THE RECENT AND FOSSIL MOLLUSKS OF THE GENUS CERITHIOPSIS FROM THE WEST COAST OF AMERICA.
By Paul Bartsch,
Assistant Curator,DivisionofMollusks, U.S. NationalMuseum.
The
first Cerithiopsisknown from
the west coast ofAmerica was
reportedby
AlcideD'Orbigny
in 1840from
Peru.1 Thiswas
fol-lowed twelve years later
by two
additional species, discoveredby
Prof. C. B.Adams
atPanama and
described as Cerithium neglectum C. B.Adams and
Triforisinfrequens C. B.Adams.
2In 1857 Dr. P. P. Carpenter published that part of his Catalogue of
Mazatlan
Shellswhich
deals withthemembers
of thisgenus, citing the following species:tuberculidesCarpenter. sorexCarpenter.
tuberculoid.es albonodosa Carpenter. convexa Carpenter.
cereaCarpenter. decussata Carpenter.
pupiformisCarpenter. assimilata C. B.
Adams.
Of
these, C.convexaisnow
placedinthegenus Metaxia. C.decussatais a Bittium,
and
Cerithiopsis assimilata Carpenter=
Cerithium assimilatumC. B.Adams must
be referred to the genusSeila.In 1865 Doctor Carpenter described Cerithiopsis intercalaria3
and
atthe
same
timereferred Cerithium bimarginatumC.B.Adams
to this genus.At
present both of these species are placed in the genus Eumeta. In theSupplementary Report on
the Present StateofOur Knowledge
withRegard
totheMolluscaoftheWest
CoastofAmerica
4 Doctor Carpenter published a tersediagnosisofthe followingspecies:Cerithiopsis columna. Cerithiopsis purpurea.
Cerithiopsis munita. Cerithiopsisfortior.
All of these were later
more
fully described. Three of them, C. munita, C. purpurea,and
C. fortior, arenow
placed inthe genus Bittium. In 1867De
Folinadded
another species,5 Cerithium destrugesi,which may
not belongto ourfauna, asthe localityiscited asPanama
or Negritos Island.iVoy.Amer.Merid.,p. 443, pi. 77,figs.9-10.
*Ann.Lye. Nat.Hist,ofN.Y.,1852,pp.379-80.
aProc. Zool.Soc,1865, p. 281.
*Rep.Brit.Ass.Adv.Sci.for 1863,publishedin 18C4.
«LesMeleagrinicoles,p. 71, pi.6, fig. 12.
Proceedings U.S.National Museum,Vol.40—No. 1823.
327
328 PROCEEDINGS OF THE NATIONAL MUSEUM.
vol.40.After this
no
additions weremade
for seventeen years,when
Dr.W. H.
Dall described Cerithiopsisstejnegeri,from
Alaska.1Two
yearslaterthe
same
authoradded
another species,2alsofrom
Alaska,under
thename
Cerithiopsis stejnegeri truncata.After another lapse of seventeen years
we
find another speciesadded
to our list, this time a fossil described as Bittiumwilliamsoniby
Dr.Ralph
Arnold,3from
the PleistocenebedsofSan Pedro and San
Diego, California. Five }Tears later the present writer described:4Bittium
tumidum.
Cerithiopsis cosmia.Bittium quadrifilatum ingens. Cerithiopsis pedroana.
the first
two
ofwhich must
be referred to the present genus. Since thenDoctor
Dall hasadded
Cerithiopsis excelsus,5and
the present writer described Cerithiopsis stephensse.6The
nuclear characters ofthe species studied are quite interestingand
can be used toadvantage
in subdividing the genus intominor
groups. Unfortunately,twenty
oftheforty-fiveformsunder
consid- erationhave
lost these early turns,which
leaves a rather large per- centageinan
uncertainposition.Of
the twenty-five speciesinwhich
the nuclearwhorls areknown,
eighteenbelongto Cerithiopsisproper,havingsmooth
nuclear whorls.These
are:fatua Bartsch. dbreojosensis Bartsch.
oxys Bartsch. berryi Bartsch.
cerea Carpenter. galapagensisBartsch.
sorex Carpenter. cesta Bartsch.
carpenteriBartsch. stejnegeriDall.
pedroana Bartsch. stejnegeridinaBartsch.
tuberculides Carpenter. neglecta C. B.
