Frond venation palmate (basal actinodromous) with four or five asymmetrical lobes; the basal sinus of the lobes incised either from the base of the blade or from more apical

Lygodiaceae

P

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F

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M

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G

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T

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. # T

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. #

L

OCA

CJ61

Lygodium sp. ^M

^ORPHOTYPE

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palmate - actinodromous basal

dichotomizing

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RHIZOMECHARACTERS

microphyll

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VENATION OF THEPINNAE AND/ORPINNULE

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DIAGNOSTICFEATURES

Incomplete pinnae. Laminar size microphyll (the specimen is ~2.6 cm long and ~2.9 cm wide). The base of the pinnae is decurrent and a short petiole is preserved (~3 mm long).

Frond venation palmate (basal actinodromous) with four or five asymmetrical lobes; the

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DIAGNOSTICFEATURES

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DESCRIPTION

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DESCRIPTION

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-2° V

EIN

F

REQUENCY

;"5",,%,8058486+

C

^{OURSE OF}

P

^ERCURRENT

3°

I

^NTERCOSTAL

3° V

^EIN

A

^NGLE

V

^ARIABILITY

E

PIMEDIAL

3°

A

^DMEDIAL

C

^OURSE

E

XMEDIAL

C

OURSE

E

^XTERIOR

3° C

^OURSE

DIAGNOSTICFEATURES

F/05/# "70 +%$870"5/%+ "5% +1861 "70 46,1/+15"70%0< G.8 +6#+%1+ 8* $,8+%,3 +;"$%0 ?%/7+ 12"1 ",1%57"1% "70

;"5",,%, 12% +%$870"5/%+ "5% ;5%+%71< E%$870"5/%+ "70 12% 5%+1 8* ?%/7 850%5+ "5% +1587=,3 $58.0%0 "70 0%$655%71 87 4/0?%/7< G.8 T/70 8* 2/=2%5 850%5 ?%/7+h $58++%0 ?%/7+ ,3/7= "1 ?"5/"#,% "7=,%+ "70 ?%/7+ 485% 85 ,%++-;"5",,%,-18-12%-1.8-+6#+%1+-8*-?%/7+<

SIMILARMORPHOTYPES

67,8#%0

F.E.V.

S

T

ERMINAE

;"5",,%,8058486+

*5%%,3-5"4/*3/7=

4"$58;23,, 4%=";23,,

)%"* +/O% 4"$58;23,, 18 4%=";23,, J12% #/==%+1 +;%$/4%7 /+ PAK $4 ,87= "70 :Y $4 ./0%N< F"5=/7

%71/5%< B5/4"53 ?%/7 *5"4%.85T ;58#"#,3 ;/77"1%< )"4/7"5 +2";% ";;"5%71,3 8#,87=< F/05/# "70 +%$870"53 ?%/7+ +1861 "70 46,1/+15"70%0[ :M +%$870"5/%+ "5% ;5%+%5?%0 87 12% #/==%+1 +;%$/4%7<

E%$870"53 ?%/7 +;"$/7= *584 M<: $4 18 Y<M $4< W"5/"1/87 8* 4"V85 +%$870"53 "7=,% 18 4/0?%/7

*584 _KH 18 ^KH> 12%7 12% "7=,% 0%$5%"+% %D4%0/",,3< G.8 +6#+%1+ 8* ;"5",,%,8058486+ "70 $,8+%,3 +;"$%0 ?%/7+ 12"1 ",1%57"1% /7 #%1.%%7 12% +%$870"5/%+< G2%+% +6#+%1+ "5% 5%$8=7/O%0 #3 12%/5 ./012> "70 /70%;%70%71,3 8* 12%/5 850%5 12% +;"$/7= 5"7=%+ *584 K<^ 44 18 :<X 44< G2% */5+1 +6#+%1 ";;%"5+ *584 X 18 _ 1/4%+ /7 #%1.%%7 " ;"/5 8* +%$870"5/%+< G2% +%$870 +6#+%1 $87+1/161%+

