxb. Herb
Flemingia strobilifera (L.) W.T.Aiton Shrub
Indigofera astragalina DC. Tree
Indigofera cassioides DC. Tree
Indigofera hirsuta L. Herb
Indigofera linifolia (L.f.) Retz. Herb
Indigofera linnaei Ali Shrub
Indigofera tinctoria L. Shrub
Indigofera trita L.f. Herb
Lablab purpureus (L.) Sweet Herb
Lathyrus aphaca L. Herb
Lathyrus sativus L. Herb
Medicago polymorpha L. Herb
Melilotus indicus (L.) All. Herb
Millettia extensa (Benth.) Baker Herb
Millettia peguensis Ali Tree
Mucuna pruriens (L.) DC. Climber
Phyllodium pulchellum (L.) Desv. Shrub
Pongamia pinnata (L.) Pierre Tree
Pterocarpus marsupium Roxb. Tree
Rhynchosia capitata (Roth) DC. Herb
Rhynchosia minima (L.) DC. Herb
Sesbania bispinosa (Jacq.) W.Wight Herb
Sesbania sesban (L.) Merr. Shrub
Smithia conferta Sm. Herb
Tephrosia pumila (Lam.) Pers. Herb
Tephrosia purpurea (L.) Pers. Herb
Tephrosia strigosa (Dalzell) Santapau &
Maheshw. Herb
Teramnus labialis (L.f.) Spreng. Herb
Trifolium alexandrinum L. Herb
Trigonellafoenum-graecum L. Herb
Uraria lagopodoides (L.) DC. Herb
Uraria lagopus var. neglecta (Prain) H.Ohashi Herb
Uraria picta (Jacq.) DC. Herb
Vicia hirsuta (L.) Gray Herb
Vicia sativa L. Herb
Vigna aconitifolia (Jacq.) Marechal Herb
Family Taxa Growth form
Vigna mungo (L.) Hepper Herb
Vigna radiata (L.) R.Wilczek Herb
Vigna trilobata (L.) Verdc. Herb
Vigna umbellata (Thunb.) Ohwi & H.Ohashi Herb
Zornia gibbosa Span. Herb
Zornia glochidiata DC. Herb
Gesneriaceae Didymocarpus pygmaeus C.B.Clarke Herb
Gentianaceae Canscora alata (Roth) Wall. Herb
Canscora diffusa (Vahl) R.Br. ex Roem. & Schult. Herb
Hoppea dichotoma Willd. Herb
Swertia ciliata (D. Don ex G. Don) B.L. Burtt Herb
Hydrophyllaceae Hydrolea zeylanica (L.) Vahl Herb
Family Taxa Growth form
Lamiaceae Anisomeles indica (L.) Kuntze Herb
Hyptis suaveolens (L.) Poit. Herb
Leonotis nepetifolia (L.) R.Br. Herb
Leucas aspera (Willd.) Link Herb
Leucas cephalotes (Roth) Spreng. Herb
Ocimum americanum L. Herb
Ocimum basilicum L. Herb
Ocimum tenuiflorum L. Herb
Orthosiphon pallidus Royle ex Benth. Herb
Premna herbacea Roxb. Herb
Salvia plebeia R.Br. Herb
Lauraceae Litsea glutinosa (Lour.) C.B. Rob. Tree
Litsea monopetala (Roxb.) Pers. Tree
Lecythidaceae Barringtonia acutangula (L.) Gaertn. Tree
Careya arborea Roxb. Tree
Lentibulariaceae Utricularia aurea Lour. Herb
Linaceae Linum usitatissimum L. Herb
Linderniaceae Lindernia anagallis (Burm.f.) Pennell Herb Lindernia ciliata (Colsm.) Pennell Herb Lindernia crustacea (L.) F.Muell. Herb Loranthaceae Dendrophthoefalcata (L.f.) Ettingsh. Shrub
Loranthus philippinensis Cham. & Schlecht.,
extract Shrub
Taxillus tomentosus Tiegh. Shrub
Lythraceae Ammannia baccifera L. Herb
Ammannia multiflora Roxb. Herb
Lagerstroemia indica L. Shrub
Lagerstroemia parviflora Roxb. Tree
