During the first three months of 1932 the writer visited many of the Bahamas and the West Indies, and British and Dutch Guiana, while a member of the AlHson V. Diagnosis. - The shape of the gonopods and the first and second legs, as shown in Fig. 3.bto e, distinguish this species.
12 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 PROSTEMMIULUS WHEELERI (Silvestri)
Ventral striae do not extend beyond the borders of the legs, except on some anterior segments. The assignment of this species to the genus Orthoporus, on the basis of female characteristics alone, is somewhat tentative, and final judgment must be postponed until the male has been examined.
14 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 ANETHOPORUS CLARKI Charaberlin
Males in which the coxae of the third legs are shaped into triangular lobes; similar lobes, decreasing in size, are found on the next four legs. Segments 2 to 4 become smaller in width behind the first segment; the other segments are of equal width until near the posterior end of the body, which narrows rather gradually.
20 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 Males with first two pairs of legs conspicuously smaller than the ad-
The posterior segments with the transverse impression are very distinct in the back of all but six or eight of the anterior segments, where it is. Exposed part of the anterior half of the segments with surface similar to the posterior half, which is rather coarsely reticulate and with a few fine scratches.
26 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 Head without a median sulcus on the vertex ; clypeal setae four on
Succeeding segments each with a strong intervening transverse depression, before which the surface is sculptured with circular and crescentic impressions; posterior half of segment smooth, higher and much more strongly convex, large pore half-way to posterior border and opening from apex of broad but distinctly visible. The apical halves of the posterior lobes bent and carried outside the body, and their shape indicates that they are unable to be retracted inside the body at any time. First segment with a very pronounced, narrow lip, extending from the lower corner of the eye to the posterior lip; smooth surface.
The swellings in the pores are not very obvious from above, but when viewed from the side they are immediately visible.
28 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 Apex of the last segment rather broadly rounded and not separately
89ment20,uniform chestnut brown throughout, like the antennae and ment20,uniform chestnut brown throughout, like the antennae and legs are porifous segments with the entire keel and some of the adjacent area of the back white. The living color of the animals is chestnut brown with the keels and a triangular area on each yellow segment, with the base of the triangle extending along the posterior margin, contrary to the description of A. Diagnosis.—Although no males have been seen, the validity of this genus seems well supported by a combination of strong juvenile characters. the lateral and posterior edges of the segments near the middle of the body, and the different coloration of poriferous and non-poriferous segments.
Head wider than body, mandibular crest particularly prominent, long, subrectangular; antennae long and slender, nodes 2 to 6 inclusive of uniform thickness. 16,a), after which the tooth disappears (fig. 16, h); in the middle segments the tooth is larger and produced outwards and backwards; The posterior angles of the non-poriferous segments are all produced strongly caudad, especially in the mid-body region; in the segments of the porifera thick rims surrounding the pore occupy much of the lateral edge of the keel and replace the angle produced. The posterior margin of the segments with a large tooth are of the posterior angle of each keel and usually with its much smaller teeth; these teeth break off.
Anterior half of all segments brown below, the dorsum with a white medial area, wide to the anterior margin, which extends backwards and connects some of the last segments with the white portion of the posterior subsegment.
34 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 59 Since the female of this species closely follows the description and
Numbers of specimens have been found in the forest on the Bar de risle above Castries, St. In remarkable contrast to the diversity of the secondary characters, the gonopods show a degree of simplicity probably unequaled by any other member of the order. Dorsum and lateral edges of the keels with a few thick-eded setae, but without any polygonal areas ortubercles, the surface finely granular.
Pores in the usual order, opening on the dorsal surface from the base of the posterior corners produced. Males with a prominent transverse tubercle anteriorly, just below the antennae; also with a continuous ridge joining the lateral corners of the head; on each side the ridge begins at an angle and passes in front of the antenna and curves upward between the antenna and the frontal cusp, joining the ridge from the opposite side high on the apex where it is less prominent; labrum widened, the leading edge almost straight.
36 SMITHSONIAN MISCELLANEOUS COLLECTIONS
Three males, four females and four young were collected in the heavy forest near the top of the ridge at Kings Bay, Tobago, 20 February 1932. Head with nomadic furrow, the surface reticulate and with very small erect hairs, those of the vertex shorter than otherwise - true. Repugnatorial pores large, borderless, opening from the top of a light cone-shaped swelling of the dorsal surface near the posterior corner of segments 5,7,g and17.
When the apical joints are exercised, one crosses the other and they all face the opposite side of the body. Behind segment 4, these setas at the end of the posterior row are usually reduced in size or missing altogether, leaving only the four innermost setae. On the porous segments, a small seta rises from the top of the posterior projection of the lateral margin of each carina above and just behind the pore.
Body more convex than in other species and sculpting thicker; thesulci between areas of anterior margin of first.
46 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 pied by the four large areas, which are even more conspicuous than
Head with a deep median depression across the coarsely granulated area of the vertex, which is separated from the front of the head by a deep canal just above the antennae; the clypeal area is raised above and separated from the front by a deep furrow, especially evident on the sides. Segments with raised leading edge strongly evident on carinae and adjacent dorsum, but broken into erect crenations across middle of dorsum. First joint of the fourth leg with a rounded stiffened tuberclein front, the second joint with the distal anterior corner continued into an acute lobe.
