cajpsula
Norman,
they were found to be identical and, therefore, S.levishas been placed in
synonymy
with S. capsula.Misidentifications of Sars (1888)
and
Miiller (1894) are under- standable considering the unillustrated description ofNorman
(1869),
which
could includemany
extant sarsiellids. Actually Miiller (1894) prefaced his jS. capsula identification with a "?" butthis
was
omitted in his synopticwork
in 1912.Both
malesand
females of S. capsulaand
S. levis were describedand
illustratedby
Miiller (1894).
Brady and Norman
(1896) established the genusNematohamma
to receive their
new
speciesN.
obliqua,which was
based on males collected off Valentiaand
Birterbuy Bay, Ireland.Nematohamma
obliqua
was
placed insynonomy
with ^S. levis Miillerby
Miiller (1912).As
previously stated, S. levis is asynonym
of S. capsulaNorman;
therefore,A^.obliquaisalsoasynonym
ofS. capsulaNorman.
1
have compared
carapaces of of S. capsulaNorman from
the Shet- land Islandsand
Valentia with the carapace of a specimen ofN.
obliqua
from
Valentia(Norman
Collection, BritishMuseum) and
find that they are indeed conspecific.
A
carapace of A^. obliquafrom
off Valentia (from sUde 1911.11.8,M3989,
BritishMuseum)
is illustrated in plate Se.Material. —
Information concerning specimensexamined
is as followscontents of label {location of material)
1911.11.8, M3986; Sarsiella cap- sula Norman, Madeira, off PraiaBay, 50 fathoms,A.M.N.
1897[British Museum]
1911.11.8, M3985; Sarsiella cap- sula Norman, Types, Shetland 1863
&
Valentia 1870 [BritishMuseum]
1911.11.8,37013;Sarsiellacapsula Norman, Birterbuy Bay, Ire- land, 1874[British Museum]
remarks
A
dryslide containing aleftand right valve. Absence of a posterodorsal prominence indicates it is not con- specific with*S. capsula Normanbutis probably a new species. Occur- renceofthe species offMadeira was reportedin Brady (1911).
Specimens in this dry slide are illus- trated in figures 16a-d. The back oftheslidecontains these notations:
"a, 10 miles oflf Balta, Shetland 73 fathoms; b. off Valentia, 112 fath- oms." These data do not agree withlocality anddepth information forShetland givenin Norman(1869, p. 293). The date (1863) does not agree with the date of 1869 given by Brady and
Norman
(1896).A
vial containing a whole specimen preservedinpreservative.SARSIELLA
— KORNICKER 35
contents of label {location of material) 4. 1900-3-6-456; Sarsiella capsula,
oflf Capri, Bay of Naples, April, 1887[British Museum]
5. 1900-3-6-457;Sarsiellacapsula,off Valentia, Ireland, 1870 [British
Museum]
6. 1900-3-6-458; Sarsiella capsula, off Valentia, Ireland, 1870 [British Museum]
7. 1911.11.8, M3987; Nematohamma
obliqua Bra[dy] and Nor[man], Type, Valentia Ireland, 1870 [British Museum]
8. 1911.11.8, M3988; Nematohamma
obliqua Bra[dy] and Nor[man], Type, Birterbuy Bay, Ireland, 1874 [British Museum]
9. 1911.11.8, M3989: Nematohamma
obliqua Bra[dy] and Nor[man], Type, off Valentia, 112 fath.
1870 [British Museum]
10. Ace. Cat. No. 25019; Sarsiella capsula
Norman
[ZoologicalIn- stitute of Greifswald]11. Ace. Cat. No. 25020; Sarsiella levis
M
tiller [Zoological Insti- tuteof Greifs^A'ald]12. Sarsiella capsula Norman, 20-30 M, 19/21/39, Station 07722, Rocquerbrane cf [Zoological Station of Naples]
remarks
A
dry slide containino mount,a female.a dry whole
A
slide containing mounted append- agesofanadult female.A
slide containing mounted append- ages of an adult female. Illustra- tionsofappendagesin thispaperare fromappendages on thisslide.A
slide containing appendages of a dissected specimen. Specimen was reported in Brady andNorman
(1896).
A
slide containing appendages of a dissected specimen. Specimen was reported in Brady andNorman
(1896).
A
dry slide containing a whole speci-men
in good condition. Specimen wasreportedin BradyandNorman
(1896). The bottom of the slide contains the pencilled notation:
"Philomedes Folinii, off Valentia, 112 fath. 1870." This specimen is illustrated in figure I6e.
Vialcontainingabout15 specimensin- cluding 1 adult male. Thesespeci-
menswerecollectedbyG.W.Miiller
from the Gulf of Naples and re- ported in Miiller (1894).
Vial containing about 6 specimens in- cluding 1 adult male. Thesespeci-
menswerecollectedby G.W. Miiller in the Gulf of Naples and reported in Miiller (1894).
A
slide containing a whole specimen mountedinbalsam. Itisa juvenile of a species similar to S. capsula of Miiller, notNorman. Collected byDom.