Lagerstroemia speciosa (L.) Pers. Tree
Lawsonia inermis L. Shrub
Rotala indica (Willd.) Koehne Herb
Trapa natans L. Herb
Woodfordia floribunda Salisb. Shrub
Woodfordia fruticosa (L.) Kurz Shrub
Family Taxa Growth form
Malvaceae Abelmoschus manihot (L.) Medik. Herb
Abelmoschus moschatus Medik. Herb
Abelmoschus crinitus Wall. Shrub
Abelmoschus esculentus (L.) Moench Shrub Abutilon ramosum (Cav.) Guill. & Perr. Shrub
Abutilon indicum (L.) Sweet Shrub
Byttneria herbacea Roxb. Shrub
Ceibapentandra (L.) Gaertn. Tree
Corchorus aestuans L. Herb
Corchorus capsularis L. Herb
Corchorus tridens L. Herb
Corchorus olitorius L. Herb
Grewia asiatica L. Tree
Grewia hirsuta Vahl Tree
Grewia tiliifolia Vahl Tree
Hibiscus cannabinus L. Shrub
Hibiscus lobatus (Murray) Kuntze Herb
Hibiscus micranthus L.f. Shrub
Hibiscus mutabilis L. Shrub
Hibiscus panduriformis Burm.f. Shrub
Hibiscus sabdariffa L. Herb
Kydia calycina Roxb. Tree
Malvastrum coromandelianum (L.) Garcke Herb
Pavonia cancellata (L.) Cav. Herb
Pterospermum acerifolium (L.) Willd. Tree Sida cordata (Burm.f.) Borss.Waalk. Shrub
Sida cordifolia L. Shrub
Sida spinosa L. Herb
Sida acuta Burm.f. Herb
Sida rhombifolia L. Herb
Thespesia populnea (L.) Sol. ex Correa Tree
Triumfetta pentandra A. Rich. Shrub
Urena lobata L. Herb
Family Taxa Growth form
Martyniaceae Martynia annua L. Herb
Meliaceae Azadirachta indica A. Juss. Tree
Melia azedarach L. Tree
Soymidafebrifuga (Roxb.) A. Juss. Tree
Toona ciliata M. Roem. Tree
Menispermaceae Cissampelos pareira L. Shrub
Cocculus diversifolius DC. Climber
Cocculus hirsutus (L.) W.Theob. Climber Tinospora cordifolia (Willd.) Miers Climber Menyanthaceae Nymphoides cristata (Roxb.) Kuntze Herb
Moraceae Artocarpus heterophyllus Lam. Tree
Family Taxa Growth form
Artocarpus lacucha Buch.-Ham Tree
Ficus benghalensis L. Tree
Ficus benjamina L. Small tree
Ficus elastica Roxb. ex Hornem Tree
Ficus hispida L.f. Tree
Ficus palmata Forssk. Small tree
Ficus pumila L. Climber
Ficus racemosa L. Tree
Ficus religiosa L. Tree
Ficus virens Aiton Tree
Morus alba L. Tree
Morus indica L. Tree
Streblus asper Lour. Tree
Moringaceae Moringa oleifera Lam. Tree
Myrsinaceae Embelia tsjeriam-cottam (Roem. & Schult.) A.DC. Tree Myrtaceae Callistemon citrinus (Curtis) Skeels Tree
Eucalyptus tereticornis Sm. Tree
Psidium guajava L. Tree
Syzygium cumini (L.) Skeels Tree
Syzygium salicifolium (Wight) J.Graham Tree
Nyctaginaceae Boerhavia diffusa L. Herb
Boerhavia chinensis (L.) Rottb. Herb
Boerhavia repens L. Herb
Bougainvillea glabra Choisy in DC. Shrub
Mirabilis jalapa L. Herb
Nymphaeceae Nymphaea nouchali Burm. f. Herb
Nymphaea pubescens Willd. Herb
Olacaceae Olaxscandens Roxb. Shrub
Oleaceae Jasminum multiflorum (Burm. f.) Andrews Shrub
Jasminum arborescens Roxb. Shrub
Nyctanthes arbor-tristis L. Small tree
