Station3362, in 117.5 fathoms,
mud,
off Cocos Island in the GulfofPanama
; bottom temperature,36.8° F. No. 107275, U.S.N.M.A
singlespecimenof thismodest littleshellwas
obtained. It differsfrom L. davidsonl,
Adams,
in its incomplete foramenand
lessflexuous margin; itis less inflatedand somewhat
smaller.From
theyoung
of L. nva, Broderip, which is (notwithstanding it has been called"smooth") a finely regularly microscopically striate shell,it
maybe
distinguished
by
its smooth surface. L. arctica, Friele,isless like itthan Bavidsoni is;and, in short, while the shell has no very
marked
characters,it can not be safely referred to
any
described species.Though
small,by
thesolidity of its hinge armatureand
themanner
inwhich the foramen is
worn by
the peduncular motion, itwould seem
to bean adult shell.
Though
the shellwas
in adrj^ conditionwhen
takenup
for study, the softpartswerewellpreserved,and
present thepeculiarityofhaving thelateralbands
of brachia quiteclose to oneanotherand
rather long;the space between
them
is smoothand
occupiedby
astretchofmem-
branous tissue,while thecentralwhorlof brachials below, and, looking verticallydown upon
the valve placed horizontally, is invisible, the cirrhi, of course, being contractedby
drying. It is only on looking sidewiseat the valve that thecoil is seen under themembrane
above mentionedand
lower thanthe lateral brachial loops.On
soaking the remains in fresh water theyexpanded
considerablyand assumed
a fairly natural elasticity, but the relative position of themedian
brachial coilremainedthesame.The
externalappearanceof thissliell isalmost exactly like that of Macandrevia cranium of thesame
size.The
species isnamed
in honorof J. M. Clarke,esq., associateof Prof.Hallin therevision of the Paleozoic Brachiopoda.
Itis not likelythatthisspeciescan berelatedtolAothyrinastearnsii, Ball
and
Pilsbry, whichis a native ofJapan
(pi. xxx, figs. 8, 9, 11), as that species has a complete foramen, but the figures are given for comparison with the otherspecies.Genus
TEREBRATULINA, Orbigny.
TEREBRATULINA. CAPUTSERPENTIS, L
in nseus.Plate XXXII,figs.2,5.
TerebratuUna caputserpentis var. unguicula, Davidson, Mon. Rec.Brach.Pt.i,p.
25,1886.
Terehratnla unguiatla, Carpenter, Proc. Zool.Soc,1865,p. 201, figs.
1^.—
Dall, Am.Jouni. Conch.,vi,1870,p.102.Stations2849, 3311, 3330, 3350,
and many
others, in from low waterto 500 fathoms, temperature 40° to 44° F., from the southern part of Bering Seasoutheast to the coastof Californiainlatitude33° N.,and
southwesttoJapan and
Korea. Also the NorthAtlantic, the upper Tertiary rocks ofEurope,etc. Figured specimen No.123155,U.S.N.M.For
some
time Iwas
disposed to regard theNorth Pacificform as720 DEEP WATER MOLLlSKS AND BBACHDJPODS—DALL.
vol.xvil distinct from that of the Atlantic, following Canjeuter, likewhom
Ihad
only the stunted specimens from shallow water.But
the deep- water dredj^ings of the Albatross having supplied a sufficient series of normallygrown
specimens of all agesand
sizes, I have convinced myself, after a thorough comparison ofmany
specimens from each region, that there is no good ground for a separation of them, even varietally.The
Pacific form isthe typical T. caxndserpentis (not the easternAmerican
septentrionaUs)and
neitherin the shells norin the soft parts does there appearto beany marked
or constantdifference.They
could notbe separated ifoncemixed
in thesame
tray.TEREBRATULINA
KIIENSIS, Ball and Pilsbry, Plate XXXII, figs.8,9.Terehratulina (nnguicula,Carpentervar.f) kiicnsis,DAi.Land Pilsbry,NautilKS, V,p. 18, pi. 1, tigs. 4, 5,1891.
Stations2871,3316,
and
3205, in559 fathoms,oft" the entrancetoFuca
Strait; in309fathomsoft"theislandofUnalaska in Bering Sea;
and
in 240 fathoms oft' Santa Cruz, Cal.; temperature, 38° to 44° F. Also from the coast of the province of Kii, Japan, Stearns;and
from the PhilippineIslands,NE.
from Mindanao,in 82fathoms. Challengerex-pe- dition. Figured specimen No. 128463, U.S.N.M. 'This fine brachiopod, which
when young
approaches closelysome
broadvarieties ofT. caj)utserpentis, isshown by
the Albatross material to be a distinct species. Itmay
beknown by
its rounder outline, largersize,and
thefactthat the sculpture of the peripheral parts of theshellbecomesobsolete,and
isrepresentedby
grooveswithflattened,much
widerinterspaces, instead of theroundedthreads,characteristic of the surface of T. caputserpentis at all ages,and
T. Kiiensiswhen
young.The
extension of its range,made known by
the Albatross dredgings,isveryremarkableand
interesting.Family Terebratellid^.