The following segments without longitudinal or transverse lines dividing the surface into large polygonal areas, and without any definite series of large tubercles; instead, the dorsal surface, except for the usual smooth sulcate areas of the posterior margin and lateral carinae, is densely scattered with small tubercles of various sizes, the median ones somewhat rounded, but those on the sides of the dorsal and adjacent carinae are decidedly cuspidate, and each is surmounted of very dirty. . 14 MILLIPEDES OF THE WEST INDIES AND GUIANA LOOMIS 49 Anal valves flattened, with moderately raised margins; disc of each valve with an angular fold or ridge extending upwards from the lateral corner of the scale. Females with the anterior ventral margin of the third segment gradually raised from each side into a form of a low ridge or ridge, i.
The posterior subsegments are raised abruptly high above the anterior subsegments, the front of the elevation being distinctly depressed and receiving the posterior edge of the preceding segment when the body is held straight; anterior margin of the segments with a definite scallop between the outer and inner row of tubercles and tubercles on each side. two other scallops between the interior of the tubercles.
50 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 portion sharply raised above the broadly expanded anterior margin,
14 MILEPADES OF THE WEST INDIES AND GUIANA LOOMIS 5 Females with a very short ventral ridge on the third segment just posterior to this second pair of legs. Two males, one of the same type, and two females were collected under a mango tree on the lower slopes of Morne Brigand, near Bayeux, Haiti, July 16, 1927, by H. In living specimens, the dorsal bone is dark coal black and usually somewhat covered with dirt; the underside of the posterior subsegments is also dark. the sterna, and the raised part of the head between and above the antennae, is black as coal; rest of head, antennae, legs, sternum, anal valves, preanal scale, papilloform apex of posterior segment, and entire anterior subsegments uniformly white or colorless.
First segment with anterior median border almost straight, sides extending obliquely outwards and backwards; lateral parts of posterior border directed inward and backward to straight, median transverse border. Central area of segment subglobular, very abruptly raised above anterior margin greatly expanded, with anterior row of four large conical tubercles and a posterior row of six smaller tubercles, each tubercle surmounted by a long seta, except for many small, raised ones scattered over the entire surface of the segment. A small group on the middle border of each of the areas of the expanded anterior border.
On subsequent segments, the dorsal surface, including the large tubercles, is smooth with each of the large tubercles supporting long bristles.
52 SMITHSONIAN MISCELLANEOUS COLLECTIONS
The first segment is somewhat flattened, much less convex than the next segment, and with the margin and lateral angles noticeably higher than the keels of this segment. Viewed from the side, the keels of the segments do not extend below the ventral third of the body. Second segment with lateral margin of each keel but little longer than keel margins of next two segments, marginal lobe not noticeably larger than these segments.
Non-poriferous segments anterior to segment 15 with three lateral lobes each sheath, segments 16-19, with 4-lobed calyces; Segment 5 with one lobe in front of pore-process, all other pore-bearing segments with two lobes in front of process, which projects from posterior corner of column (fig. 27, b). Segments 2 to 18 bilobed at posterior margin of outer margin of large dorsal tubercles, outer lobe largest and slightly elevated. From the description and drawings of P.sequcns (Chamberlin). it is evidently closely related to P. crcsccntis but is larger in size; the basal nodes of the gonopods of segmented specimens 19 and 20 of P. crescentis are wider than long, but in the P. sequences the basal nodes are distinctly longer than wide; apical node of gonopods absent in immature Florida specimens and broken in mature.
Specimens of P. f/raiiulofro)is (Chamberlin) have been found at several localities in Haiti, and a description of them has been prepared which allows comparison with P.
56 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 Numerous specimens, all of which are females with not more than
The gonopods are somewhat like those of Cynedesmus simplex Loomis, but the interior of each is shorter, heavier, and simpler. His assumption that Lophodesrnus Pocockisasynonimofthisgenus does not seem justified after a careful examination of the characters of the two genera. The statement that no type description of Cynedcsmus has been published is incorrect, as C.
Surface of segments with four longitudinal rows of four high, rounded tubercles; in the caudal segments, the lateral rows of tubercles decrease and the median rows increase; surface elsewhere obscurely granular, granular. Second segment with lateral margin of each keel 3-lobed and again over half length of margins of segment 3 or 4. Posterior angle of lateral keel each side reaching about apex of median ridge produced.
Second segment with four narrow longitudinal ridges, apex of each divided into three parts; all edges of the segment.
64 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 Last segment fully exposed from above ; the dorsum with two
More than a score of female specimens were found on the underside of a log near Paramaribo, Dutch Guiana, March 1, 1932. Subsequent segments without any distinctly projecting lateral carinae, indicated by rather prominent shoulders on the anterior segments extending to the back of the body becomes unclear. Live animals dull black; the anterior subsegments black posteriorly, white anteriorly, the dividing line very distinct, biform.
14 THOUSANDS OF THE WEST INDIA AND GUIANA LOOMIS 67 First segmenthemispherical or somewhat crescentic; with the anterior margin transverse and strongly marginated, the middle third low and simple, the outer third occupied on each side by three raised lobes, the outer being twice as broad as the two undersized inner ones together; disk broadly depressed in center, finely erosic-granular and hispid, each side slightly inflated and coarsely granular. The succeeding segments are coarse-grained and very dull, with short and long bristles mixed together, the entire surface usually having a moderate deposit of dirt which obscures the pores. The outer margin of the keels, which follows the second segment, is simple and sharply rounded, becoming more broadly rounded on the middle segments, where it is distinctly bilobed, and on the posterior segments it finally becomes distinctly trilobed.
On the anterior subsegments the dark posterior portion is uniformly punctate, the surface, including the pits, very minutely granular; white anterior part of subsegment minutely honeycombed.
68 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 Last segment broadly rounded behind, the median third of the
EXPLANATION OF PLATES
