TRILOBITA
R. I W., 4 miles southeast of Hennepin, Arbuckle Mountains, Oklahoma
Cotypes.
— U.S.N.AL
no. 93025.Elvinia missouriensis, n. sp.
Eh'inia rocmcri Bridge (part), U. S. Geol. Surv. Prof. Pap. 186-M, p. 2511, pi. 69, figs. 12, 13, 1937.
32
SMITHSONIAN MISCELLANEOUS COLLECTIONS
VOL. 97 Bridge figured one of tlietwo
species present at the locality.The
holotype is a well-preserved cranidium.
This species is characterized
by
its relative narrownessimmedi-
ately in front of the eyes.
The
eyes are not parallel to the dorsal fur- row, butmake
a wide angle with the axis. Eyelines are rather heavy.The
cranidium isarched considerablylaterally, this being accentuated by thepronounced downward
depression of the anterior angles of the brim. Longitudinally the head is also highly arched, the greatest curvature being in the anterior half.The
preglabellar area is bulged a bitand
the thickened rim is sharply upturned.Upper
Cambrian, Uavis; (loc. iik) near shaft of FederalLead Mine No.
4, Flat River, Missouri.Holotype.
— U.S.N.M.
no. 93011.Elvinia dakotensis, n. sp.
Elvinia rocmcri Bridge (part), U. S. Geol. Surv. Prof. Pap. 186-M, p. 251, pi. 69, figs. 14, 16, 1937.
This species is nearest likeE. missouriensis, dififering in the round- ing of the anterior portion of the glabella
and
the slightlynarrower
width immediately in front of the eyes. E. dakotensis also has lessrugged
relief in the various portions of the brim. In E. dakotensis the glabella does not rise somuch
as the fixigenes,and
the longi- tudinal curvature,which
is considerable,forms
a rather even curve.The
libragenes are largeand
wide, with a strongly curved outer margin, particularlytoward
thegenal angle.The
genal spine is shortand
small.The
associatedpygidium
has an axiswhich
stands well above the flat pleural lobes.Upper
Cambrian,Deadwood
; (loc. 88a) northern part ofDead-
wood, Black Hills,South
Dakota.Cotypes.
— U.S.N.M.
no. 93024.Elvinia utahensis, n. sp.
Elvinia rocmcri Bridge (part), U. S. Geol. Surv. Prof. Pap. 186-M, p. 251, pi. 69, fig. 22, 1937.
This species is characterized
by
rugged reliefdue
to the depth ofall furrows, the arching of the preglabellar area
and
of the fixigenesand
the depression of the anterior angles of the brim.The
eyes are largeand
situated rather far forward.Upper
Cambrian,Orr;
(loc. 32t)Fandango
Spring Canyon, east sideDugway
Range, Utah.Holotype.
— U.S.N.M.,
no. 93026.NO. lO
CAMRRIAN
FOSSILS,4TH CONTRIBUTION
RESSER 33 Elviniamatheri (Walcott)Ptychoparia mafheriWalcott, Smithsonian Misc. Coll., vol. 57, no. 9, p. 268, pi. 44, figs. 15-17, 1912; Bridge (part), U. S. Geol. Surv. Prof. Pap.
186-AI, p. 254, pi. 69, fig. II, 1937.
Upper
Cambrian,Potsdam;
(loc.no)
east side of Whitehall,New
York.Cotypcs.—U.S.^M.
nos. 58585-7.GENE
VIEVELLA
Lochman, 1936 GcncvicveUaLochman,
Journ. Pal., vol. 10, no. i, p. 40, 1936.Llanoaspis Lochman, idem, vol. 12, no. i, p. 80, 1938.
When
this genuswas
established, only the genotypewas known.
