Museum collections on various occasions, such as Alberto Tremoleras, from the same city. Haevis^'' from the vicinity of the city of Mendoza, in the province of the same name.
THE SPECIES OF AEGLA — SCHMTTT 441
The presence of two or more species in one place, as in and around Buenos Aires, and perhaps also in Montevideo, may be the result of massive floods that affected at least the lower reaches of the several rivers that flow into the Rio de la Plata. object. In general, most species appear to be quite restricted in their distribution (but it should be remembered that the number of records we have for any one species is still very small).
THE SPECIES OF AEGLA — SCHMTTT 443
444 CONTINUATION OF THE NATIONAL MUSEUM vol.91 Ortmann (1902,p.341) in his discussion of the geographical distribution of freshwater crustaceans and their bearing on ancient geography. Since the forms with Andean-type rostrum are found in each of the latter "environmental complexes" of Haseman, they must have something in common, be it geological history, environment or something else.
THE SPECIES OF AEGLA — SCHMTTT 445
CHARACTERS USED IN DIAGNOSTIC KEY AND SPECIFIC DESCRIPTIONS (Fig. 41)
The chelae in the female are undersized and underdeveloped, and more closely resemble the pattern of the small chela of the male, which in turn might be described as more or less. In three species the grasping edge of the fixed finger is not such a lobular tooth: A.
THE SPECIES OF AEGLA — SCHMTTT 447
The anterior and posterior lines mark the muscle attachments; the middle one constitutes "'a plane of fracture' in which the division of the limbs takes place in [this and] many [other crustaceans] Reptantia" (Caiman, 1909, p. 273). In describing the armor of the "inner margin of the ventral surface of the ischium."
THE SPECIES OF AEGLA — SCHMITT 449
450 PROCEE'DINIGS OF THE NATIONAL MUSEUM vol.91 Width of orbital and extraorbital sinuses, where mentioned,. Most species of Aeglae show a tendency towards softness and softness, even to suppression of spines in older and more developed specimens.
THE SPECIES OF AEGLA — SCHMITT 451
KEY TO THE SPECIES OF AEGLA
Inconspicuously strong spine near distal end of ventral posterior margin of ambulatory merus, at most a relatively small spine, tubercle, or scale in this position; epimeron of second (internal view, apparently . first) abdominal somite with anteroventral border more or less straight, at most only slightly concave; ventral angle rounded;. Usually only the first hepatic lobe is well defined and pointed anteriorly, the second and third lobes scarcely more than indicated (occasionally one of the other lobes is fairly well marked on one side or the other of the carapace); mobile finger definitely with lobes on outer edge near base; in most specimens, especially those of medium and small size, the lobe is furnished with a small spine or sharp scale, in many larger specimens like the type the lobe is often unarmed, but always distinctly present and more or less angled; seta of chelipeds with not particularly prominent tubercle (not spine) provided with pink tip or scale at distal end of inner margin of ventral surface.
THE SPECIES OF AEGLA — SCHMITT 453
B\ Carapace conspicuously inclined or carinated throughout its length; rostral carina anteriorly disappears in distal third of free part of rostrum and merges with thickened distal end; lateral margin of posterior part of carapace (behind cervical groove) conspicuously denticulate, sharply incised and armed with prominent serrated or flattened triangular dorsal teeth; orbital spine of good size; extraorbital sinus well formed, anterior part distinct, though moderately narrow, perhaps not more than one-fourth width of orbital sinus; anterolateral spines reaching one-third to one-fourth length of cornea; palmar ridge thick, conspicuously spiny; movable property. C\ The anterior third, or even nearly half in some cases, of the upper surface of the free part of the rostrum is gently convex or concave from side to side with usually only a trace of the forward-extended rostral carina or scales with which its carina is furnished; distal part of rostrum typically and usually strongly and more or less abruptly curved; rostral outline moderately broad triangular, carina short but well defined, furnished with one row of irregularly alternating pink scales; the orbital spine may or may not be developed;.