12% 4"V85/13 8* ?%/7+< G.8 ?%53 12/7 13;%+ 8* 2/=2%5 850%5 ?%7"1/87h ?%/7+ 12"1 $58++ 12% +6#+%1+ "1 8#,/e6%-"7=,%+>-"70-?%/7+-./12-"7-/55%=6,"5-$865+%>-;"5",,%,-18-12%-+6#+%1+<

DESCRIPTION

/715"4"5=/7",-+%$870"53

0%$655%71

#

Pteridaceae

P

LANT

F

AMILY

M

AJOR

P

LANT

G

ROUP PTE

O

RGAN

T

YPE Leaf

T

YPE LOC

. # T

YPE SPEC

. #

L

OCA

CJ66

Acrostichum sp. ^M

^ORPHOTYPE

^#

M

ORPHOTYPE

N

AME

absent

M

ARGINAL

U

LTIMATE

V

ENATION

A

REOLATION

V

EINLETS

C

OURSE

L

AMINAR

S

IZE

(

SMALLEST

)

pinnate

reticulodromous

apparently uniform regular - crowded

V

ARIATION OF

2° V

EIN

A

NGLE

M

AJOR

2° V

EIN

F

RAMEWORK

M

AJOR

2° S

PACING

1° C

OSTA

F

RAMEWORK

M

ORPHOTYPER

P

RINCIPAL

V

EIN

T

ERMINATION

T

OOTH

S

PACING

T

OOTH

S

HAPE

#

OF

O

RDERS

T

EETH

/

CM

S

INUS

S

HAPE

CERREJON FLORA

RHIZOMECHARACTERS

notophyll

L

AMINAR

S

IZE

(

LARGEST

)

mesophyll

P

INNAE

A

TTACHMENT

S

URFACE

T

EXTURE

TEETHFEATURES

T

OOTH

S

HAPE

P

RINCIPAL

V

EIN

A

CCESORY

V

EIN

C

OURSE OF

A

CCESORY

V

EIN

F

EATURES OF THE

T

OOTH

A

PEX

VENATION OF THEPINNAE AND/ORPINNULE

#

OF

B

ASAL

C

OSTA

I

NTERIOR

S

ECONDARIES

fimbrial vein

P

ERIMARGINAL

V

EINS

slightly decurrent

M

AJOR

2° V

EIN

A

TTACHMENT

DIAGNOSTICFEATURES

Frond organization unknown. Laminar size from nanophyll to mesophyll. Frond shape elliptic. Margin entire. Costa framework pinnate, very thick (0.3-0.4 cm) compared to secondaries (<0.1 cm). Secondary veins reticulodromous, weekly decurrent on costa, closely spaced (from 0.07 to 0.13 cm), and arise from the costa from 40° to 60°. Fimbrial vein. Polygonal areoles become smaller near costa and at the margin, but more elongated in between. The areoles are 3-5 sided free of veinlets, and their sizes range from 05-1.0 cm x 0.07-0.13 cm.

SIMILARMORPHOTYPES

untoothed

B

ASE

S

YMMETRY

A

PEX

S

HAPE

B

ASE

S

HAPE

M

ARGIN

T

YPE

A

PEX

A

NGLE

B

ASE

A

NGLE

L

OBATION

F

ROND

S

HAPE elliptic

M

ARGIN

F

EATURES

unlobed

V

EINLETS

T

ERMINAE

G

ENUS Acrostichum

R

HIZOME

T

YPE

R

HIZOME

L

ENGHT

S

PECIAL

F

EATURES

FRONDCHARACTERS

F

ROND

T

YPE

P

ETIOLE

F

EATURES

F

ROND

A

TTACHMENT

F. O

RGANIZATION

FEATURES OF THEPINNAE

S

CALES

& H

AIRS

C

OSTULE

F

RAMEWORK

#

OF

B

ASAL

C

OSTA

SPORANGIA

S

ORI AND

I

NDUSIA

S

ORUS

P

OSITION

VASCULARBUNDLES

T

YPE

DESCRIPTION

Frond organization unknown. Costa framework pinnate. Margin entire. Secondary

veins reticulodromous that anastomose to form small polygonal areoles without veinlets.