R. Rome.Description
ofadult
female.—
Shell (pi. 3a, b):Oval
in lateralview with posteroventral caudal process; posterodorsal part of shell inflated,
median
part depressed. Ornamentation: prominent ridge witliinand
parallel to dorsaland
ventral margins of shellbecoming
faintordisappearinganteriorly
and
posteriorlybut formingcrestalong dorsalmargin ofposterodorsalinflation;7 radialridgeswithin anteriorand
ventralsubmarginalregions; 3 anterior, 3 ventral,and
1 extending onto caudal process. Surface punctate.36 PROCEEDINGS OF THE NATIONAL MUSEUM
vol. 122Typical of sarsiellids, the caudal process of S. capsula is
more
anteriorly located on adults than on juveniles (pi. 3c, d).
The
shell of S.capsulaillustratedby Brady and Norman
(1896,pi.LX
: figs. 1-4) hasthe caudal processposterior, suggesting thatit is ajuvenile.The
lateral outline of the carapace is similar to that of the specimen illustrated in plate 3c of this paper.
Both
adult femalesand
juvenileshave
3 prominent radial ridges anteriorly.Three
ventral radial ridgeson
juvenilesmay
be present on adults also; however, it is not possible to be certain of the ridges on the adults because the ventral margins of both adult shellson
the type slide are considerably distorted. Juveniles, adult females, and males (A'^. obliqua) (pi. 3c)have
the flat-topped, crestUkeprominence on the posterodorsal part of the shell.Measurements:
Because adult specimenson
shde no. 1911.118,M3985
(BritishMuseum)
aresomewhat
distorted, the dimensions given belowmust
be considered as approximate (the lettersused to designate specimens refer to specimensillustrated in plate 3)specimen
SARSIELLA
— KORNICKER 37
processcarrying oneshortseta."A
secondspineis clearlydiscernable on BritishMuseum
slide no. 1900-3-6-458.Mandible
(fig. 16c):Each
joint of endopodite with long curvedFigure 16.
—
Sarsiella capsula Norman (slide 1900-3-6-458, British Museum): a, 1st antenna;b,2nd antenna;c,mandible;d,maxilla;e,7thHmb;/,furca.
claw ventrally.
The
mandible on each slide is in poor condition permitting discrimination of onlygross features.A
faintrow
ofshort spines borders the ventral margin ofthe coxaleofamandiblemounted
38 PROCEEDINGS
OFTHE NATIONAL MUSEUM
vol. 122on slide no. 1900-3-6-457 (British
Museum)
but not on mandibles on slide no. 1900-3-6-458 (BritishMuseum).
Maxilla (fig. 16f/):
2nd
joint of endopodite with 2 slenderbristleson outer side, 1 short bristle on inner side,
and
5 stout terminal bristles with marginal spines.The
maxillawas
the onlyappendage
illustratedby Brady and Norman
(1896, pi. 9: fig. 18). Itis essentially thesame
as themaxilla on slideno. 1900-3-6-458 (BritishMuseum)
illustrated in this paper(fig. 16^).
Seventh limb (fig. IQe): 6 bristles in terminal group; 2 bristles in proximal group; allbristles with 2-4 distalbells.
The
distal tip of the 7thappendage
can not be observed clearly onBritishMuseum
slides (1900-3-6-457; 1900-3-6-458). This limbwas
not describedby Brady and Norman
(1896).Furca (fig. 16/): each lamella with 5 curved claws decreasing in length posteriorly; claw no. 1 joined to lamella; claws 1-3 withlateral
and
medial spines inrow
along concave margin.Brady and Norman
(1896, p. 678) described the furca as having 6 claws. InbothBritishMuseum
slides (1900-3-6-457; 1900-3-6-458) the fm-ca is compressed,making
the claws difficult to distinguish however, only 5claws canbe observed on eachlamella.Sarsiella disparalisDarby
Figures 17-19; Plate 4 Sarsiella disparalis Darby, 1965,p. 40,pis. 30, 31.
Holotype:
Museum
ofPaleontology,UniversityofMichigan,48819, female.Darby
(1965, p. 40) described S. disparalis basedon
5 females collected off Sapelo Island, Ga.The
shells of his specimens varied considerably in ornamentation,some weakly and
others strongly ornamented, or the right valveornamented and
the left valvealmost plain. I received 2mature
females of thisunusual species, both with asymmetricalshells,inacollectionfrom
the AtlanticshelfoffBeaufort,N.C. To
determine whether the asjniimetrj^and
variability of the shellsmight
be causedby
parasitism, both specimens wereexamined
for parasites, but
none
were found. Leftand
right appendages werecompared
to determineifasymmetry
in valveswas
reflected in them, but differences were found to be minor.Examination
of a slide(UMMP
48818, no. 86) of a paratype preparedby Darby
revealed the presence of a parasitic copepod of the family Choniostomatidae, but it is not considered likely that it is the cause of shellasymmetry
in S. disyaralis because the specimen containing the copepod is fairly symmetrical.