Onagraceae Ludwigia octovalis (Jacq.) P.H.Raven Herb
Ludwigia perennis L. Herb
Orobanchaceae Orobanche aegyptiaca Pers. Herb
Oxalidaceae Averrhoa carambola L. Tree
Biophytum reinwardtii (Zucc.) Klotzsch Herb
Biophytum sensitivum (L.) DC. Herb
Oxalis corniculata L. Herb
Papaveraceae Argemone mexicana L. Herb
Fumaria indica (Hausskn.) Pugsley Herb
Pedaliaceae Sesamum mulayanum N.C.Nair Herb
Sesamum indicum L. Herb
Phyllanthaceae Bridelia retusa (L.) A. Juss. Tree
Cleistanthus collinus (Roxb.) Benth. ex Hook.f. Tree Flueggea virosa (Roxb. ex Willd.) Royle Shrub
Family Taxa Growth form
Phyllanthus emblica L. Tree
Phyllanthus urinaria L. Herb
Phyllanthus virgatus G. Forst. Herb
Phyllanthus reticulatus Poir. Small tree
Plantaginaceae Lindenbergia indica Vatke Herb
Scoparia dulcis L. Herb
Plumbaginaceae Plumbago zeylanica L. Herb
Putranjivaceae Putranjiva roxburghii Wall. Tree
Polygalaceae Polygala chinensis L. Herb
Family Taxa Growth form
Polygala longifolia Poir. Herb
Polygala arvensis Willd. Herb
Polygonaceae Antigonon leptopus Hook. & Arn. Climber
Persicaria barbata (L.) H.Hara Herb
Persicaria hydropiper (L.) Delarbre Herb
Polygonum aviculare L. Herb
Polygonum plebeium R.Br. Herb
Rumex dentatus L. Herb
Portulacaceae Portulaca oleracea L. Herb
Primulaceae Anagallis arvensis L. Herb
Punicaceae Punica granatum L. Small tree
Ranunculaceae Ranunculus sceleratus L. Herb
Ranunculus scleratoides Perfil. ex Ovczinn. Herb
Rhamnaceae Ventilago maderaspatana Gaertn. Tree
Ziziphus nummularia (Burm.f.) Wight & Arn. Small tree Ziziphus oenopolia (L.) Mill. Small tree
Ziziphus jujuba Mill. Small tree
Ziziphusxylopyrus (Retz.) Willd. Small tree Rubiaceae Catunaregam longispina (Link) Tirveng. Small tree
Ceriscoides turgida (Roxb.) Tirveng. Tree Dentella repens (L.) J.R.Forst. & G.Forst. Herb
Gardenia latifolia Aiton Tree
Haldina cordifolia (Roxb.) Ridsdale Tree
Knoxia sumatrensis (Retz.) DC. Herb
Mitragyna parvifolia (Roxb.) Korth Tree Neolamarckia cadamba (Roxb.) Bosser Tree
Spermadictyon suaveolens Roxb. Herb
Oldenlandia diffusa (Willd.) Roxb. Herb
Oldenlandia corymbosa L. Herb
Oldenlandia herbacea (L.) Roxb. Herb
Spermacoce articularis L.f. Herb
Tamilnadia uliginosa (Retz.) Tirveng. & Sastre Small tree Wendlandia heynei (Schult.) Santapau &
Merchant Tree
Rutaceae Aegle marmelos (L.) Correa Tree
Citrus limon (L.) Osbeck Tree
Citrus medica L. Tree
Family Taxa Growth form
Glycosmis pentaphylla (Retz.) DC. Shrubs Hesperethusa crenulata (Roxb.) M. Roem. Tree
Limonia acidissima Groff Tree
Murraya koenigii (L.) Spreng. Tree
Murraya paniculata (L.) Jack Tree
Salicaceae Casearia tomentosa Roxb. Tree
Flacourtia indica (Burm.f.) Merr. Tree
Sapindaceae Sapindus emarginatus Vahl Tree