Adams.
tuberculides albonodosa Carpenter. halia Bartsch.
pupiformis Carpenter. aureaBartsch.
Two
specieshave
the early portion of the nuclear whorlssmooth and
the later part axially ribbed; for these Iwould
suggest the sub- genericname
CeritMopsina.The two
species are Cerithiopsis (Cerithiopsina) necropolitanaBartschand
Cerithiopsis (CeritMopsina)adamsi
Bartsch.The
first of thesemay
beconsidered the type.Two
specieshave
the early portion of the nuclear whorls smooth,which
is succeededby an
axially ribbed part,which
in turn is fol-lowed
by
a portion bearingtwo
spiral cords in addition to the axial ribs.The
last ornamentation resembles the sculpture of the early post-nuclear whorls, but is less strong, withmany more
axial ribs'Proc.U.S.Nat. Mus.,vol.7,1S84,pp.345-6,pi.2, fig. 4.
2Proc.U.S.Nat.Mus.,vol.9,1886, p. 304, pi.4, fig. 5.
8Mem.CaliforniaAcad.Sci.,vol.3,1903, p. 295, pi.6, fig. 11.
*Proc.U.S.Nat. Mus.,vol. 33, 1907,pp.79-82.
6Prof.Paper No.59,U.S.Geol. Surv.,1909, p. 75.
6Proc.U.S.Nat. Mus..vol. 37, 1909,pp. 399-400andfigure.
no.1823.
MOLLUSKS OF THE GENUS CERITHIOPSIS— BARTSCH. 329
than are presenton
the early post-nuclear turns.To
this group Iwould
apply the subgenericname
CeritMopsida.The two
speciesare Cerithiopsis (CeritMopsida) diegensis Bartsch,which may
be consid- ered the type of the subgenus,and
Cerithiopsis (CeritMopsida) rowelliBartsch.A
fourthgroup, consisting ofthreespecies, has the earlyportionof the nuclear turns smooth; the rest finely, axially ribbed, with the intercostal spaces finely, spirally lirate.For
these Iwould
suggest the subgenus CeritMopsidella, with Cerithiopsis (CeritMopsidella) cosmia Bartsch as type.The
othertwo
species are Cerithiopsis (CeritMopsidella) antejilosaBartschand
Cerithiopsis (CeritMopsidella) alcima Bartsch.The
species inwhich
the nuclearwhorls areunknown
are:excelsa Dall.
antemunda
Bartsch.curtataBartsch. diomedeae Bartsch.
fossilisBartsch. wittiamsoni Arnold.
gloriosaBartsch. truncataDall.
columna
Carpenter. stephensae Bartsch.infrequensC. B.
Adams.
montereyensis Bartsch.paramoea
Bartsch. ingens Bartsch.arnoldiBartsch. tumidaBartsch.
bicolor Bartsch. peruviana D'Orbigny.
magellanicaBartsch. destrugesi
De
Folin.Only
five of the forty-fourknown
species are fossils,and
none, as far asknown,
occur both recentand
fossil.Of
these five,two have
their nuclear characters well preserved
and
thus enable us to assignthem
to their properpositions.These
are Cerithiopsis (Cerithiopsis) fatuaBartschand
Cerithiopsis (Cerithiopsina) necropolitana Bartsch.Both
ofthese arefrom
the PleistoceneofSan
Pedro, California.The
other three species without the nuclear characters are Cerithiopsis excelsa Dall,
from
theEocene
ofOregon; Cerithiopsis fossilis Bartschand
Cerithiopsis wittiamsoni Arnold, from, the Pleistocene ofSan
Pedro, California.Itis
hoped
that the presentpapermay
stimulate thewestAmerican
collectors to be
on
the lookout for thesecharming
little shells,and
thattheir efforts will result inaspeedyeliminationofthespeciesnow
referred to
an
uncertainposition.11Inthe preparationofthe present diagnoses the followingterminologyisused:
" Axial sculpture," themarkings which extend fromthesummitofthewhorlstowardthe umbilicus.
Theaxialsculpturemay be—
"Vertical,"whenthemarkingsareingeneral parallelismwiththe axisoftheshell.
" Protraetive,"whenthemarkingsslantforwardfromthepreceding suture.
