Consequently, the genus has often reached most of the high oceanic areas in the Pacific Ocean and elsewhere. 2 mm long, externally minutely scabby on the lobes, otherwise apparently glabrous, internally apparently glabrous, anthers and style ca. 6 mm long, elongated, externally hirtellous or glabrous except minutely hirtellous towards the apex, internally villosulous in the basal half, buds thickened adaxially into hooked projections, anthers 2.5-2.8 mm long, attached below the top of the tube, including tips, style 5- 7 mm long.
Shrub up to 3 m high, branches 7-8 mm in diameter, dense reddish brown, distally compressed to the menthos, soon glabrous, bark light brown, wrinkled. 1—1.5 X 1 cm (without corollas), subsessile, subcapitate cymose, subglobose or slightly oblong, short axes obscured by dense reddish-brown serrate-tomentose pubescence up to 3 mm, flowers ca. 20 mm long, sparsely forked within distally, lobes 4 (-5?), ovate-lanceolate, 4-5 X 2 mm, apices with thickened appendages; appendage, 1.5-2 mm long, attached to low tip of tube, style not visible.
Shrub or small tree up to 5 m high, twigs with a diameter of 4-5 mm, when young sometimes with matte, light brown tomentum, quickly bald, bark brown, smooth. 15-50 mm long, ovoid to ovoid-oblong, thinly matted tomentose and soon glabrous or glabrous and hairy only along the sutures, deciduous, sheath 10-40 mm long, lobes 2, elliptic to obovate, 5-10 mm long, deciduous, a few short brown hairs remain. Flowers monomorphic, with hy panthium 1-1.5 mm long, obconical, glaze. brittle; calyx limb 0.5-1.5 mm long, cupuliform to shallow cyathiform, glabrous externally and internally, truncated margin, glabrous or ciliolate, disc glabrous, 0.5 mm long; corolla hypocrateriform, 12-18 mm long, white, externally glabrous or minutely hairy at apex, tube 7-15 mm long, internally densely hairy in distal lobes 5, 4-5 mm long, elongated, externally glabrous except apically minutely hairy, internally densely hairy or only hairy towards the base, apices thickened adaxially into hook-like appendages, anthers 2.5–. 4 mm long, attached under the top of the tube, supplied, style approx. Fruits ripen orange or red, ellipsoidal, 8–. 10 X 6-7 mm, ribbed, spherical; pyrenes hemispherical in cross section, dorsal surface with 2-3 prominent ribs and thickened margins, ventral surface flat.
This species can be distinguished. by its large, elliptical, broad ellipse. tic, oblong or ovate-elliptic leaf blades beneath covered with a thin layer of pale brown arachnoid tomentum when young, but soon glossy, numerous secondaries. mm long, large inflorescences with 45-90. but that species has fewer secondary veins and smaller stipules and fruits.
- Fosberg (1939) recognized plants from Tahuata (762—884 m) as P. lebron
- Collections from Fatu Hiva (732—820
It can be distinguished from its Marquesan congeners by the following combination of characters: thick twigs with short nodes between them and visible leaf scars; leaves with relatively short leaves, usually winged,. The considerable morphological variation in this species may be a reflection of its wide geographic and ecological range. Because these distinctive vegetative and floral characters vary independently and to a continuous degree between extremes and do not appear to have any ecological or habitat correlates, we have not formally recognized any infraspecific variants here.
Fosberg (1939) has leaves with sturdy and broadly winged petioles 6-15 mm long, broad obovate leaves with abruptly short acuminate to blunt or rounded apices, compact inflorescences, flowers with stem cells 3-5 mm long, corollas 15-30 mm long with externally glabrous to minutely hairy lobes and anthers 3-3.5 mm long.
BISH, PTBG), 10190 (BISH, PAP, PTBG);
2-7 mm long, lobes 4, 3-6 mm long, narrow rows triangular-subulate or oblong, glabrous, deciduous leaving a fringe of reddish-brown hairs. It grows in wet forests with Weinmannia, Fagraea, Crossostylis, Hibiscus tiliaceus and Pandanus tectorius with Angiopteris eecta in the understory. Schäfer (Laboratoire de Botanique, Montpellier), who in 1975 made almost 700 botanical collections in the Marquesas.
