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:

*

!"#$%#

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E-mail: pourjam@modares.ac.ir

0912

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F.afghanicus (Khan & Khan, 1978) Siddiqi, 1986 F. andrassyi (Szczygiel, 1969)

Andrássy, 1979 F.baloghi (Andrássy, 1958) Siddiqi, 1986

F.cylindricaudus (Wu,

1969) Siddiqi, 1986 F. facultativus (Szczygiel, 1970) Raski & Geraert, 1987

F.misellus (Andrássy, 1958) Raski & Geraert, 1987 F.paravesiculosus Karegar &

Geraert, 1998 F. polyhypnus (Steiner & Albin, 1946) Meyl, 1961

F. sandneri

(Wasilewska, 1965) Raski & Geraert, 1987 F.vulgaris (Brzeski, 1963) Lownsbery

& Lownsbery, 1985 F.elegantulus Raski & Geraert, 1986

F. longicaudatulus Zell,

#

1988

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BI ; ? ; +. ? V

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!

&

$? U$V %

Meyl

.

(1961)

9

> $

Filenchus

=% #

) = 6 $? U$V > R P

!

# > W

F. filiformis (Butschli, 1873) T. (F).

= 6

duplexus

= 6 $? X / > 9# #

Tylenchus

valkanovi Andrássy, 1958

> =% #

Filenchus

$? :Y0

Brzeski

.

(1963)

N % Z : 4

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= 6

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^ "

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1 - 0 A, 9# #

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Tylenchus

(Filenchus) filiformis

G #H6

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! [

"

= 6 $` L? 9#

T. vulgaris Brzeski,

I. )

1983 Lownsbery & Lownsbery (1985)

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^ "

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1

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!

$ 4 [.# > bV.

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(Spicies

$?

incuired)

. d)

Brzeski

!

(1982)

# $%

P ) =? &

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(P#

T. filiformis

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"

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1

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=Y`C

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=Y`C

% 4

#

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63

%

F. filiformis

= 6

F. vulgaris

$? g " # .

Tylenchidae

# / h`C

$/3

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3 4 g T #

,2008

)

Geraert

(

# # 4 42 9 i =? [.# >

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Atylenchinae,

Boleodorinae, Tylodorinae Ecphyadophorinae

.

Tylenchinae

4 1 6

!

$ . k\% >

9

Tylenchinae

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=0 # #$Z

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& $06 H% = 6

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(De Grisse, 1969)

= [a$6 g T #

.

& q

1 -

> 1 . 4 " = 6 [ !,

Filenchus

-#$ #

m`

G #H6 hC - %H! - " !6

= 6

4 -# " = $a = L]

Filenchus acris (Brzeski, 1986) Raski & Geraert,1987 _#$6

3 H

g# % =0 # F. afghanicus (Khan & Khan, 1978) Raski & Gereart,1986

6 T%

1 #$ # f F. andrassy (Szczygiel, 1969) Andrássy, 1979

4 -# "

Equisetum arvensis F. baloghi (Andrássy, 1958) Siddiqi,1986

6 _ ? ` 6 AZ %3 - 0 %

= # " =T 1 ! 9 r!.

F. cylindricaudus (Wu, 1969) Siddiqi,1986

4 -# "

F. discrepans (Andrássy, 1954) Raski & Geraert,1987 _#$6

2 4H?$ -#$A)

F. ditissimus Siddiqi,1987. #

4

&! # F. facultativus (Szczygiel, 1970) Raski & Geraert,1987 =`

5 [

F.fortis (Siddiqi & Lal, 1992) Brzeski, 1997 i $%

6

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1 #$ # f F. hamatus (Thorne & Malek, 1968) Raski & Geraert, 1987

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Z s]

F.helenae (Czszygiel, 1969) Raski & Geraert, 1987

4 H

F. misellus (Andrássy, 1958) Raski & Geraert, 1987 #

5

&! # F. paravesiculosus Karegar & Geraert, 1998 =`

2 X$?

r!.

s] 1 ! 9 t $(6 =T F. parvus (Siddiqi, 1963) Raski & Geraert, 1986

1 -S!6

F. polyhypnus (Steiner & Albin, 1946) Meyl, 1961 i $%

7

&! # F. quartus (Szczygiel, 1969) Lownsbery & Lownsbery, 1985 =`

2 X$?

g 6 s] 1 ! 9 r!.

