46 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 89 pied by the four large areas, which are even more conspicuous than

on the preceding segments. Anterior rim of the subsegments con- spicuous onthe carinaebutlessevident across thedorsum. Posterior angles of the carinae not definitely produced

backward

until seg-

ment

16 or 17.

Last segment with two prominent dorsaltubercles.

Fig.22.

—

Docodesmussculpfuratus. Gonopod.

Gonopods

as

shown

infigure22,the basaljoint large,hemispherical.

Legs of the males without special modifications except that the sternum betweenthefourth legsbears

two

tinyroundedtubercles.

Females with the anterior ventral margin of segment 3 elevated for a long distance into a very prominent ridge directed obliquely backward, the median portion thin-edged, the lateral angle oneach side higher and several times as thick.

Type.—

U.S.N.

M.

no. 1102.

DOCODESMUS TRINIDADENSIS

Chambeilin Plate4, fig. 2

DocodesmustrinidadensisChamberlin, Bull. Mus. Comp.Zool.,vol. 62, p.219, 1918.

Specimens were collected in Trinidad at Sangre Grande, in the Aripo Valley, and in the

Arena

Forest area; others were foundat Scarborough and

Man-of-War

Bay,Tobago.

The

largestspecimen

was

12

mm

longand2.2

mm

wide.

The

living color ranged

from

blackwiththe carinaedeep reddish-brown to en- tirelyblackwiththeexception of a

narrow

reddish area along the front margin ofsegmenti.

Body

moderatelyconvex^;rather coarsely sculptured butrelatively lessprominent than D.sculpturatus;thetransverseconvexpolygonal areas of the

dorsum

are

more

pronounced and there are

more

fine, irregular granulations in addition to the large tubercles.

The

areas separatedbysulcialong the posteriormarginof thesegments project

NO. 14 MILLIPEDS OF

WEST

INDIES

AND GUIANA

LOOMIS 47 as faint crenulationsbut areproduced ^intoacute projections only on segment 19.

Head

with adeep medianimpression across the coarsely granular area of the vertexwhich isseparated

from

the front of thehead by adeepchanneljustabovetheantennae;clypeal region elevatedabove andseparated

from

the frontby a deep furrow, especially evidenton thesides.

Segmentswith raised anterior rim strongly evidentonthe carinae andadjacent

dorsum

butbrokeninto erectcrenations across themid- dle of the dorsum.

Fig. 23. Docodcsmustrinidadcnsis. Gonopod.

Gonopods

as

shown

in figure 23, the basal joint relatively small, subangularly compressed,not forming an almost truehemisphere.

Males withthesternum betweenthe fourthlegsbearing

two

small, rounded tubercles. First joint of the fourth legs with a rounded setiferous tuberclein front,the second joint withthe distal anterior cornercontinued intoan acute lobe.

Females with theanterior ventralmargin of segment3 carried

up

into a high, thin, slightly reflexed ridge, highest at the middle and without prominentlateralangles.

Inspiteofthefactthat

my

specimensare smaller,itseemsevident thatthis is Chamberlin's species, although

few

distinctivecharacters weregivenin its description.

DOCODESMUS ROBUSTUS,

n.sp.

Plate4,fig.3

A

male (type)

and

a female were found at Kings Bay,

Tobago

Island, February20, 1932.

Diagnosis.

—

^{Besides being}

much

shorterin proportion toitswidth than

any

other species of

Docodesmus,

theirregular, cuspidate tuber- culation of the

dorsum

andthe lackof seriesof large tubercles

make

thisthemosteasilyrecognized

member

of the genus. ^It

may

bere- motely relatedto D. trinidadcnsis,as the structure of thehead and thegonopods

seem

toindicate.

4

48

SMITHSONIAN MISCELLANEOUS

COLLECTIONS VOL. 89 Description.

— Body

elongate-ovaland lessthan four times as long asbroad;bothspecimensmeasuring 13

mm

inlengthand 3.7

mm

in width.

Dorsum

stronglyconvex,

much more

sothanin trinidadensis, andthelateralcarinae descend

more

obliquely.

The

living animals were dull black above throughout, lighter be- neath.