"Retractive,"whenthemarkingsslantbackwardfromthe suture.
"Spiralsculpture," themarkingsfollowing the directionsofthecoilsofthe whorls.
330 PROCEEDINGS OF THE NATIONAL MUSEUM.
vol.40.KEY TO THE GENUS CERITHIOPSIS.1
Shellwithvarices excelsa.
Shellwithoutvarices.
Spiralcordsbetweenthe suturesthree.
Spiralcordatsummitequaltothe othersinstrength.
Sutures stronglychanneled.
Shellofmediumsize, lessthan6.5
mm.
long fatua.Shellminute; adultlessthan4
mm.
long.Base withtwospiralcords.
Shell elongate-conic oxys.
Shellelongate-ovate curtata.
Base withmorethantwospiralcords.
Spiralcordsofbase three cerea.
Spiralcordsofbasenine sorex.
Sutures notchanneled.
Whorls shoulderedatthesummit Base withstrongspiralcords.
Shelllarge; adultmorethan7
mm.
long.Spacesinclosedbetweenaxial ribsandspiralcordsonthespire, large squarishpits.
Base witha strong keelnexttoperipheral keel fossilis.
Base with nostrongkeelnexttoperipheral keel gloriosa.
Spacesinclosedbetweenaxial ribsandspiralcordsonthespire,small roundpits.
Peripheral keelcompletely exposedinthe sutureinadultshells,
columna.
Peripheral keel never completely exposed in the suture in adult shell.
Basal cordsthree,strong cosmia.
Basal cordsingle,feeble " carpenteri.
Shellsmaller.
Adultbetween3and5
mm.
long.Basal cordbroad and round.
Axialribsonlastwhorl about32 pedroana.
Axialribsonlastwhorlabout20 tuberculoides.
Basal cords sublamellar.
Adultlessthan3
mm.
long infrequens.Shellstout pupiformis.
Shell slender abreojosensis.
Base withfeeblesculpture only.
Basalfasciolepresent.
Base withasingle fasciole (encirclingthecolumella) only.
Shellbroadlyelongate-conic paramoea.
Shellslender elongate-conic necropolitana.
Base withmorecordsthanasingle fasciole arnoldi.
Basalfascioleabsent.
Whorlsnotshoulderedatthesummit magellcmicacesta.
Spiralcordatsummitweaker thantherest.
Sutures strongly channeled.
Shellrobust.
Basal keels strong berryi.
1Since somanyofthe speciesunderdiscussion lackthe nucleus,itwasdeemedadvisableto treatthe genusasa unitandignorethesubgenericdivision inthiskey,byutilizingcharacterspresentonthe post- nuclearwhorlsonly.
no.1823.
AIOLLUSKS OF THE GENUS CERITHIOPSIS— BARTSCH. 331
Basal keelobsolete.
Axial ribs retractive antemunda.
Axial ribs protractive diomedese.
Shell very slender galapagensis.
Sutures notchanneled.
Tubercles ofmiddle keeltruncated on both posteriorand anteriormargin.
Axialribsandspiralcords inclosingstrong,squarishpits.
Base withtwostrongkeels.
Adultshellmore than5
mm.
long alcima.Adultshelllessthan3
mm.
long williamsoni.
Base withafeeblefascioleonly truncata.
Axialribsandspiralcords inclosingroundedpits.
Spiralcords slender cesta.
Spiral cordsverybroad.
Shellveryelongate-conic stephensse.
Shellbroadlyconic.
Adultshelllessthan6
mm.
long stejnegeri.Adultshellmorethan7
mm.
long s.dina.Tubercles of middle keel not truncated on both posterior and anterior margin. Tuberclesof middlekeeltruncated onposteriormarginonly.
Basemarkedbya slenderfascioleonly rowelli.
Basemarked byspiralcords.
Spiralcordsofbase three antefilosa.
Spiralcordsofbasetwo neglecta.
Tuberclesofmiddlekeel not truncated.
Whorlsstrongly, slopingly shouldered atthesummit halia.
Whorlsnot slopingly shoulderedatthe summit.
Shellrobust.
Base withasinglekeelattheperiphery diegersis.
Base withthree keelsnear the periphery montereyensis.
Shellslenderandminute adamsi.
Spiralcordsbetweenthe suturesfour.