THE SPECIES OF AEGLA — SCHMTTT 455
Rostrum more or less hoofed (tends to be tongue-shaped rather than sharply triangular), lateral margins often more or less subparallel in the middle part of the free part of the rostrum; rostrum in lateral view noticeably bent downwards, distally curved;. Orbital spine usually absent, usually outer end of orbital margin merging with inner slope or edge of anterolateral spine with little or no demarcation; sometimes a slight sinuosity or a more or less insignificant oblique deviation develops.
THE SPECIES OF AEGLA — SCHMITT 457
Rostrum long, slender spine-like, sharply carinate, ridged, triangular in cross-section, exceeding eye stalk by two to three times the length of cornea; crestofrostralcarina furnished with a close double row of good-sized corneous scales about to level of corneae, anterior to which the row becomes single with scale soft, close, sometimes separated from another; posteriorly, the carina ends just before the anterior margins of protogastric lobes, larger scales of carina often stop at level of epigastric prominences. Epigastric prominences well marked, although low tubercular, furnished with one or more, usually several, corneous scales, individually about the size of the scales on the rostrum; anterior margin of protogastric lobes, though only slightly elevated, distinctly marked, at least in part, by a short row of substantial corneous scales, the apical one of which is larger and heavier than the others.
THE SPECIES OF AEGLA — SCHMITT 459
Dorsal longitudinal margin of merus of kelipata armed with row of large, sharp, well-developed spines; at middle of anterior margin of merus a strong spine about as large as anterior spine of dorsal longitudinal margin. The inner margin of the ventral surface of the seat of the cheliped is armed with at least two fairly long, stout, more or less equal pink-tipped spines; among Aeglas, two sessile spines of this size and prominence are found only in this species and A.
THE SPECIES OF AEGLA — SCHMITT 461 The chelipeds and chela for the greater part have the same general
462 PROCEEDINGS OF THE NATIONAL MUSEUM
THE SPECIES OF AEGLA — SCHMITT 463 Larger hand very smooth appearing, but under glass finely granu-
Anterior margins of protogastric lobe broadly incurved, not at all tuberculate at apex of angle. 34;This figure of Rudolfo von Liiering is original and undoubtedly based on one of the many specimens collected by his father Hermannvon Ihering, in the state of Rio Grande do Sul (collector no. 619).
THE SPECIES OF AEGLA — SCHMITT 465
466 PROCEEDINGS OF THE NATIONAOL MUSEUM
Anterolateral angles of all three hepatic lobes well marked, at least the first (in the type all three) sharply acute and canted; first spine long and slender and considerably exserted; the second about half the length of the. THE SPECIES OF AEGLA — SCHMITT 469 to the inside^^ for inside and often a little before the anterionnost.
THE SPECIES OF AEGLA — SCHMITT 469 inward ^^ for inside and often a little in advance of the anterionnost
Rostrum sharp, spine-like ridge, exceeding eyes by at least twice corneal length; therostral carina is furnished with several longitudinal rows of irregularly placed tiny pink scales; Karina. All three hepatic lobes are well marked and pinkish-tipped, and each well spaced from the others, so that the lateral margin of the anterior part of the carapace tapers stepwise from the jugular groove to the anterolateral spine.
THE SPECIES OF AEGLA — SCHMITT 471
472 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 91 . smaller spikes) lined up on the inner edge of the palm slightly. THE SPECIES OF AEGLA — SCHMITT 473 The stepwise arrangement of the well-marked lobes of the liver and.
THE SPECIES OF AEGLA — SCHMITT 473 The stepwise arrangement of the well-marked hepatic lobes and
Border above the inner spiny margin of the carpus, armed with conical tubercles, the greater part of which are more acute conical spines; the anterior internal lobe or angle of the carpus is broadly conical and tipped with a small sharp horny spine; upper margin of merus armed with slender spines, the most anterior and sometimes the largest located directly on the anterior margin of the merus, the next spine may be slightly larger or slightly smaller than the anterior spine. The inner edge of the ventral surface of the ischium also has a strong conical spine at the anterior end, and generally in addition a smaller one of variable size and sharpness at the posterior end, and one or two smaller ones in between.