T ? 1 ! 9 g# % F. sandneri (Wasilewska, 1965) Raski & Geraert,1987

4 -# "

Equisetum arvensis F. sheri Siddiqi, 1986

4 D(0f hC

D(0f - " !6 F. vulgaris (Brzeski,1963) Lownsbery & Lownsbery,1985

1 - g#

j-# @ " $a )

2001 ( 2 - 4$!/

) : 1970 ( 3 -

$6 ? : 1989 ) 3 (

4 5 6 - R #$6 $6 ? :

1995 ) 4

a ( 1998 ) 5

b ( 1998 ) 6 ( 7 -

$` Z j ! ) 1997 (

(3)

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Filenchus

>

. 1

= 6 =? [a$6 #$Z

# 9# S`Z -3 G #H6 -#$

#

#$% -3 = 6 4

- a

)

"

4

# -#$

:d' C =% = # # =? 0 "

"# / # U$

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Filenchus elegantulus Raski & Geraert,

.1987

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:

1 :@

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`p) 9# E% (%

! [

1 / 1 @ K$C =%

. [, O =% 4 O ) g

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= 6 %

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#

% = 6 &

!

$0L

&

= 6 % # [" `

"

F. andrassyi (Szczygiel, 1969)

4

Andrássy, 1979 F.marinus (Timm, 1956) Raski

& Geraert, 1987

- L 1 9 =? "

Y $

% =

!

&

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= 6 = 6 &

"

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4 )

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f 1 :@

(

= 6 9#

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= 6 9#

F. marinus

- % [ =V.# =% "

=Y(O 4 "

1q$F 0.# $6 - % # # - % 0 [(

H

# [.

.

[!E m

3 4 - .#$/ 9#

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[f 1.

( ? h% V) 1

(P# U$ % 1

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/$% IYa 1

v/ 9#

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4 [f T . 1 R ')

QH 1

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.

# C# 9# = 6 &

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K

$'. H

, `,3

m -S6$ 9# Bf%

3 4 . 1 .

Filenchus longicaudatulus Zell, 1988

= # 0.# - %

:

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`p) 9# E% J] ? :@

! [

R P =%

=0 %

C

? 1 9 % 1 3 , 1 r, 7 :@

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C

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@!) ?

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2 `O % (%

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O

! (. ) K @ = 1

% g

! (%

MS ? f 1 :@

[.#

.

:@

1

Filenchus elegantulus

-

A

: - % 1(? 4

:

:

B

$. =!O

C

:

D E

g :

F

1`YF = !? 1(. ) K @

O [.# L G #H6

:

.

(? 9# '0.# %

! R #$6 ) 2008 ( =Q# # U$

9 I. )

1988

) (

=E, V = 6

u \, =%

[f 1.

[f T . 1 A ) :% Z h% V) 1

= 6 %

F. longicaudatulus

- L 1

"

.

# I. ) = 6 &

@.9$%

1 ) 1997 (

= 6 %

F. helenae

, g "

1

R #$6 ) 2008 (

#

$? gSF# $`0E # = 6 &

. = 6

. $%

1

%!

$0L

= 6 % # [" ` &

"

F. helenae

4

(szczygiel, 1969) Raski & Geraert, 1987 F. infirmis (Andrássy, 1954) Siddiqi, 1986

- L

1

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.

# % , =% # = 6 9# &

!

[ :

$) - %

=Y(O

"

4 q$F 1

&A - % O - % $)

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X$f ) 1(. ) K @ =(P a =% g )

2

:% Y 6 / 2 (

# ?c = 6 9#

1 . g = 6

!

=% H

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! D

$) ) 4 - 5 / 3

:% Y 5 / 7 - 5 / 6

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$0 $@

( J] ? $. - % 9# $)

= 6 0 k\%

H 1

$6 .

# , # = 6 &

! :; 9# % &

"

4 - ,3 G#

m 3 4 G #H6

Zell, 1988

) .(

# ) &

h!Y\

=L K#$C# 9# = 6 & # H!

{.

! - 0.$A K#$C# #

m = f 3

3 4 . 1

#$%

4 , #

!

# 9# % &

-#$

G #H6 1 .

:@

2

Filenchus longicaudatulus

-

A

: : - % 1(? 4

:

B

4$ =!O :

C

g =!O :

D

B [A 1q$F G$%

1` %

(5)

2 -

[f R dfL

1T .