S5%$"$%"%

PLANT FAMILY/ORDER

M

AJOR

P

LANT

G

ROUP

FIU O

RGAN

T

YPE

)%"* T

YPE LOC

. # T

YPE SPEC

. #

L

OCA

6718812%0 B

ASE

S

YMMETRY

A

PEX

S

HAPE

B

ASE

S

HAPE

M

ARGIN

T

YPE

(9_^

incertae sedis ^M

^ORPHOTYPE

^#

M

ORPHOTYPE

N

AME

M

ARGINAL

U

LTIMATE

V

ENATION

A

REOLATION

F.E.V.

S

C

OURSE

S

TOMATA

C

UTICLE

F

EATURES

L

AMINAR

S

IZE

(

SMALLEST

)

;"5",,%,8058486+

I

^NTER

-2° V

^EIN

P

^ROXIMAL

C

^OURSE

V

ARIATION OF

2° V

EIN

A

NGLE

(

COSTAL

) A

GROPHIC

V

EINS

M

AJOR

2° V

EIN

F

RAMEWORK

(

COSTAL

)

M

AJOR

2° S

PACING

(

COSTAL

) 1° V

EIN

F

RAMEWORK

M

ORPHOTYPER

L

AMINAR

L:W R

ATIO

P

ETIOLE

F

EATURES

Q

UATERNARY

V

EIN

F

ABRIC

Q

UINTERNARY

V

EIN

F

ABRIC

I

NTERCOSTAL

3° V

EIN

F

ABRIC

A

NGLE OF

P

ERCURRENT

3°

P

RINCIPAL

V

EIN

T

ERMINATION

T

OOTH

S

PACING

T

OOTH

S

HAPE

#

^OF

O

^RDERS

T

EETH

/

CM

S

^INUS

S

^HAPE

L

EAF

A

TTACHMENT

A

PEX

A

NGLE

B

ASE

A

NGLE

L

OBATION

CERREJON FLORA

SECTIONI. LEAFCHARACTERS

L

EAF

A

RRANGEMENT

L

^EAF

O

RGANIZATION LEAFLETORGANIZATION LEAFLETATTACHMENT

FEATURES OF THEBLADE

P

OSITION OF

B

LADE

A

TTACHMENT

L

AMINAR

S

IZE

(

LARGEST

)

L

AMINAR

S

HAPE

M

EDIAL

S

YMMETRY

M

ARGIN

F

EATURES

T

ERMINAL

A

PEX

F

EATURES

S

URFACE

T

EXTURE

S

URFICIAL

G

LANDS

SECTIONIII. TEETH

T

^OOTH

S

^HAPE

P

RINCIPAL

V

EIN

A

^CCESORY

V

^EIN

C

^{OURSE OF}

A

^CCESORY

V

^EIN

F

EATURES OF THE

T

OOTH

A

PEX

SECTIONII. VENATION

N

AKED

B

ASAL

V

EINS

#

^OF

B

^ASAL

V

^EINS

I

NTERIOR

S

ECONDARIES

M

INOR

2° C

OURSE

P

ERIMARGINAL

V

EINS

M

AJOR

2° V

EIN

A

TTACHMENT

(

COSTAL

)

I

NTER

-2° V

EIN

L

ENGHT

I

^NTER

-2° V

^EIN

D

^ISTAL

C

^OURSE

I

NTER

-2° V

EIN

F

REQUENCY

;"5",,%,8058486+

C

^{OURSE OF}

P

^ERCURRENT

3°

I

^NTERCOSTAL

3° V

^EIN

A

^NGLE

V

^ARIABILITY

E

PIMEDIAL

3°

A

^DMEDIAL

C

^OURSE

E

XMEDIAL

C

OURSE

E

^XTERIOR

3° C

^OURSE

DIAGNOSTICFEATURES

(8+1";",4"1% 85 ;",4"1% ,%"?%+< F"5=/7 %71/5%< E%=4%71+ ./0% ./12 " 4/0?%/7< !865 850%5+ 8*

;"5",,%, ?%/7+< (58++?%/7+ ./0%> +15/T/7=,3 +/7686+> ./12 "7 /55%=6,"5 $865+%> ;"5",,%, 18 87%

"7812%5<

SIMILARMORPHOTYPES

F.E.V.

