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6- Anderson J. J., Priester T. M., and Shalaby L. M. 1989. Metabolism of metsulfuron-methyl in wheat and barley. J. Agric. Food & Chem., 37:1429-1434
7- Baghestani M. A., Zand E., Mesgaran M. B., Veysi M., PourAzar R., and Mohammadipour M. 2008.
Control of weed barley species in winter wheat with sulfosulfuron at different rates and times of application. Weed Biology and Manage., 8:181-190.
8- Ball D. A., Frost, S. M., and Bennett, L. H. 2007. ACCase-inhibitor herbicide resistance in downy brome (Bromus tectorum) in Oregon. Weed Sci., 55:91-94.
9- Bayer Crop Science. 2006. Safety data sheet. Panther 18482 4X5L GB.
10- Brown A. H. D., Nevo E., Zohary D., and Dagan O. 1978. Genetic variation in natural populations of wild barley (Hordeum spontaneum). Genetica, 49:97-108.
11- Christopher J. T., Powles S. B., and Holtum J. A. M. 1992. Resistance to acetolactate synthase inhibiting herbicides in annual ryegrass (Lolium rigidum) involves at least two mechanisms. Plant Physiol., 100:1909-1913.
12- Geier P. W., and Stahlman P. W. 1996. Dose responses of weeds and winter wheat (Triticum aestivum) to MON 37500.Weed Technol., 5:3-17.
13- Harlan J. R. 1992. Crops and man. Madison, Wisconsin: University of Wisconsin Press.
14- Kelley J. P., and Peeper T. F. 2003. Wheat (Triticum aestivum) and rotational crop response to MON 37500. Weed Technol., 17:55-59.
15- Moseman J. G, Baenziger P. F., and Kilpatrick R. A. 1980. Hordeum spontaneum- an overlooked source of disease resistance. Europe and Mediterranean Cereal Rust Foundation, Pp. 91-93.
16- Ritz C., and Streibig J. C. 2005. Bioassay analysis using R. Journal of Statistical Software, 12:1-22.
17- Sensmen S. A., 2007. Herbicide Handbook. (9th ed). Weed Science Society of America, U.S.A.
18- Shinn S. L., Thill D. C., and William J. P. 1999. Volunteer barley (Hordeum vulgare) control in winter wheat (Triticum aestivum) with MON 37500. Weed Technol., 13:88-93.
19-Volis S., Mendlinger S., Turuspekov Y., Esnazarov U., Abugalieva S.,and Orlovsky N. 2001. Allozyme variation in Turkmenian populations of wild barley (Hordeum spontaneum Koch.).Annals of Botany, 87:435-446.
20- Zand E., Baghestani M. A., Soufizadeh S., Eskandari A., PourAzar R., Veysi M., Mousavi K., and Barjasteh A. 2007. Evaluation of some newly registered herbicides for weed control in wheat (Triticum aestivum L.) in Iran. Crop Protec., 26:1349-1358.
21- Zhang Q., Saghai-Maroof M. A., and Kleinhofs A. 1993. Comparative analysis of RFLPs and isozymes within and among populations of Hordeum vulgare spp. spontaneum. Genetics, 134:906-916.