Shellverylarge;adultmorethan12
mm.
long.Intercalatedcordsabsentorveryfine ingens.
Shell smaller; adultlessthan7
mm.
long.Spiralcordswider thanthe spaceswhichseparatethem.
Shellelongate-conic tumida.
Shellelongate-ovate peruviana.
Spiralcordsnarrower thanthe spaceswhichseparatethem.
Spiral cordssubequal aurea.
Spiralcordsdecidedly unequal destrugesi.
CERITHIOPSIS (CERITHIOPSIS) FATUA, new species.
Plate36, fig.5.
Shell very elongate-conic,
creamy
white. Nuclear whorls three, strongly rounded, separatedby
wellmarked
sutures, forming aslen- der, regularspire. Post-nuclear whorls well rounded,separatedby
a deeply channeled suture,ornamented
with three nodulose, spiral cordsoneofwhich
is atthe summit, thethird immediatelyabove
the periphery, while thesecondismedian between
the two. Thesecords are alittle wider than the spaceswhich
separate them. In addition to the spiral sculpture, the whorls aremarked by
retractive, axial332 PROCEEDINGS OP THE NATIONAL MUSEUM.
vol.40.ribs
which
are as strong as the spiral cordsand
render the junctions with these nodulose.Of
these ribs 16 occurupon
the first to fifth,18
upon
thesixth, 20upon
theseventh,and
22upon
thepenultimate turn.The
spaces inclosedbetween
the axial ribsand
spiral cords appear as strongly impressed,rounded
pits.The
tubercles are slightly truncated posteriorly, sloping gently anteriorly. Periphery of the last whorlmarked by
a channel like those occurring between the spiral cordson
the spireand
likethem
crossedby
the continua- tions of the axial ribs.Base
rather long, well rounded, freefrom
sculptureexcepting very fine, incrementallines. Aperture regularly ovoid, strongly channeled anteriorly; posterior angle obtuse, outer lip thin, renderedwavy by
the external sculpture; columella stout, decidedly curvedand somewhat
twisted, projecting considerablybeyond
the anteriormargin
ofthe outerlip.The
typeand
seven specimens (Cat. No. 195194, U.S.N.M.)come from
theLower
Pleistocene(Lower San Pedro
Series),Deadmans
Island, California.
The
type has twelve whorlsand
measures:Length
6mm.,
diameter 1.6mm.
CERITHIOPSIS(CERITHIOPSIS) OXYS, newspecies.
Plate36,fig.2.
Shellminute, elongate-conic, dark brown. Nuclear whorls almost four, forming aslender, elongate-conic spire, having the whorls well rounded. Post-nuclearwhorls wellrounded, separated
by
astrongly channeled suture,marked
b}^ three equally strong, spiral cords, the firstofwinch
isat thesummit,
the thirdslightlyabove
the periphery, while the second ismedian between
these two. In addition to the spiral cords, the whorls aremarked by
vertical, axial ribs, almost as strong as the spiral cords.Of
these ribs, 16 occurupon
the first,20
upon
the second, 18upon
the third to fifth,and
24upon
the penultimate turn.The
junctions of the axial ribsand
spiral cordsform
strong tubercles ofwhich
those ofthe posteriorcordarerounded
while the othertwo
are truncated posteriorlyand
slope gently ante- riorly, while the spaces inclosedbetween
the spiral cordsand
axial ribs are strongly impressed,rounded
pits. Periphery of the last whorlmarked by
astrong cordwinch
isfeeblynoduloseand
separatedfrom
thetuberculate keel posterior toitby
a channelaswideas those occurringon
the spire; tins channelismarked by
the continuations of the axial ribs.Base
moderately long,marked by
astrong, spiral cord about halfwaybetween
the peripheralcordand
the insertion of the columella, the spaceswhich
separate itfrom
the peripheral cordon
onesideand from
the columellaon
the other, appearingas deep,rounded
channels. Aperture rather small, irregularly ovate, very strongly channeled anteriorly; posterior angle obtuse; outer lip ren-no. 1823.
MOLLUSKS OF THE OENUS CERITHIOPSIS—BARTXCH
.333
deredsinuousby
theexternal sculpture; columellaverystout,twisted revolute,and
reflected; parietalwall glazed with a thincallus.The
typeand
another specimen (Cat. No. 106505, U.S.N.M.)come from
Point Abreojos,Lower
California.The
type has seven post- nuclear whorlsand
measures:Length
3.3mm.,
diameter 1.2mm.