THE SPECIES OF AEGLA — SCHMITT 475
476 PROCEEDIN'GS OF THE NATIONAL MUSEUM vol.9i In turnover, these specimens straightened very handily, an ability not so evident in the larger parana specimens collected at Rio Negro, Parana, Brazil. In getting back the considerable number of specimens I had brought with me, I was most helpfully assisted by Harry Preston Midkiffs, of the Institut Christao, by Camille Cunha and several of his nephews, and by Werner Nickol, Conrado Pusch, Amacleto Baptista, and one of their friends who took me on a full day drive to Hacienda Marumby where we got additional material.
THE SPECIES OF AEGLA — SCHMITT 477 Palmar crest low, narrow; obscurely and irregularly serrate,
Rostrum rather broadly triangular, scarcely exceeding eyes in length of cornea; the median carina is sharply ridged to its anterior extremity, making the rostrum undoubtedly covered with a ridge, especially in the anterior half or third of its free part, although the lateral slopes of the dorsal surface of the rostrum towards the base of the rostrum are somewhat concave; rostral carina throughout or greater part of its extent with one row of pink scales of good size, at least on that part of rostrum lying anterior to posterior margins of orbits. The epigastric ridges are also low, barely better developed than the anterior margin of the protogastric lobes.
THE SPECIES OF AEGLA — SCHMITT 479 Large hand short, stout, inflated, and smooth appearing; short
Holotype.- The largest of three male specimens, measuring approx. 29 mm, in length of shield and rostrum together, collected by An-. Rostrum moderately broad-triangular, scarcely or at all exceeding eyestalks by as much as length of cornea; one-.
THE SPECIES OF AEGLA — SCHMITT 481
Cheliped ridgeofcarpus above inner margin, armed with four to five sharp spines, occasionally with several much smaller spines in between, sometimes, as in one of the females, these spines. Dorsal border of merus of celipeds armed anteriorly with large, stout, pointed spine, followed by perhaps two or three much smaller ones; a spine similar to the large spine at the dorsal end of the merus, but of even larger size, flanks the anterior end of the joint; often this spine has a slightly pointed ridge or spine on the inner or outer side of its base.
THE SPECIES OF AEGLA — SCHMITT 483
Each of the three specimens belonging to the Philadelphia Academy, referred to in the notes to the 'Distribution' of this species below, shows a distinct but small orbital spine on each side, separated from the corresponding anterolateral spine by a narrow notch . . Anterolateral angle of the carapace forming a sharply acute, rather slender conical spine, extending to and slightly beyond the posterior margin of the cornea, in some cases approximately to the center of the cornea.
THE SPECIES OF AEGLA — SCHMITT 485
Although the anterior dorsal angle of the epimeron of the second abdominal somite may rarely, and I think only accidentally, bear a small corneal scale or two, or even a small spinule, it appears that A. The several suture lines which meet form the anterolateral angles of the cardiac region of the carapace together form a short, fairly longitudinally oriented bar (Fig. 54).
THE SPECIES OF AEGLA — SCHMITT 487
If one considers the largest spine of those arming the inner margin of the carpal as the most anterior of that particular series, in this species we find on the innermost anterior slope of the base of that first spine a smaller, but still conspicuous, strong spine . in roughly a triangular area bounded by the first spine, the carpal border and the anterior inner lobe of the carpal. This "inserted" spine may sometimes be closer to the large first spine of the series, but not normally fused to the inner edge of the carpal, than to the carpal lobe or carpal border.
THE SPECIES OF AEGLA — SCHMITT 489
The first hepatic lobe started from the anterolateral lobe with a fairly wide, relatively good level; anterolateral angle of first hepatic lobes slightly produced and subacute, bearing a small corneous grain or denticle, lateral margin of lobe scabrous; The second and third hepatic lobes, although not much more, are slightly better. Inner margin of palm with a well-developed, impressed or excavated ridge, having its outer margin somewhat parallel to dorsal margin of palm, not nearly so subdisiform as in A.
THE SPECIES OF AEGLA — SCHMITT 491 were formed by the merging of two spines of normal size to form one
Compared to the eye, the rostrum of the actual species appears slightly longer; also, it appears to be relatively slightly more recurved distally; the rostrum is more nearly equal in the subspecies. In the subspecies, they are relatively heavier, stockier (thicker, more raised or inflated), with considerably shorter, wider (stubber) fixed fingers; the outer margin of.