Schleichera oleosa (Lour.) Merr. Tree Sapotaceae Madhuca longifolia (J.Koenig ex L.) J.F.Macbr. Tree Scrophulariaceae Bacopa monnieri (L.) Wettst. Herb
Centranthera nepalensis D.Don Herb
Kickxia ramosissima (Wall.) Janch. Herb Limnophila heterophylla (Roxb.) Benth. Herb
Limnophila indica (L.) Druce Herb
Mazus pumilus (Burm.f.) Steenis Herb
Mecardonia procumbens (Mill.) Small Herb
Scoparia dulcis L. Herb
Veronica anagallis L. Herb
Simaroubaceae Ailanthus excelsa Roxb. Tree
Solanaceae Datura innoxia Mill. Herb
Datura metel L. Shrub
Lycopersicon lycopersicum (L.) H.Karst. Shrub
Nicotiana plumbaginifolia Viv. Herb
Nicotiana tabacum L. Herb
Physalis minima L. Herb
Physalis peruviana L. Herb
Solanum americanum Mill. Herb
Solanum incanum L. Herb
Solanum lycopersicum L. Shrub
Family Taxa Growth form
Solanum melongena L. Herb
Solanum nigrum L. Herb
Solanum surattense Burm. f. Herb
Solanum viarum Dunal Herb
Solanum virginianum L. Herb
Solanum tuberosum L. Herb
Sterculiaceae Byttneria herbacea Roxb. Herb
Eriolaena hookeriana Wight & Arn. Tree
Firmiana simplex (L.) W.Wight Tree
Helicteres isora L. Shrub
Melochia corchorifolia L. Herb
Sterculia villosa Roxb. Tree
Waltheria indica L. Herb
Ulmaceae Holoptelea integrifolia ( Roxb. ) Planch. Tree
Parasponica rugosa Blume Tree
Family Taxa Growth form
Verbenaceae Clerodendrum phlomidis L.f. Shrub
Lantana camara L. Shrub
Lippia javanica (Burm.f.) Spreng. Shrub
Phyla nodiflora (L.) Greene Shrub
Tectona grandis L. Tree
Violaceae Hybanthus enneaspermus (L.) F.Muell. Herb Hybanthus linearifolius (Vahl) Urb. Herb Vitaceae Ampelopsis glandulosa (Wall.) Momiy. Climber
Cayratia trifolia (L.) Domin Climber
Leea asiatica (L.) Ridsdale Shrub
MONOCOT
Alismataceae Sagittaria sagittifolia L. Herb
Amaryllidaceae Crinum asiaticum L. Herb
Aponogetonaceae Aponogeton crispus Thunb. Herb
Aponogeton natans (L.) Engl. & K.Krause Herb
Araceae Alocasia macrorrhizos (L.) G. Don Herb
Amorphophallus paeoniifolius (Dennst.) Nicolson Herb
Colocasia esculenta (L.) Schott Herb
Pistia stratiotes L. Herb
Remusatia vivipara (Roxb.) Schott Herb
Arecaceae Phoenix acaulis Roxb. Shrub
Phoenix rupicola T.Anderson Tree
Asparagaceae Asparagus racemosus Willd. Shrub
Colchicaceae Gloriosa superba L. Shrub
Commelinaceae Commelina benghalensis L. Herb
Commelina caroliniana Walter Herb
Commelina diffusa Burm.f. Herb
Commelinapaludosa Blume Herb
Cyanotis axillaris (L.) D.Don ex Sweet Herb Murdannia malabarica (L.) G.Bruckn. Herb
Murdannia nudiflora (L.) Brenan Herb
Cyperaceae Courtoisina cyperoides (Roxb.) Sojak Herb
Cyperus compactus Retz. Herb
Cyperus compressus L. Herb
Cyperus cyperoides (L.) Kuntze Herb
Cyperus difformis L. Herb
Cyperus exaltatus Retz. Herb
Cyperus iria L. Herb
Cyperus glomeratus L. Herb
Cyperus leucocephalus Retz. Herb
Cyperus malaccensis Lam. Herb
Cyperus michelianus subsp. pygmaeus (Rottb.)
