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ارزیابی زیستگاه دارکوب سرسرخ (Dendrocopos medius) در پارک ملی گلستان

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دراد راﺮﻗ ﻪﻌﻟﺎﻄﻣ و ﻆﻔﺣ اﺬﻟ ، آ

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Dendrocopos medius ... (

ﻴﻳ ﻗﻼ ﻲ

Alnus subcordata

Quercus castaneifolia

Carpinus betulus

Acer cappadocicum

Zelkova carpinifolia

Parrotia persica

Populus caspica

Diospyros lotus

!!

Celtis australis

Tilia begonifolia

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Ulmus glabra

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داﺪﻌﺗ ﻪﻧﻮﻤﻧ ﻪﻌﻄﻗ ﻪﻧﻮﻤﻧ يﺎﻫ

يرادﺮﺑ

= 106

هﮋﻳو راﺪﻘﻣ

Eigenvalue

085

/ 0 04 / 0 03 / 0

ﻲﻬﻴﺟﻮﺗ ﺲﻧﺎﻳراو ﺪﺻرد

34

16

12

راو ﻲﻌﻤﺠﺗ ﺪﺻرد ﻲﻬﻴﺟﻮﺗ ﺲﻧﺎﻳ

34 50

62

Factor loading

طﻮﻠﺑ 245

/ 0 - 495 / 0 324 / 0 -

زﺮﻤﻣ

432

/ 0 - 187 / 0 065 / 0

رادﺮﻴﺷ اﺮﻓا

360 / 0 267 / 0 632 / 0

ﻲﻠﻴﺠﻧا

960

/ 0 085 / 0 283 / 0 -

ﺎﻜﺳﻮﺗ

ﻴﻳ ﻗﻼ ﻲ 026 / 0 186 / 0 - 259 / 0

دازآ

011

/ 0 - 583 / 0 - 204 / 0

يﺪﻨﻣﺮﺧ

025 / 0 - 479 / 0 - 100 / 0 -

نﺎﻏاﺪﻏاد

231 / 0 - 526 / 0 - 182 / 0 -

ﺖﻠﭘﺪﻴﻔﺳ

186 / 0 - 514 / 0 - 184 / 0 -

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لوا رﻮﺤﻣ نﺎﻏاﺪﻏاد

رادﺮﻴﺷ اﺮﻓا طﻮﻠﺑ

ﻲﻠﻴﺠﻧا زﺮﻤﻣ

ﺎﻜﺳﻮﺗ

يﺪﻨﻣﺮﺧ ﺖﻠﭘﺪﻴﻔﺳدازآ

(9)

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Dendrocopos medius ... (

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.

- Alizadeh, A. 2006. Identifying bird species as biodiversity indicators for terrestrial ecosystem management, PhD thesis, RMIT University, Melbourne, Australia, 173p.

- Antongiovanni, M., and J.P. Metzer. 2005. Influence of matrix habitats on the occurrence of insectivorous bird species in Amazonian forest fragments. Biological Conservation 122: 441-451.

- Bahadori, F. 2008. Habitat suitability modeling of Eurasian Nuthatch (Sitta europaea) at Northern Alborz, Iran. , MS Thesis, University of Tehran, Iran, 92 p.

- Bartoszewicz, M.; H. Okarma, A. Zalewski and J. Szczesna. 2008. Ecology of the raccoon (Procyon lotor) from western Poland. Annales Zoologici Fennici 45: 291-298.

- Beauvias, G.P.; D.A. Keinath; P. Hernandez; L. Master and R. Thurston. 2006. Element distribution modelling. University of Wyoming, Wyoming, 42p.

- Campbell, D. 2001. The encyclopedia of British birds. The Foundry Creative Media Company Ltd, Crabtree Hall, Fulham, London, 384p.

- Castelletta, M.; J.M. Thiollay and N.S. Sodhi. 2005. The effects of extreme forest fragmentation on the bird community of Singapore Island. Biological conservation 121: 135-155.

- Claridge, M.F. and H.F. Evans. 1990. Species-area relationships: relevance to pest problems of British trees? In: Watt, A.D., Leather, S.R., Hunter, M.D., Kidd, N.A, (Eds), Population Dynamics of Forest Insects.

Intercept, Andover: 59-69.

- Darvish Sefat, A. 2006. Atlas of protected areas of Iran. Department of the Environment, Iran. 157p.

- Diaz, I.; J.J. Armesto; S. Reid; K.E. Sieving & M.F. Willson. 2005. Linking forest structure and composition: aian diversity in successional forests of Chiloe Island, Chile, Biological Conservation 123: 91-101.

- Hasanzadeh Kiabi, B; B. Zehzad; B. Farhang Darehshouri; H. Majnoonian and H. Gashtasb Meygooni.

1993. Golestan National Park. Department of Environment. 203p.

- Hooper, R.G. 1996. Arthropod biomass in winter and the age of longleaf pines. Forest Ecology and Management 82: 115-131.

- Kuchler, A. W. 1967. Vegetation Mapping. The Ronald Press Company Net Work, 472p.

- Marsden, S.J.; M. Whiffin and M. Galetti. 2001. Bird diversity and abundance in forest fragments and Eucalyptus plantations around an Atlantic forest reserve, Brazil. Biodiversity and Conservation 10: 737-751.