THE SPECIES OF AEGLA — SCHMITT 495
496 PROCEEDINGS OF THE NATIONAL MUSEI3M
TYPES OF AEGLA - SCHMITT 497 that the surface scratches of the finger differ in it.
THE SPECIES OF AEGLA — SCHMITT 497 than the scabrosities of the surface of the finger otherwise in the
The podium looks quite thick, is more or less broadly flattened - triangular, noticeably pressed forward, curved down, so much. Orbital sinus quite narrow, more or less V-shaped; orbital spine small, well placed on the inner slope or margin of the anterolateral spine and separated from it by only a small notch.
THE SPECIES OF AEGLA — SCHMITT 499
Geographically in the province of Jujuy, these species are found barely more than 70 miles apart, but environmentally, or at least climatologically, they are far removed from each other. One of these specimens was selected as the type of A.affinis (M. A. C.N. tag No.9817), the los eliped (M. A.C. N. tag No. 4186) represents the same species; the remaining specimen proved to be amale Aeglaabtao (28.0 mm. in length of carapace and rostrum).
THE SPECIES OF AEGLA — SCHMITT 501
Anterolateral spine of good size, with anterior extremity reaching near or approximately level with the center of the cornea; the dorsal surface of the anterolateral lobes is strongly flattened, almost or slightly excavated, giving the impression that the anterolateral spines are oblique.
THE SPECIES OF AEGLA — SCHMITT 503
Furthermore, the several suture lines that meet to form the anterolateral angles of the cardiac region of the carapace run together to form a short transverse or obliquely transverse bar inA. Kostrum more or less ungulate (more tongue-shaped than sharply triangular), lateral edges more or less subparallel in middle of free part, exceeds eyes by I14 times to nearly twice.
THE SPECIES OF AEGLA — SCHMITT 505
Extraorbital sinus very small, sometimes obsolete and represented by no more than a marked, usually abrupt, often almost rectangular offset between the tip of the orbital margin and the inner slope or margin of the anterolateral spine; an orbital spine, or more accurately spinule, commonly present, usually much smaller in size. Upper longitudinal margin of merus bears series of apically scabby, raised tuberculiform elevations, the anterior being the largest; middorsal point of anterior margin of merus without red or swelling and otherwise unarmed or onorna-.
THE SPECIES OF AEGLA — SCHMITT 507
Lieutenant Gilliss, of the United States Naval Astronomical Expedition of 1849-52, and determined by William Stimpson; three small males (15 to 21 mm. long) and one female (17.5 mm.) from near Melipilla, Province of Sanitago, Chile, which were collected by Dr. In the smaller specimen collected for me a small extraorbital sinus and a tiny orbital spinule are present on the right side; on the left, the anomaly is usually found on the inside of the anterolateral spine in the absence of an orbital spine or spinule.
THE SPECIES OF AEGLA — SCHMITT 509
Rostral carina ridge almost fades near distal end of rostrum, which is pubescent similar to A. Epigastric ridges low and blunt; anterior margins of protogastric lobes not particularly distinct from rest of carapace, but well so.
THE SPECIES OF AEGLA — SCIBMITT 511
However, the conical tubercles on the dorsal longitudinal margin of the merus of the cheliped resemble single or spinule points; the inner ventral margin of the ischium is as in the type on the right cheliped; on the left there are two small elevations of which the anterior is the larger and the small spinule point between the anterior and posterior spine. The anterior dorsal corner of the epimeron of the second abdominal somite is armed with two small spines or spinules on the left, with one only on the right.
THE SPECIES OF AEGLA — SCHMITT 513 AEGLA RIOLIMAYANA, new species
The inner anterior lobe of the carpus of the chelipeds appears to be armed or otherwise revolute in both. The more or less straight back of the lateral furrows or areola furrows are subparallel in A.
LITERATURE CITED Bennati-Mouchet, Simone
THE SPECIES OF AEGLA — SCHMITT 517
THE SPECIES OF AEGLA — SCHIVIITT 519
INDEX TO DESCRIPTIONS OF AND DIAGNOSTIC NOTES ON THE SPECIES AND SUSPECIES OF AEGLA
NOTE