Asch. & Graebn. Herb
Cyperus niveus Retz. Herb
Cyperus pangorei Rottb. Herb
Cyperus rotundus L. Herb
Family Taxa Growth form
Cyperus tenuispica Steud. Herb
Eleocharis geniculata (L.) Roem. & Schult. Herb Eleocharis palustris (L.) Roem. & Schult Herb Eleocharis quinqueflora (Hartmann) O.Schwarz Herb Fimbristylis bisumbellata (Forssk.) Bubani Herb
Family Taxa Growth form
Fimbristylis dichotoma (L.) Vahl Herb
Fimbristylisfalcata (Vahl) Kunth Herb
Fimbristylis miliacea (L.) Vahl Herb
Fimbristylis squarrosa Vahl Herb
Fimbristylis tetragona R.Br. Herb
Fuirena ciliaris (L.) Roxb. Herb
Kyllinga brevifolia Rottb. Herb
Pycreus flavidus (Retz.) T.Koyama Herb Pycreus polystachyos (Rottb.) P.Beauv. Herb
Pycreus pumilus (L.) Nees Herb
Schoenoplectiella articulata (L.) Lye Herb Schoenoplectiella juncoides (Roxb.) Lye Herb
Scirpus supinus L. Herb
Scleria levis Retz. Herb
Eriocaulaceae Eriocaulon guinguangulare Linn Herb
Hydrocharitaceae Blyxa aubertii Rich. Herb
Hydrilla verticillata (L.f.) Royle Herb
Najas minor All. Herb
Vallisneria spiralis L. Herb
Lemnaceae Lemna aequinoctialis Welw. Herb
Wolffia microscopica (Griff.) Kurz Herb
Musaceae Musa balbisiana Colla Large Shrub
Orchidaceae Vanda tessellata (Roxb.) Hook. ex G.Don Herb Zeuxine strateumatica (L.) Schltr. Herb
Poaceae Alloteropsis cimicina (L.) Stapf Herb
Andropogon fastigiatus Sw. Herb
Apluda mutica L. Herb
Apocopis vaginata Hack. Herb
Arundo donax L. Shrubs
Aristida adscensionis L. Herb
Aristida funiculata Trin. & Rupr. Herb
Aristida redacta Stapf Herb
Aristida setacea Retz. Herb
Arthraxon hispidus (Thunb.) Makino Herb Arthraxon lancifolius (Trin.) Hochst. Herb Arthraxon prionodes (Steud.) Dandy Herb
Avena sativa L. Herb
Arundinella setosa Trin. Herb
Bothriochloapertusa (L.) A.Camus Herb
Bambusa bambos (L.) Voss Tree
Chloris virgata Sw. Herb
Family Taxa Growth form
Chrysopogonfulvus (Spreng.) Chiov. Herb
Coix aquatica Roxb. Herb
Cymbopogon martini (Roxb.) W.Watson Herb
Cynodon dactylon (L.) Pers. Herb
Dactyloctenium aegyptium (L.) Willd. Herb Dendrocalamus strictus (Roxb.) Nees Herb Desmostachya bipinnata (L.) Stapf Herb Dichanthium annulatum (Forssk.) Stapf Herb Digitaria bicornis (Lam.) Roem. & Schult. Herb Digitaria ciliaris (Retz.) Koeler Herb Digitaria longiflora (Retz.) Pers. Herb
Digitaria stricta Roth Herb
Dimeria ornithopoda Trin. Herb
Echinochloa colona (L.) Link Herb
Echinochloa crus-galli (L.) P.Beauv. Herb Echinochloa stagnina (Retz.) P.Beauv. Herb
Eleusine indica (L.) Gaertn. Herb
Enteropogon dolichostachyus (Lag.) Keng Herb Eragrostis amabilis (L.) Wight & Arn. Herb Eragrostis japonica (Thunb.) Trin. Herb