Two
specieswere added from
Texas,and
the descriptions of elevenfrom
the Appalachians are in press; consequently this is a well- represented trilobite genus.Thus
far the genus is confined to the Crepicephalus zone.Llanoaspis
modest
aand
L. undiilataLochman become
Gene- vievella.IRVINGELLA
Ulrich and Resser, 1924Irvingella
was
applied to specieswhich
differfrom
Chariocephalus in a limited degree. Chariocephalus has large eyes situated rather far forward.The
facial suture joins the dorsalfurrow
anterior to the eye, bywhich arrangement
the anterior fixigene is eliminatedand
thebrim
is caused tobe a singlebar not attached to the lixigenes at theanterior angles.On
the otherhand
Irvingella has e\'en larger eyes than Chariocephalus, the extra length being attained by the rear portion of the palpebral lobes extending farther back.At
thesame
time the eyes are slantedoutward from
the axis of the head.Above
all,Irvingelladiffers
from
CJiariocephalusby havinganterior fixigenes which unite with the brim.Irvingella tumifrons (Hall and Whitfield)
Chariocephalus tumijrons H.\ll and Whitfield, U. S. Geol. Expl.40t]i Par., vol. 4, p. 224, pi. 2, figs. 38, 39, 1877.
Upper
Cambrian, SecretCanyon; Pogonip
^Mountain,White
Pine District, Nevada.Holofype.
— U.S.N.M.
no. 24561.KOCHINA
Resser, 1935 Kochina? lux (Walcott)Ptychoparia lux Walcott, Smithsonian Alisc. Coll., vol. 67, no. 3, p. 90, pi. 12, fig. 5, 1917.
34
SMITHSONIAN MISCELLANEOUS COLLECTIONS
VOL. 97Lower
Caml)rian,Mount Whyte
; (loc. 6id) southwest slopeMount
Shaffer, British Columbia.
IIoIotypc.—U.S.^M.
no. 64387.LONCHOCEPHALUS
Owen, 1852 Lonchocephalus verrucosus (Whitfield)Conoccphalites vcvntcosus Whitpield, Amer. Mus. Nat. Hist. BuH. i, p. 146, pi. 14, figs. 9-12, 1884.
Upper
Cambrian,Potsdam;
near "Post office." AusableChasm,
New
York.CoO'/'^-^—
A.M.N. H.
no. 280.MENOMONIDAE
WalcottMENOMONIA
Walcott, 1916Mcnomonia Walcott, Smithsonian Misc. ColL, vol. 64, no. 3, p. 161, 1916.
MUlardia Walcott, idem, p. 163, 1916.
Walcott referred three genera to the Proparian family
Menomo-
nidae, but he failed to recognize that
two
ofthem
were identical.Evidence for a primitive aspect about
Mcnomonia and
Drcsbachiamay
be questioned. Evidently Walcott based his idea of this on thenumerous
thoracic segments ofM.
calynicnoides.Examination
of availablespecimens failstoprove that this extraordinarily longthorax belongs toMcnomonia,
although association in three instances sug- gests the possibility.Even
if thegenotype has 42 thoracic segments, one can hardly consider this alone as evidence of primitive structure.It
seems
rather that theMenomonidae
are a highly specializedgroup
descendedfrom
speciesnow
placed in Alokistocarc.Menovwnia
is a characteristic trilobite of the earlyUpper Cam-
brian Ccdaria zone; thus far being confined to that zone. Whitfield's original specimens of the type species are
on
small pieces of rock with the types of Ccdaria woosfcri.MUlardia was
distinguishedfrom Mcnomonia
by differences ofbrim and
in having fewer thoracic segments.There
is a difference in thebrim
of AI. calymcnoidcsand
ofM.
scmclc but it involves no altered structureand
thegap between
thetwo
species is bridged by intermediate forms.M.
calymcnoidcs has amuch
swollen brim, whereas other species obtain thesame
rigidity by arching the short brim.Many
specimens referred toMUlardia
retain the libragenes, givingthem
a different aspect,which
caused their reference to a separate genus.Drcsbachia is a distinct, but closely related genus.