Leaves with foliage 12-20 x 4.5-12 cm, obovate to broad obovate or elliptic-obovate, chartaceous, sparsely short-villous above, densely rufous villy below on veins, domatia absent, moderately villy elsewhere, edges flat , base cuneate, sometimes narrow, apex rounded and abruptly short-acuminate, acute 3-7 mm long, sometimes acute, secondary veins 15-20 pairs, eucampto dreamy, tertiary and higher order veins inconspicuous; petiole 1.5-3.5 cm long, densely red-vellified, barely or not winged; stipules calyptrate, ovoid, integument. 16-24 mm long, lobes 2.6-8 mm long, oblong, each sometimes about halfway bilobed, reddish brown, deciduous leaving a fringe of reddish brown hairs. 9-11 mm long, lobes 5, 6-8 mm long, elongated, externally hemispherical, internally shiny, apices adaxially thickened into hooked appendages, anthers 2.5-3 mm.
Known only from Nuku Hiva, where it occurs from 825 to 1175 m in the undergrowth of montane wet forest in and along valleys. There are correlated differences in pubescence (blade tufted or sometimes glabrous above in P. taupotiinii and consistently glabrous above in P. toviana), number of seconds. and 10-12 pairs in P. toviana), barbate domatia (absent in P. taupotinii and present in P. toviana), structure of inflorescence (once branched in P. taupotinii and twice in P. toviana), number of flowers per flower. However, there are no obvious differences either in the length of the anther or in the point of insertion of the anther, and the anthers are always included in the tube or only with it.
PAP, PTBG, US)
BISH-578766)
It is known only from Nuku Hiva where it is one of the most collected species. Psychotria toviana occurs in moist montane forest from 800 to 1170 m, usually in or along streams, together with Wein mannia, Metrosideros collina, Hernandia nukuhivensis F. QUESAS ISLANDS: Nuku Hiva: Spur of Mt. au-dessus du réservo3 ridges from near the complex l'Economie Rurale to the summit of Ooumu, Lorence et al. Tapueahu Gorge drainages northwest of Toovii above summit ridge, Perlman & Wood 15017. New Airport Road west of summit ridge. airport road, between airport road and Te kao, main ridge above Toovii, Perlman.
- Psychotria uapoensis Lorence & W
- Perlman 6472 (Holotype PTBG;
- mm long, obconic, glabrous, calyx limb
Small tree or shrub up to 5 m high, moderately branched, branches 3-4 mm in diameter, internodes compressed, bark brown, smooth. Blade leaves 5-11 X 3-7 cm, opposite, obovate-elliptic or occasionally elliptic, yellow to subcoriaceous, slightly shining, glabrous above or proximally glabrous, glabrous low, domatia absent, margins straight, base narrow to broadly cuneate, rarely slightly decurrent, apex obtuse to rounded and very shortly acuminate, acuminate length 2-4 mm, secondary veins 9-11 pairs, eukamptodromous, tertiary vein oblique, raised and prominulized on both surfaces, quaternary vein finely reticulate, raised on both sides; petiole 0.5-1 cm long, distally narrowly winged, adaxially brown scaly, abaxially glabrous; stipules ca liptrate, oblong, sheath 20-22 mm long, lobes 2, elliptic, 1-2 mm long, red-vilosulous outside, leaves leaving ru fos hairs in leaf axils. This new species is most similar to Psychotria hivaoana, from which it differs in its leaves with glabrous blades below and with fewer secondary veins (9-11 pairs), pedicels and bones villous above proximally, flowers with longer pedals, 1-7 mm long, and smaller fruits 6-7 X 5-6 mm.
ACKNOWLEDGMENTS
Systematics of the tropical shrub genus Psychotria (Rubiaceae): origin, species, and breeding systems in Hawai'i. Reorganization of the genus Psychotria and tribe Psychotrieae (Rubiaceae) inferred from ITS and rbcL sequence data. Carpology and pollen morphology of Psychotrieae (Rubiaceae-Rubioideae), towards a new tribal and generic delimitation.
Review of the neotropical genus Notopleura (Rubiaceae: Psychotrieae), with the description of some new species. Rubiacearum Americanarum Mag na Hama Pars VI: New species and morphological notes on Psychotria subg. Jean-Yves Meyer on Hiva Oa and Fatu Hiva came to our attention and is quoted below.