Two
additional specimens (Cat. No. 195181, U.S.N.M.)come from San Pedro
Bay, California.CERITHIOPSIS (CERITHIOPSIS) CEREACarpenter.
Plate37, fig. 6.
CerithiopsiscercaCarpenter, Cat.Maz. Shells, 1857, p. 443.
Shell small, broadly conic, semitransparent, light horn-color.
Nuclear whorls well rounded. Post-nuclear whorls separated
by
a strongly channeled suture,marked by
three spiral cords, the first ofwhich
is a little anterior to the summit, the second median,and
the third alittleposterior to the suture. In addition to the spiral cords, the whorls aremarked by
axial ribswhich
equal the spiral cords in strength.Of
these ribs, 12 occurupon
thefirst to third, 14upon
the fifthand
sixth,and
22upon
the penultimate turn.The
junctions of the spiral cordsand
the axial ribsform
prominent tubercles, while the spaces inclosedbetweenthem
are elongated, oval pitson
the early whorls, having their long axes parallel to the spiral sculpture.On
the last volution they are rounded. Periphery of the last whorl
marked by
a strong channel.Base
moderately long, well rounded,marked by
three strong, spiralkeels, ofwhich
thefirstis immediately below the peripheral sulcus,and
the second halfway between thisand
the cordwhich
surrounds the insertion of the columella. In addition to the spiral cords, the base ismarked by
numerous, fineincrementallines. Aperture irregular, very strongly channeled ante- riorly; posterior angle obtuse; edge of outer lip rendered sinuous
by
the external sculpture; columella moderatelylong, twisted, with the edgereflected.DoctorCarpenter'stypeison Tablet No. 2030,Liverpoolcollection, British
Museum, and was
collectedon
Spondylus at Mazatlan. It has seven post-nuclear whorlsand
measures:Length
2.4mm.,
diameter 1mm.
CERITHIOPSIS (CERITHIOPSIS)SOREXCarpenter.
Plate37,fig.2.
CerithiopsissorexCarpenter, Cat.Maz. Shells, 1857, p. 444.
Shell minute, pupiform. Nuclear whorls three, smooth, well rounded, forming a slender,
mucronate
apex.The
first three post- nuclear whorls increasing very rapidly in size, well rounded, orna-mented
with three spiral cords, ofwhich
the first is a little anterior334 PROCEEDINGS OF THE NATIONAL MUSEUM.
vol.40.tothe summit, the second median,
and
the thirdalittle posterior to the suture. In addition to the spiral cords, the whorls aremarked by
retractive axial ribswhich
are almost as strong as the spiral cords.Of
these ribs, 16 occurupon
the second, 18upon
the third,and
20upon
thepenultimate whorl.The
junctions of the axial ribsand
the spiral cordsform prominent
tubercles, while the spaces inclosedbetween them
are elongated pitson
the second whorland
well-rounded, well-impressed pitson
the remaining ones. Sutures channeled. Periphery of the last whorlmarked by
a deep sulcuswhich
is crossedby
the continuations of the axial ribswhich
extend to the posterior edge of thefirst basal cord.Base
moderately long, concave in the middle,marked by
eight low, rounded, subequally spaced, spiral cords,which grow
successivelyweaker from
theperiph- erytoward
the tip of the columella. Aperture irregular, decidedly channeled anteriorly; posterior angle acute; outer lip rendered sinuous at the edgeby
theexternal sculpture; columella moderately long, twisted, theedge reflected,and
joined to the posterior angle of the apertureby
a strong calluswinch
covers theparietal wall.The
type,which
ison
Tablet No. 2032, Liverpoolcollection,BritishMuseum,
hasfivepost-nuclearwhorlsand
measures:Length
1.6mm.,
diameter 0.7mm.
Itand
three additional specimens were foundon
Spondylus atMazatlan.CERITHIOPSIS(CERITHIOPSIS)CARPENTERI,newspecies.
Plate38, fig. 9.