Genus
EUDESIA, King.
This
name was
proposed at thesame
time as Waldheimia,King
[=Ma()eUania, Bayle), whichwas
discarded as preoccupied. Subse- quently it has been treated as a subgenus of thenewer name
Magellania. Beecher has
shown
that the austral forms typifiedby
Magellania, on account of their dift'erent development,must
be sepa- rated in adifferent subfamily from thoseof thenorthern hemisphere.Eudesia belongs with the latter. These again are separable into at least twogeneric groups, DaUlna,of Beecher(apparently adescendant ofAntiptychina,Zittel),whichincludes thosewitha continuous cardinal plate, strong median septum in the brachialvalve, and no buttresses tothe teeth of the pedicle valve.
The
other groupcomprises Eudesiaand
Macandrevia, and has the cavity of the pediclevalve under the hinge separated into three cavitiesby two
buttresses which support1894.
PROCEEDINGS OF THE NATIONAL MUSEUM. 721
theteeth. In the brachial valve the cardinalplateisdivided medially
by
asinus.The
cardinal processisobsolete,and
the medialseptum
eitherwholly absent or represented only
by
a short, low ridge not attached to the cardinal buttresses,and
not extending- forwardinto thecavity of the valve between the buttresses.As
EiuJesia ismuch
theolder
name
itmust
be retained forthese forms, unless Macandrevia exhibits characters strongenough
to give it generic independence.So far as known, the only ditterences between
them
consist in the smoothsurface of thevalvesand
the feebleness of the brachialseptum
inMacandrevia. These can hardly raise the latter above subgeneric rank, as the plication of the valves is often inconstant in the
same
species,while the
same
si^ecies (and doubtless thesame
individual) at different ages willshow
a septummore
or lessdeveloped, from quite obsolete in the young, to quite perceptible inthe senile stag'e. Ihave
verihed this onthetypeof Macandrevia, though the septum is never prominent. If thebeak
of the brachial valve of an old individual wereto be groundoff, aseptum would
beperceptible thereverymuch
as figured
by
Oehlerttor thebeak
of Endcsia cardium. ConsequentlyI feel obliged to regard Macandrevia, at least forthepresent,as form- ingmerely asubgeneric group under thegenus Eudema.
As
regards the partlyaustral speciesabouttobedescribed,since thereisnomeans
of decidingwhethertheirdevelopmentagrees with thoseformsreferable to Magcllaniinte or not,and as the adult shells exhibitno characters which could be regarded as diagnostic of a genus different from Eudesia, I feel obliged forthe presentto referthem
to that group. Itmay
be observedthat thereis nothingto prevent thefreemigration of northern forms intothe south Pacific along the coastof theAmericas.The
writer has already the evidence toshow
that several species, indeep water, do extend from Bering Sea south to the vicinity of the Grala])agos Islands and,in the case of one species, Solemyajohnsoniy Dall,
more
than a thousand miles farther south,with theknown
great range ofmany
brachiopods, there would be no apparent reasonwhy
species of the
Pauamic
region, forinstance, belongingto the northern type of development should not extend their range southward, ifopportunityarose. I regard it then, as quite likely, that the species I refer to
may
beMacandrevian
in their development as well asin their adultstate, though, forthemass
of characteristically austral species, the reverse might be thecase.SubgeuusMacaxdrevia,
King.
TypeTerebratidacranium, Miiller.
MACANDREVIA AMERICANA,
newspecies.PlateXXXII, iigs. 1,4,7.
Eudcsia foniaineana, Dall(notOiuugxy)Proc. U. 8. Nat. Mns.,xii,1889, p. 231.
Shell of moderatesize, rounded ovate,brownishwhite,smoothexter- nally except for
numerous
incrementalHuesbest visible under a lensj margins not flexuous; pedicle valve moderatelyarched, thin, witha
Proc. :N^.
M.
9J: 46722 DEEP WATER MOLLVSKS AND BRACniOrOVS—
BALL, yolivu.recurved short bealc, rather large foramen incomplete below to the extent of about one-fourth its circumference, with a small obsolete deltidial plate on each side; teeth strong, short, snpported each
by
a strong buttress witha recess behind it, and in old specimens with a smooth deposit ofcalluson the snrface of the valve between the two buttresses; nomedian
septum, the nniscnlarimpressionsfaint, situated in theupperfourth of the valve; brachial valveflatter, orbicular,Avith a small verylow cardinalprocess produceddownward,
three times itswidth, on the surfaceof thebeak between the crural plates, as a low ridge rounded above; crural plates strong, supported
by
buttresses forhalf their length, ratherclose together, supi^orting a thin delicate brachidium, longitudinally grooved nearand
at the recurvation, with a few spinuleson the outer edge, thehaemal border of the bightof the loop showing a small projection opposite the crural ])rocess of eachside, the brachidium reaching three-fourths of the distance froui the cardinal border to the basalmargin
and
over all one-third as wide as the valve; pallialsinusesfollowingmuch
thesame
courseasinM.
venasa but straighter, less branched,and
of a whitish instead of reddishbrown
color. Height of shell, 22; width, 11); diameter, 9.5mm.