S

T

ERMINAE

*5%%,3-5"4/*3/7=

4%+8;23,, 4%=";23,,

!5"=4%71+ 8* $8+1";",4"1% 85 ;",4"1% ,%"?%+< F"5=/7 Z71/5%< E%=4%71+ PY:<_j::M 44 ./0% ./12

" 4/0?%/7 :<Y 44 ./0%< B"5",,%, 18 12% 4/0?%/7> 12%5% "5% ?%/7+ 8* *865 850%5+h */5+1 850%5 ?%/7+

:<:^j:<_ 44 ./0%> +%;"5"1%0 #3 AQ@AQ<ML 44[ +%$870 850%5 ?%/7+ K<Q@:<: 44 ./0%[ ;5/4"53 "70 +%$870"53 ?%/7+ +%;"5"1%0 #3 K<M 44 ";;58D/4"1%,3[ 12/50 850%5 ?%/7+ K<:@K<M 44 ./0%>

+%;"5"1%0 #3 M<KjA<XX 44< G%51/"53 "70 +%$870"53 ?%/7+ +%;"5"1%0 #3 _jQ 44< !86512 850%5

?%/7+ ?%53 12/7> ",48+1 /70/+$%57/#,%> +%;"5"1%0 #3 K<M 44 "70 ";;58D/4"1%,3 +%?%7 /7 764#%5

#%1.%%7 1.8 12/50 850%5 ?%/7+< (58++?%/7+ K<:KjK<:M 44 ./0%> +15/T/7=,3 +/7686+> ./12 "7 /55%=6,"5 $865+%> ;"5",,%, 18 87% "7812%5 #%1.%%7 %"$2 812%5> +;"$%0 :@Y 44 ";"51 "70 V8/7/7= 12%

4/00,% 85 ;5/4"53 ?%/7+ 0%$655%71,3 "1 " XYH "7=,%< S."3 *584 12%/5 0%$655%71 V67$1/87 ./12 12/$T%5 ?%/7+> 12% $58++?%/7+ #%70 18 $58++ 12% +%$870"53 ?%/7+ "1 "7 ",48+1 +15"/=21 "7=,%[ 12%3

$"7 $877%$1 1.8 ;5/4"5/%+ 85 " ;5/4"53 "70 " ,%++%5 850%5 ?%/7< (58++?%/7+ "5% 781 ",."3+ ?/+/#,%>

87,3-/7-+84%-,%"*-*5"=4%71+-.2%5%-;"51/",-0%$84;8+/1/87-188T-;,"$%<

DESCRIPTION

#

S5%$"$%"%

PLANT FAMILY/ORDER

M

AJOR

P

LANT

G

ROUP

FIU O

RGAN

T

YPE

)%"* T

YPE LOC

. # T

YPE SPEC

. #

L

OCA

6718812%0 B

ASE

S

YMMETRY

A

PEX

S

HAPE

B

ASE

S

HAPE

M

ARGIN

T

YPE

(9_Q

incertae sedis ^M

^ORPHOTYPE

^#

M

ORPHOTYPE

N

AME

M

ARGINAL

U

LTIMATE

V

ENATION

A

REOLATION

F.E.V.

S

C

OURSE

S

TOMATA

C

UTICLE

F

EATURES

L

AMINAR

S

IZE

(

SMALLEST

)

;/77"1%

;"5",,%,8058486+

I

^NTER

-2° V

^EIN

P

^ROXIMAL

C

^OURSE

V

ARIATION OF

2° V

EIN

A

NGLE

(

COSTAL

) A

GROPHIC

V

EINS

M

AJOR

2° V

EIN

F

RAMEWORK

(

COSTAL

)