Shell broadly elongate-conic, dark chocolate brown. Nuclear whorls three, smooth, well rounded, separated
by
a slender suture, formingan
elongate-conic spire. Post-nuclear whorls well rounded,marked by
three strong, broad, spiralbands which
are aswide as the spaces that separate them.Of
these, the firstand
widest is at thesummit,
thenextis median, while the third is alittleposterior to the suture. In addition to the spiral keels, the whorls aremarked by
slender, rounded, axial ribs
which
are abouthalf as wide asthespiral cords.Of
these ribs, 20 occurupon
thefirstand
second,22upon
the third tofifth, 24upon
the sixthand
seventh, 28upon
theeighth,and
30upon
the penultimate turn.The
junctions of the axial ribsand
spiral cords
form
strong, elongated tubercles, having their long axes parallel with the axial sculpture.The
tubercles at thesummit
are wellrounded
anteriorlyand
posteriorly;thoseofthemedian
series are alittlemore
suddenlyrounded
anteriorlythanposteriorly;while those belonging to the supraperipheral cord are truncated suddenly pos- teriorlyand
slope gently anteriorly.The
spacesbetween
thespiral cordsand
axial ribs are small, moderately rounded, well-impressed pits. Suture constricted,showing
a portion of the basal cord.Peripheryofthelastwhorl
marked by
a deep,spiralgrooveas wideasno.1823.
MOLLUNKti OF THE GENUS CERITHIOPSIS— BARTSCH. 335
that separating thefirst supraperipheral cordfrom
themedian
spiral cordand, likeit,crossedby
the continuationsoftheaxial ribs.Base
short,wellrounded,
marked by
a broad,spiralcordwhich
istruncated posteriorlyand
slopes gently anteriorlytoward
the shallow, well-marked
groovewhich
separates itfrom
the baseproper. Entiresur- faceof spireand
basemarked by
numerous, veryslender,incrementallines. Apertureirregularly oval, verystrongly channeled anteriorly;
posterior angle obtuse; outerlipthin,
showing
the external sculpture within, rendered decidedly sinuous at the edgeby
the external sculp- ture; columella stout, moderately long, the edge reflected; parietal wallglazedwitha thincallus.The above
descriptionisbasedupon
thetwo
cotypeswhich, together with 130specimens, areregistered asCat. No. 109510, U.S.N.M.,and come from
TerminalIsland, California.One
of these specimens is ayoung
individual consisting of the three nuclear whorlsand
sixpost- nuclear whorls; the otherisan
adultwhich
haslostthe nuclearwhorlsand
has ten post-nuclear whorlswhich
measure:Length
8.1mm.,
diameter 2.8mm.
Named
for the late Dr. Philip P. Carpenter.Specimensexamined.
Catalogue No.
i109510 109511 173088 73725 195178 152758 159330 134857
'15653
i130584 56005 32219 32287 130315 3566 32220 56006 123401
Locality.
TerminalIsland,California.
do ....do
SanPedro, California
White'sPoint,San Pedro Bay,California.
AlamitosBay,California SanDiego, California ....do
.do.
SanDiego, California (on sponge).
SanDiego, California do
.do.
FalseBay,SanDiego, California
San Diego Bay,California(3fathoms,sandandshellbottom, bottom temperature58°)
SanDiego, California ....do
SanDiego, California(livingonsponges)
Numberof specimens.
132 1 21 1 2 35 5 3
1 4 17 15 114 1
1
48 18
1Cotypes.
CERITHIOPSIS (CERITHIOPSIS)PEDROANABartsch.
Plate38,fig.2.
CerithiopsispedroanaBartsch, Proc.U.S. Nat.Mus.,vol.33, 1907,pp. 181-182.
Shell small, slender, dark brown. Nuclear whorlsthree, yellowish- white, smooth. Post-nuclear whorls strongly differentiated
from
the nuclear ones,showing
the sculpture characteristic of the adult shellfrom
the very beginning. This sculpture consists of three equally spaced tuberculatespiralkeelsbetween
thesutures, the posteriorone ofwhich
isslightlysmaller thantheother two. Thesekeels aresepa-'336
PROCEEDINGS OF THE NATIONAL MUSEUM.