Station2783, in 122fathoms
mud,
bottom temperature 480F., offthe westcoast of Patagonia, in latitude51°2' south;and
station 3360, in 1672 fathoms, sand, temperature 42oF.,in theGulf ofPanama.
Nos.87547
and
122859, U.S.N.M.When
firstexamined
one of these specimenswas
erroneously sup- posedto be theyoung
of Terehratulafontaincana, Orbiguy, a species which should doubtlessbereferred to MiKjeJlania venosa,Solander,as a synonym. Laterand more
careful study of a second specimen has enabledme
to correctmy
mistake.From
theyoung
ofM.
venosa the incompleteforamen enables itto be discriminated, without examining theinteriorof the shell. In a generalway
this specieslooksverymuch
likean adolescent &-pecimenofLaquensJeffrey sH, in generaloutline,but
is flatter.
MACANDREVIA
CRANIELLA, newspociea.PlateXXX,fig. 1.
Shell
much
resembling in sizeand
form the specimen figuredby
Davidson,* belowreferred to, but rathermore
rectangularwith a lessprominentbeak,
and
anarrowerandmore
slender brachidium. Surface of shellsmooth, exceptfornumerous
strong concentriclines ofgrowth and
prominent punctation.Under
alens a microscopicradial sculp- tureisvisible onpartsof theshell, resemblingthefibrous surfaceof aworn
lihynchonellaTntherthanregularstriation.Form
ofshellrounded, rectangular, with a low beakand
large foramen; pedicle valve rather inflated, thesideand
basalmargins slightlyexcavated, the basal cor- nersrounded but slightly prominent, from which the obscurerectan-•SeeMacandreviasp.,Davidsoit, Mou.Rec. Brach.,i, pi.xii, fig. 13,1889.
1894.
PROCEEDINGS OF THE NATIONAL MUSEUM. 723
gularityof the shell arises; teeth strong, foramen large, incomplete, thedeltidia almostlinear, butlong; dental buttresses strong,receding as they approach the archof the valve; brachial valve flatter; teeth strong, with very oblique buttresses; brachidium four-fifteenths the length of the valve,barelyone-third aswideasthevalve, slender, with a singlespineatthe point of recurvation,
and
a slightthickening, but noseptum, atthe cardinal apex; muscularimpressions smalland
con- fined totheupper fourth of the valve. Heightof shell, 17; width, 12;diameter,9
mm.
Station3302, in 1,175 fathoms,
mud;
temperature, 36.8^ F.; in the GulfofPanama.
No. 122858, TJ.S.N.M.This species resembles
MageUania
[Waldheimia) wyvillei, Davidson, but wantstheseptum and
cardinal process of the brachial valve of that species, which also has less oblique, shorter,and
wider deltidia. It hasmuch
resemblance also to Macandreviacranium, but isamore
rec- tangular,plump,and
compactshell,and
appears not to reach the size of ili. cranium. M. tenera hasa shorter loopand more
obviousseptum
in theapex of the brachial valve. It is also a smaller species
when
adult.
MACANDREVIA
DIAMANTINA, new species.PlateXXX,fig.5; platexxxii, figs.3, 6.
Shell smooth polished, dorso-ventrally compressed, flexuous, of a roundedlozengeor "
diamond
"-shaped outline; surface with faint in-cremental lines,but noradial sculpture,
waxen
white; pedicle valvesub- pentagonal,widestnear the middle, theconvergingsidesbelowproduced, the basal margin concave;beak
short, wide, the foramen incomplete below,with well-developeddeltidiaoneachside;teeth strong, supportedby
widebuttresses,forming wide recesseson each side; muscular im- pressions indistinct,situated intheupperthirdof theshell; nomedian septum
exists in this valve.The
genital glands are of a reddishbrown
color,shiningthroughtheshell&s
two
shortparallelstreakson eachside ofthe adductorimpressions; brachial valvewiderthan high, with the baseflexuousupward;
teethstrong,with veryoblique buttresseshidden under the dental platesand
forming small foveol*; cardinal process short, stout, prominent,but not produced into a septum; brachidium veryslender, extending tothe basal thirdof thevalve,the transverse limbatthe bightof the loopbeing wider thanany
partof the lateral limbs,the bight itself being, of course, narrower, as usual; there are twoor three spiuules at the recurvation; the genital glands in this valveshow
as asinglebrown
streak an each side of the attachment of the adductors. Itis, however, longer than the paired streaks of the pedicle valve. Heightof shell, 18; width, 17; diameter, 7mm.
Station 33G2, in 1,175 fathoms,
mud;
temperature,30.8^ F.; in the Gulf ofPanama.
No. 122860, U.S.N.M.This elegant little species recalls lAotliyrina wyvillei, Davidson, in itsform, though
more
lozenge-shapedand
less sharplyflexuous.724 DEEP WATER MOLLVSKS AND BRACHIOI'ODS—DALL.
vol.xvn.Itis sufficientlydistinctfrom all described recent species to