M

AJOR

2° S

PACING

(

COSTAL

) 1° V

EIN

F

RAMEWORK

M

ORPHOTYPER

L

AMINAR

L:W R

ATIO

P

ETIOLE

F

EATURES

Q

UATERNARY

V

EIN

F

ABRIC

Q

UINTERNARY

V

EIN

F

ABRIC

I

NTERCOSTAL

3° V

EIN

F

ABRIC

A

NGLE OF

P

ERCURRENT

3°

P

RINCIPAL

V

EIN

T

ERMINATION

T

OOTH

S

PACING

T

OOTH

S

HAPE

#

^OF

O

^RDERS

T

EETH

/

CM

S

^INUS

S

^HAPE

L

EAF

A

TTACHMENT

A

PEX

A

NGLE

B

ASE

A

NGLE

L

OBATION

CERREJON FLORA

SECTIONI. LEAFCHARACTERS

L

EAF

A

RRANGEMENT

L

^EAF

O

RGANIZATION

8;;8+/1%-18-+%4/",1%57"1%

LEAFLETORGANIZATION

+%++/,%

LEAFLETATTACHMENT

$84;8670

FEATURES OF THEBLADE

P

OSITION OF

B

LADE

A

TTACHMENT

L

AMINAR

S

IZE

(

LARGEST

)

L

AMINAR

S

HAPE

M

EDIAL

S

YMMETRY

M

ARGIN

F

EATURES

T

ERMINAL

A

PEX

F

EATURES

S

URFACE

T

EXTURE

S

URFICIAL

G

LANDS

SECTIONIII. TEETH

T

^OOTH

S

^HAPE

P

RINCIPAL

V

EIN

A

^CCESORY

V

^EIN

C

^{OURSE OF}

A

^CCESORY

V

^EIN

F

EATURES OF THE

T

OOTH

A

PEX

SECTIONII. VENATION

N

AKED

B

ASAL

V

EINS

#

^OF

B

^ASAL

V

^EINS

I

NTERIOR

S

ECONDARIES

M

INOR

2° C

OURSE

P

ERIMARGINAL

V

EINS

M

AJOR

2° V

EIN

A

TTACHMENT

(

COSTAL

)

I

NTER

-2° V

EIN

L

ENGHT

I

^NTER

-2° V

^EIN

D

^ISTAL

C

^OURSE

I

NTER

-2° V

EIN

F

REQUENCY

;"5",,%,8058486+

C

^{OURSE OF}

P

^ERCURRENT

3°

I

^NTERCOSTAL

3° V

^EIN

A

^NGLE

V

^ARIABILITY

E

PIMEDIAL

3°

A

^DMEDIAL

C

^OURSE

E

XMEDIAL

C

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E

^XTERIOR

3° C

^OURSE

DIAGNOSTICFEATURES

B/77"1% ,%"?%+< F"5=/7 %71/5%< k"$2/+ +1861 ./12 " ,87=/160/7", 5/0=%< )%"*,%1+ "5% ;,/$"1%>

5%=6,"5,3 "55"7=%0 /7 87% ;,"7%> 8;;8+/1% 18 +%4/",1%57"1%< G.8 850%5+ 8* ;"5",,%, ?%/7+<

G5"7+?%5+%-?%/7,%1+-"5%-781-%?/0%71<

SIMILARMORPHOTYPES

F.E.V.

S

T

ERMINAE

4%+8;23,, 4%=";23,,

!5"=4%71+ 8* ;/77"1% ,%"?%+< F"5=/7 %71/5%< k"$2/+ +1861 JPMA 44 ./0% "1 12% #"+% "70 ^@:^ 44 ./0% /7 485%

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$61/$,% *5"=4%71+ .%5% %D15"$1%0 *584 1.8 +;%$/4%7+> "70 $87+/+1%0 8* 1.8 13;%+h (61/$,%+ ./12 +184"1"> $%,,+