Compared
withMenomonia,
the glabella is essentially thesame
but the fixigenes areNO. 10
CAMBRIAN
FOSSU.S,4TH CONTRIBUTION
RESSER 35much
larger.The
large Hbragenes—
the part of this trilobitemost
commonly
preserved—
are attached to a very short brim, which isreduced to little
more
than a line, terminating in a swollenknob
like the rimMenouwnia.
In otherwords
Dreshachia is characterized by the lateral shortening of the preglabellar area to amere
line for the attachment of the Hbragenes which lieswholly in front of theglabella.Besides thegenotype and the species formerlyreferred toMillordia, several
new
speciesfrom
the Appalachian Nolichucky formation are in press.Menomonia avitas (Walcott)
Millardia avitas Walcott, Smithsonian Misc. Coll., vol. 64, no. 3, p. 165, pi. 28, figs. 5-5e, 1916.
Upper
Cambrian,Warrior;
(loc. 107k) 2 miles north of Benore, Center County, Pennsylvania.Holotype.—U.S.N. M.
no. 61629.Menomonia magnagranulata (Lochman)
Millardia iiiagiiagrauiilata Lochman, Journ. Pal., vol. 12, no. i, p. 84, pi. 18, fig- 3, 1938.
Upper
Cambrian,Cap Mountain;
southwest side of LionMoun-
tain,
Highway
29, 9 miles northwest of Burnet, Texas.Holotype.
— U.S.N.M.
no. 95015.Menomonia optata (Hall)
Coiioccphalitcs optatus Hall, i6th Ann. Rep.
New
York State Cab. Nat.Hist., p. 222, pi. 5A, fig. 7, 1863. Trans. Albany Inst.,vol. 5, p. 195, 1867.
Ptychoparia optata Miller, North Amer. Geol. Pal., p. 539, 1889.
Millardia optata Walcott. Smithsonian Misc. Coll., vol. 64, no. 3, p. 165, pi. 28, figs. 4, 4a-f, 1916.
One
cannot be certain that Hall's original typeand
the specimens figuredby
Walcott represent thesame
species.At
any rate Hall's original is a very small specimen.Upper
Cambrian,Eau
Claire;Willow
River Falls, Trempealeau,and
other localities in Wisconsinand
Minnesota.Holotype.
— A.M.N.H.
no.322;plesiotypes,U.S.N.M.
nos. 61626-8.Menomonia semele (Walcott)
Millardia scmelc Walcott, Smithsonian Misc. Coll., vol. 64, no. 3, p. 166, pi.28, figs.3-3C, 1916.
Upper
Cambrian.Weeks
; (loc. 30 n)Weeks
Canyon,House
Range, Utah.Cotypes.-AJ.S.N.M.
nos. 61622-5.36 SMITHSONIAN MISCELLANEOUS COLLECTIONS
VOL. 97 Menomonia texana, n. sp.Millardia az'itas
Lochman
(not Walcott), Journ. Pal., vol. 12, no. i, p. 84, pi. 18, figs. 4-S, 1938.This species evidently
was
referred toM.
avitas because it has smallergranulesthanM.
nmgnagranulata.The
referencetothePenn-
sylvania species cannot stand.M.
avitas has afew
scattered large granules on the glabellaand
fixigenes; the remainder of the cephalonis
smooth
to finely granulose orornamented
with lines.The
eyes ofM.
avitas are elevatedon
rather long stalks. Thismay
also have been the case inM.
texana as the eyes are broken off.M.
texana isfurther characterized
by
a long slender glabellaand
the eyes are rather far forward.Upper
Cambrian,Cap Mountain
; southwest sideLion
Mountain,Highway
29, 9 miles northwest of Burnet, Texas.Holotype.