Eragrostis pilosa (L.) P.Beauv. Herb
Eragrostis tremula Hochst. ex Steud. Herb Eragrostis unioloides (Retz.) Nees ex Steud. Herb Eremopogon foveolatus (Delile) Stapf Herb Eulalia trispicata (Schult.) Henrard Herb
Family Taxa Growth form Eulaliopsis binata (Retz.) C.E.Hubb. Herb Hackelochloa granularis (L.) Kuntze Herb Hemarthria compressa (L.f.) R.Br. Herb Heteropogon contortus (L.) P.Beauv. ex Roem. &
Schult. Herb
Hordeum vulgare L. Herb
Hygroryza aristata (Retz.) Nees ex Wight & Arn. Herb Imperata cylindrica (L.) Raeusch. Herb
Isachne globosa (Thunb.) Kuntze Herb
Ischaemum rugosum Salisb. Herb
Iseilema laxum Hack. Herb
Leptochloa chinensis (L.) Nees Herb
Leptochloa panicea (Retz.) Ohwi Herb
Melanocenchris jacquemontii Jaub. & Spach Herb
Ophiuros exaltatus (L.) Kuntze Herb
Oplismenus hirtellus (L.) P.Beauv. Herb
Oryza glaberrima Steud. Herb
Oryza rufipogon Griff. Herb
Oryza sativa L. Herb
Panicum notatum Retz. Herb
Panicum paludosum Roxb. Herb
Family Taxa Growth form
Paspalidium flavidum (Retz.) A.Camus Herb
Paspalum scrobiculatum L. Herb
Pennisetum glaucum (L.) R.Br. Herb
Pennisetum pedicellatum Trin. Herb
Perotis indica (L.) Kuntze Herb
Phalaris minor Retz. Herb
Pogonatherum paniceum (Lam.) Hack. Herb Polypogon monspeliensis (L.) Desf. Herb Pseudoraphis spinescens (R.Br.) Vickery Herb Pseudosorghum fasciculare (Roxb.) A.Camus Herb
Saccharum bengalense Retz. Herb
Saccharum spontaneum L. Herb
Sacciolepis interrupta (Willd.) Stapf Herb Sacciolepis myosuroides (R.Br.) A.Camus Herb Schizachyrium brevifolium (Sw.) Buse Herb Schizachyrium exile (Hochst.) Pilg. Herb
Schoenefeldia gracilis Kunth Herb
Sehima nervosum (Rottler) Stapf Herb
Setaria intermedia Roem. & Schult. Herb Setaria pumila (Poir.) Roem. & Schult. Herb Setaria verticillata (L.) P.Beauv. Herb
Sorghum bicolor (L.) Moench Herb
Themeda quadrivalvis (L.) Kuntze Herb
Thysanolaena latifolia (Roxb. ex Hornem.) Honda Herb
Tragus racemosus (L.) All. Herb
Triticum aestivum L. Herb
Urochloa panicoides P.Beauv. Herb
Pontederiaceae Eichhornia crassipes (Mart.) Solms Herb Monochoria vaginalis (Burm.f.) C.Presl Herb
Potamogetonaceae Potamogeton crispus L. Herb
Potamogeton nodosus Poir. Herb
Smilacaceae Smilax guianensis Vitman Herb
Typhaceae Typha domingensis Pers. Herb
Typha elephantina Roxb. Herb
Xanthorrhoeaceae Asphodelus tenuifolius Cav. Herb
Zingiberaceae Cheilocostus speciosus (J. Koenig.) C. D. Specht Herb
Curcuma aromatica Salisb. Herb
Curcuma longa L. Herb
Zingiber officinale Roscoe Herb
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