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,87=/160/7",,3 18 0/"=87",,3 %D1%70%0 J7%"5 18 12% +184"1"N[ 15/$284%+ "#+%71[ +184"1" /7 787 0/+1/7$1/?% ,/7%+<

l6"50 $%,,+ M:<^Q@MY<YM!4 ,87= "70 Y<^M@Q<YQ!4 ./0%[ ,"1%5", +6#+/0/"53 $%,,+ MY<L_@AM<Y_!4 ,87= "70 M<_X

@:K<Y_!4 ./0%[ 1%54/7", +6#+/0/"53 $%,,+ ./12861 ,8#%+> L<X_@:X<L_!4 ,87= "70 :M<YX@:^<AQ!4 ./0%< (61/$,%+

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DESCRIPTION

#

!"#"$%"%

PLANT FAMILY/ORDER

M

AJOR

P

LANT

G

ROUP

&'( O

RGAN

T

YPE

)%"* T

YPE LOC

. # T

YPE SPEC

. #

L

OCA

./012-"70-/7+%51/87-"+344%1<

6718812%0 B

ASE

S

YMMETRY

A

PEX

S

HAPE

B

ASE

S

HAPE

M

ARGIN

T

YPE

(9^_

incertae sedis ^M

^ORPHOTYPE

^#

M

ORPHOTYPE

N

AME

M

ARGINAL

U

LTIMATE

V

ENATION

A

REOLATION

F.E.V.

S

C

OURSE

S

TOMATA

C

UTICLE

F

EATURES

L

AMINAR

S

IZE

(

SMALLEST

)

;/77"1%

I

^NTER

-2° V

^EIN

P

^ROXIMAL

C

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V

ARIATION OF

2° V

EIN

A

NGLE

(

COSTAL

) A

GROPHIC

V

EINS

M

AJOR

2° V

EIN

F

RAMEWORK

(

COSTAL

)

M

AJOR

2° S

PACING

(

COSTAL

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F

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M

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L

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L:W R

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;%1/8,%-#"+%-;6,?/76,"1%

P

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F

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Q

UATERNARY

V

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F

ABRIC

Q

UINTERNARY

V

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F

ABRIC

I

NTERCOSTAL

3° V

EIN

F

ABRIC

A

NGLE OF

P

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3°

P

RINCIPAL

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T

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T

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S

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T

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S

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#

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S

^INUS

S

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L

EAF

A

TTACHMENT

"$61%

A

PEX

A

NGLE

8#16+%

B

ASE

A

NGLE

L

OBATION

CERREJON FLORA

SECTIONI. LEAFCHARACTERS

8;;8+/1%

L

EAF

A

RRANGEMENT

L

^EAF

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SIMILARMORPHOTYPES

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FEATURES OF THEBLADE

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SIMILARMORPHOTYPES

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DESCRIPTION

+48812,3-/7$5%"+/7=-;58D/4",,3 0%$655%71

#

P

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CJ81

incertae sedis ^M

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M

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N

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lacking

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pinnate

craspedodromous

regular - crowded

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irregular

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variable, 1-4/cm

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angular

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CERREJON FLORA

RHIZOMECHARACTERS

mesophyll

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macrophyll

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TEETHFEATURES

st/st st/cv

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absent

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present

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DIAGNOSTICFEATURES

Frond organization varies from pinnatisect (more proximally) to pinnatifid (more distally). Frond size mesophyll to macrophyll. Pinnae organization mostly alternate.

Pinnae attachment sessile. Frond shape of the pinnae oblong (the longest specimen is ~17 cm). Pinnae base shape asymmetrical and strongly decurrent. Secondary veins craspedodromous, numerous, closely spaced (0.3 to 0.4 mm), with rare and low angle dichotomy near costa, and decurrent or excurrent on costa. Lacking areolation. Margin serrate; teeth (st/st and st/cv) with long proximal flanks; tooth spacing irregular;

number of orders one; teeth/cm variable, 1-4/cm; sinus shape angular; principal vein absent, but more than one accessory vein enters the tooth with courses straight; features of the tooth apex non-specific.

SIMILARMORPHOTYPES

asymmetrical

serrate

B

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S

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A

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S

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B

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S

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M

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acute

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OBATION

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unlobed lobed

decurrent

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FEATURES OF THEPINNAE

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CALES

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SPORANGIA

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VASCULARBUNDLES

T

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DESCRIPTION

Frond organization varies from pinnatisect (more proximally) to pinnatifid (more

distally). Pinnae oblong (up to ~17 cm long). Secondaries craspedodromous and closely

spaced. Lacking areolation. Teeth st/st and st/cv.

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