Head

with a very pronounced, broad, deep furrow just above the antennaeseparating the frontal region

from

theraised, medianly furrowed,granular vertex;clypealregionsuddenlyelevatedabovethe frontalregion,smooth andshining.

First segment abouta third narrower thanthe

midbody

segments

;

anterior margin evenly rounded, the surface behind the marginal areas with densely scattered small granules.

Fig.24.—Docodesmnsrobustns. a,last2segments, anal valvesandscale,ventral view; b,side viewof agonopod from above and slightly behind.

Ensuing segments without longitudinal or transverse lines divid- ing the surfaceinto large polygonal areas, and without any definite series of large tubercles^; instead,the dorsal surface, except the usual smoothsulcate areas of the posteriormargin andthe lateralcarinae, isdensely scattered with small tubercles of varioussizes, themedian ones

somewhat

rounded but those on the sides of the

dorsum

and adjacent carinae are definitely cuspidate and each is surmounted by averytinyand veryshortseta. Posteriormarginalareasatthemiddle of the

dorsum

veryshort,not projectingbeyondthemarginas crena- tionsexceptonseveral ofthelastsegments. Anteriorraisedrimcon- tinuous across thedorsum,

where

itsapexisslightlyundulated.

Last segment with

two

largeappressed, triangular tubercles above.

Lateral lobeslarge,themedian onebroad,rounded-transversebehind.

Ventrally, the fourapicalsetaeareprotectedon eachside

by

a strong conic tuberclewhichprojects

backward

(fig.24,a).

NO. 14 MILLIPEDS OF

WEST

INDIES

AND GUIANA

LOOMIS 49 Analvalvesflattened,withmoderatelyraisedmargins;disk ofeach valvewithanangular fold or ridgeextending

upward from

thelateral corner of the scale. Preanalscale triangular, the setiferous tubercles exceedingthe

median

angle.

Gonopods

shaped

somewhat

likethose of D.trinidadensis, but the medianportion of thelong inner

arm

isexpandedjustbelowthedistal half (fig. 24, b).

Males withthe fourth sternumbearing

two

rounded,finely hispid tubercles; first joint of each fourth legwith a tubercle ofabout the

same

size.

Females with the anterior ventral margin of the third segment gradually raised

from

each side to

form

alow ridge or rim, whichis

highestatthemiddle ofthe body.

Type.—U.^.^M.

no. 1103.

lOMOIDES,n.gen.

Type.

—

lomoideshispidiis, n. sp.

Diagnosis.

—

This genus seems tobecloselyrelatedtoloinusCook, butis

more

convex and

more

compact;the lateral carinae arenot as broadorasdeeplyincised,although withthe

same number

ofmarginal lobes.

The

rowsof large dorsal tuberclesconvergecaudadinsteadof diverging or runningparallel,andthe thirdjointofthe legsislonger than the last joint.

Description.

— Body

very compact, about fourtimes as long as wide, the

dorsum

very strongly arched; lateral carinae broad, exceeding thelegs,the anterior cornerslowerthanthebottomofthebodycavity.

Surfaceof thecarinaeandthe

dorsum

roughened,including the four longitudinal rowsof largetubercles. Segments2 to 5inclusivegradu- allywidening, theensuing segmentstothesixteenth ofuniformwidth, afterwhichthey

narrow

rapidly.

The

posterior subsegments are ab- ruptly raised high above the anterior subsegments, the face of the elevation being distinctly recessed, receiving the posterior edge of the foregoingsegment

when

thebodyisheldstraight;anteriormargin of the segments with a definite scallopbetween the outer

and

inner

row

of tubercles on each side, and

two

other scallops between the innerrows of tubercles.

Head

quite convex, completely hidden beneath the first segment surface between and above the antennae elevated and roughened, belowtheantennaethe surfaceis

somewhat

shiningandhispid.

An-

tennae strongly clavate, geniculate, joint 5 broadest andlongest.

First segment subhexagonal, greatly exceedingthe head in front, narrower than the second segment, twice as broad as long; central

50

SMITHSONIAN MISCELLANEOUS

COLLECTIONS VOL. 89