F. elegantulus

m 1(P# U$ % = Y 1, - .#$/ 9# 4 3

Origin Nortehrn Khorasan Raski &Geraert

Character Female Female

n 10 7

L 733 ± 71 (642-866) 811 ± 33 (760-858)

L' 615 ± 65.6 (542 -742) -

a 37.2 ± 1.5 (33.8-38.9) 31.5 ± 2.58 (27.6-36.1)

b 5.5 ± 0.5 (4.8-6.3) 5.8 ± 0.21 (5.5-6.1)

c 6.2 ± 0.6 (5.6-7.2) 5.8 ± 0.1 (5.7-5.9)

c' 9.3 ± 1.2 (7.7-11.4) 7.6 ± 0.38 (7.0-7.9)

V 66. 2 ± 1.2 (64.6-68.1) 65 ± 1.01 (64-67)

V' 79 ± 1.1 (77.1-80. 8) 79 ± 1.23 (77-80)

Stylet 11 ± 0.3 (10.5-11.5) 11 ± 0.92 (10-13)

MB 41.1 ± 2.5 (36.8- 46) -

Secretory-Excretory Pore 98. 2 ± 4.6 (93-106) 108.5 ± 4.64 (104-117) Oesophagus 133.5 ± 4.9 ( 125-140) 140.6 ± 2.39 (138-145)

Head-Vulva 485.9 ± 46.6 (435-572) -

PUS 12.6 ± 2.1 (10-16) 14.6 ± 2.28 (11-18)

Vulva - Anus 129.8 ± 19.8 (107-170) 144 ± 11.28 (132-157)

Tail 118 ± 11.1 (100-136) 139.4 ± 6.94 (128-147)

Tail/ v- a 0.9 ± 0.1 (0.7-1.0) 0.97 ± 0.05 (0.92-1.04)

3 -

[f R dfL

Filenchus longicaudatulus

1T . m

4 3

1(P# U$ % = Y 1, - .#$/ 9#

Origin provinceNorthern Khorasan Zell (1988)

Character Female Female

n 12 16

L 399.4 ± 19.1 (374-437) 350-447

L' 267 ± 13.5 (248-293) -

a 39.7 ± 2.7 (35.5-43.7) 25.3-43.6

b 5.8 ± 0.3 (5.4-6.4) 5.0-7.4

c 3 ± 0.1 (2.9-3.1) 2.6-3.2

c' 22.5 ± 2 (20-26.4) -

V 54.1 ± 1.3 (52.1-56.6) 48.8-58.5

V' 80.6 ± 1.7 (78.2-84.5) -

Stylet 6.9 ± 0.3 (6.5-7.5) 6.5-7.5

MB 42.1 ± 1.7 (39.7-45.7) 41.8-50

Secretory-Excretory Pore 51.4 ± 1.7 (50-56) 42-55

Oesophagus 68.3 ± 1.8 (66-72) 58-75

Head-Vulva 216 ± 11.9 (198-242) -

PUS 6.2 ± 0.7 (5.0-7.5) 7.0-10

Vulva - Anus 51.8 ± 5.3 (41-60) -

Tail 131.6 ± 6.9 (120-144) 117-164

Tail/ V-a 2.6 ± 0.3 (2.2-3.2) -

REFERENCES

1. Andrássy, I. (1954). Revision der Gattung Tylenchus Bastian, 1865. Acta Zoologica Academiae Scientiarum Hungaricae, 1, 5-42.

2. Andrássy, I. (1979). The genera and species of the family Tylenchidae

ö

rley, 1880(Nematoda). The genus Tylenchus Bastian, 1865. Acta Zoologica Academiae Scientiarum Hungaricae, 25, 1-33.

3. Brzeski, M. W. (1963). On the taxonomic status of Tylenchus filiformis Bütschli, 1873, and the description of T. vulgaris sp. n. (Nematoda: Tylenchidae). Bulletin de l'Académie Polonaise des Sciences, 11, 531-535.

4. Brzeski, M. W. (1982). Taxonomy of Ottolenchus Husain and Khan, and description of Coslenchus

(6)

polonicus sp. n. (Nematoda: Tylenchidae). Revue de Nématol, 5, 71-77.

5. Brzeski, M. W. (1997). Redescription of some species of the genus Filenchus Andrássy, 1954.

(Nematoda,Tylenchidae). Miscellanea Zoologica, 20(1), 45-64

6. Butschli, O. (1873). Beiträge zur Kenntnis der freilebenden Nematoden. Nova Acta Academiae Caesarea Leopoldino Carolinae germanicae naturae curiosorum 36, 1-144.

7. De Grisse, A. T. (1969). Redescription ou modifications de quelques techniques utilisées dans l'étude des nématodes phytoparasitaires. Mededelinyen Rijksfakulteit, 34, 351-369.

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