— Mount Holyoke Mus.
no. 655; paratype, no. 656.Menomonia lochmanae (Lochman)
Millardia avitas
Lochman,
Journ. Pal., vol. 12, no. 5, p. 469, pi. 56, figs. 27, 28, 1938.This species differs
from M.
avitas in themore
incurved anteriormargin and
considerably larger fixigenes.Upper
Cambrian, Petit Jardin;Cape
St. George,Newfoundland.
Holotype.
—
YalePeabody Mus.
no. 15822.NORWOODIA
Walcott, 1916Norwoodia Walcott, Smithsonian Misc. Coll., vol. 64, no. 3, p. 168, 1916.
IVhitfieldina Resser, idem, vol. 95, no. 22, p. 27, 1937.
Just after the 1937 paper appeared, the genus
Nonvoodia was
re- considered in conjunction with studies in the southern Appalachians.As
soon as the extraneous species then inNorwoodia were
removed,it
became
clear that IVhitfieldinais not a valid genus,because its type species is a typicalN
orzvoodia, contrary to the misleading publishedfigures.
Norwoodia quadrata (Whitfield)
Conocephalites qnadratns Whitfield, Ann. Rep. for 1879, Wisconsin Geol.
Surv., p. 47, 1880; Geol. Wisconsin, vol.4, p. 180, pi. i, figs. 15, 16, 1882.
IVhitfieldina quadrata Resser, Smithsonian Misc. Coll., vol. 95, no. 22, p. 27, 1927.
Upper
Cambrian,Eau
Claire;Eau
Claireand
other localities in Wisconsin.Cotypes.
—
Univ. Wisconsin.i
NO. lO
CAMBRIAN
FOSSILS,4TH CONTRIBUTION
RESSER 37OLENOIDES
Meek, 1877Olenoides expansus (Walcott)
Dicellocephahis ? expansus Walcott, U. S. Geol. Surv. Mon. 8, p. 45, pi. 9, fig. 19, 1884.
Dolichometopus ?expansus Walcott, Smithsonian Misc. Coll., vol. 64, no. 5, p. 368, pi. 53, figs. 5, 5a, 1916.
Middle
Cambrian, Eldorado; (loc. 55b) east slope ProspectMoun-
tain,
Eureka
District, Nevada.Holotype.—U.S.'NM.
no. 15450.ORYCTOCEPHALUS
Walcott, 1886Species of this genus have not been carefully discriminated, and specific
names were
very carelessly used.A
brief revision clears theway
for*erection of the necessarynew
species.Oryctocephalus,as
now
constituted,containstwo
groups of species.The
validspeciespreviouslyrecognizedarelistedunderthetwo
groups.Groupof O. primus (genotype) O. zvalcotti Resser O. primus Walcott O. salteri Reed O. burgessensis, n. sp. O. indicus (Reed)
O. ivalkcri Matthew O. kobayaski Saito
Group of O. rcynoldsi O. orientalis Saito O. reynoldsi Reed O. saitoi, n. sp.
In addition, one undescriljed species of the 0. priunis group
and
several of the O. rcynoldsigroup
arein hand.Oryctocephalus burgessensis, n. sp.
Oryctoccphahts primus Kobayashi, Journ. Faculty Sci. Imp. Univ. Tokyo,
sec. 2, vol. 4, pt. 2, p. 147, pi. 15, fig. I, 1935.
Kobayashi
pickedup
a photographin the NationalMuseum
collec- tionsand
published it as 0. priuins, but this illustrationwas
pre- pared by Walcott because he recognized that itwas
not 0. primus.0. bitrgcssoisis is the
most
abundant species of the genus in the Burgess shale. It is characterized by a normal cranidium, seven tho- racic segments,and
slender pygidial spines of even length.The
rear segments of thepygidium
are turned back practically parallel to the axis.The
pygidial spines of even length cause this species to lookmuch
like O. n'cilkcri, hut the rear pygidial segments are turned backmore
sharply.Middle
Cambrian, Burgess; (loc. 35k) Burgess Pass, near Field, British Columbia.Holotypc.—V.S.NM.
no. 96487.38