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:

) Dendrocopos medius ... (

- Muller, J; J. Pollath; R. Moshammer and B. Schroder. 2009. Predicting the occurrence of Middle Spotted Woodpecker Dendrocopos medius on a regional scale, using forest inventory data. Forest Ecology and Management 257: 502-509.

- Pasinelli, G. 2000. Oaks (Quercus sp.) and only oaks? Relations between habitat structure and home range size of the middle spotted woodpecker (Dendrocopos medius). Biological Conservation 93: 227-235.

- Pasinelli, G. 2003. Dendrocopos medius Middle Spotted Woodpecker. BWP Update 5: 49–99.

- Pasinelli, G. and J. Hegelbach. 1997. Characteristics of trees preferred by foraging middle spotted woodpecker Dendrocopos medius in northern Switzerland. Ardea 85: 203-209.

- Pasinelli, G. 2007. Nest site selection in middle and great spotted woodpeckers Dendrocopos medius and D.

major: implications for forest management and conservation. Biodiversity and Conservation 16: 1283–1298.

- Salman Mahini, A. 2009. Fundamentals of Environmental Protection. Rahe Daneshe Sabz. 337p.

- Schmitz, L. 1993. Distribution et habitat du pic mar Dendrocopos medius en Belgique. Aves 30: 145-166.

- Snow, D.W. and C.M. Perrins. 1998. The Birds of theWestern Palearctic. Concise Edition, vol. 2. Oxford University Press, New York. 696p.

- Spitznagel, A. 1990. The influence of forest management on woodpecker density and habitat use in floodplain forests of the upper rhine valley. In: Carlson, G., Aulen, A. (Eds.), Conservation and Management of Woodpecker Populations. Report 17, Swedish University of Agricultural Sciences, Uppsala: 117-145.

- Varasteh Moradi, H. 2004. Population dynamism of the Bovidae in Golestan National Park. Journal of Agricultural Sciences and Natural Resources. 11 (3): 163-172.

- Varasteh Moradi, H. 2005. Determination of sex ratio and age groups in two species Cervus elaphus and Capreolus capreolus in Golestan National Park. Journal of Agricultural Sciences and Natural Resources. 12 (4):

154-161

- Varasteh Moradi, H. 2004. Population dynamism of the Bovidae in Golestan National Park. Journal of Agricultural Sciences and Natural Resources. 11 (3): 163-172.

- Varasteh Moradi, H. 2005. Determination of sex ratio and age groups in two species Cervus elaphus and Capreolus capreolus in Golestan National Park. Journal of Agricultural Sciences and Natural Resources. 12 (4):

154-161

- Varasteh Moradi, H. and M. Zakaria. 2010. Responses of babblers (Timaliidae) to the forest edge-interior gradient in an isolated tropical rainforest in Peninsular Malaysia. Journal of Tropical Forest Science 22: 36-48.

- Varasteh Moradi, H.; M. Zakaria; B.M. Abdullah and Y. Ebil. 2009. Insectivorous birds and environmental factors across an edge-interior gradient in tropical rainforest of Malaysia. International Journal of Zoological Research 5: 27-41.

- Watson, J.E.M.; R.J. Whittaker and T.P. Dawson. 2004. Habitat structure and proximity to forest edge affect the abundance and distribution of forest-dependent birds in tropical coastal forests of southeastern Madagascar. Biological Conservation 120: 311–327.

- Winkler, H.; D.A. Christie and D. Nurney. 1995. Woodpeckers-A Guide to the Woodpeckers, Piculets and Wrynecks of the World. Pica Press, East Sussex, 158p.

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315 - 303 . pp , 2010 , 3 . No , 63 . Vol , Iranian Journal of Natural Resources ,

Natural Environmental Journal of

315

Habitat evaluation of Middle Spotted Woodpecker (Dendrocopos medius) in Golestan National Park

H. Varasteh Moradi*1

1 Assistant Prof., Gorgan University of Agricultural Sciences and Natural Resources, Gorgan, I.R. Iran

(Received: 26 September 2010, Accepted: 08 January 2011)

Abstract

The evaluation of ecological characteristics of Wildlife species has fundamental role in species management, because taking a decision for proper method of management is only possible by characterizing of ecological requirements of a species. In this study, suitable habitat of Middle Spotted Woodpecker was investigated in Golestan National Park. Presence and absence of birds and environmental variables (forest cover types, topography, and structural characteristics of vegetation) were obtained within a 25-m radius of each of 106 sampling points distributed throughout the study area. Data analysis was conducted using binary logistic regression. The results showed that some habitat variables including the number of snags, tree species, height of trees, d.b.h of trees,and basal area were the most significant positive predictors affecting this species presence in Golestan National Park. Old and undisturbed forest habitats with characteristics such as high values of basal area, number of snags, d.b.h, and height of trees in Golestan National Park has the capability of supporting the 'umbrella' and sensitive species to the disturbance such as the Middle Spotted Woodpecker.

Keywords: Golestan National Park, habitat modeling, Middle Spotted Woodpecker

*Corresponding author: Tel: +98 171 2245965 Fax: +98171-2245886 E-mail: hvarasteh2009@yahoo.com

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