vol.40.rated
by
deep,rounded
channels almost aswideas the keel. In addi- tion there aremany
low, rather broad axial ribs, the intersections ofwhich
withthe keelform
thetubercles.About
20ofthem
occurupon
thefirst, 22
upon
thefifth,and
30upon
thepenultimate post-nuclear turn.The
connectionsbetween
the tubercles, bothspiraland
axial,1 are about equal, inclosing deep, squarish pits. In addition to theabove
sculpture the entire surface ismarked by
fine spiral linesand
lines of growth. Sutures strongly marked, constricted,
showing
the peripheral keel in the later whorl. Peripherymarked by
a broad, low,rounded
keel.Another
ofequalwidth
islocatedupon
themiddle of the base.The
sulcuswhich
separates these keelsand
the supra- peripheral sulcus are of equal width; both are crossedby
theweak
continuations of the axial ribs,
which
graduallyweaken
as they passtoward
the columella.The
basalkeelisseparatedfrom
the columellaby
broad, shallow grooves. Aperture irregularly oval, decidedly channeled anteriorly, outerlipthin, renderedsinuousby
the external keel;columella stoutand somewhat
twisted,witha strongcalluson
itsinneredgethat extends overthe parietal wall.
The two
cotypes, Cat.No.
109512,U.S.N.M., werecollectedby Mrs/
W.
II.Eshnaur
atTerminal Island,San
Pedro, California.The
one has the nucleusand
three post-nuclear whorls; the otherhas lostthe nucleusand
has nine post-nuclear turnsand
measures:Length
5.2mm.,
diameter 1.8mm.
Specimensexamined.
Catalogue No.
i109512 56751 195179 14825 195180 737250 130584 32287a 322206 56006a 123401 109364 199179 153058 106504 32292
Locality.
TerminalIsland, California Catalina Island, California SanPedro, California
do
White'sPoint,San Pedro,California SanPedro, California
SanDiego, California do
do do do do do
GovernmentJetty,SanDiego,California.
Point Abreojos,LowerCalifornia Todos Santos Bay,LowerCalifornia
Numberof specimens.
350 1 3 1 1 1 9 40 13 3 6 1 1 7 1 1
1Cotypes.
CERITHIOPSIS(CERITHIOPSIS) TUBERCULOIDESCarpenter.
Plate 37, fig. 7.
Cerithiopsis tuberculides Carpenter, Cat. Maz. Shells, 1857, pp. 442-443.
Shell elongate-conic. Nuclear whorls five, well rounded, smooth, separated
by
a moderatelyconstricted sutureforming a well elevated spire. Post-nuclear whorls moderately rounded,ornamented
withno. 1823.
MOLLUSKS OF THE GENUS CERITHIOPSIS— BARTSCH. 337
three spiral cords ofwhich
the first is a little below thesummit,
the secondmedian and
the third a little posterior to the suture. These cords areseparatedby
spaces alittle wider than thecords. In addi- tion to the spiral sculpture, the whorls aremarked by
decidedly retractive, axial ribswhich
are about as strong as the spiral cords.Of
these ribs, 12 occurupon
the firstand
second, 14upon
the third, ISupon
the fourth,and
20upon
the penultimate whorl.The
junctions of the axial ribs
and
spiral cordsform
wellrounded
tuber- cles, while the spaces enclosedbetween them
are deep,rounded
pits.Sutures moderately constricted. Periphery of the lastwhorl
marked
with a strong, smooth, spiral cordwhich
is separatedfrom
the firstsupraperipheral cord
by
a channel as wide as thatwhich
separates the supraperipheral cordfrom
themedian
and, like it, is crossedby
the continuations of the axial ribs,which
extend to the posterior edge of the peripheral cord.Base
moderately long, well rounded,marked by two
broad rounded, spiral cords. Aperture irregularly oval, decidedly channeled anteriorly; posterior angle obtuse; outer lip rendered sinuousby
the external sculpture; columella curved, twisted, theedge reflected; parietal wall coveredwith a thick callus.Nine specimens of thisspecies werereported
by Doctor
Carpenter takenfrom
Spondylusand Chama
atMazatlan.The
cotypes areon
Tablet 2028, Liverpool collection, BritishMuseum. The
larger, a perfect specimen with seven post-nuclear whorls, measures:Length
3.75
mm.,
diameter 1.2mm. Our
figure is after acamera
lucida sketch of the typeby
Doctor Carpenter.CERITHIOPSIS (CERITHIOPSIS)TUBERCULOIDES,var.ALBONODOSA Carpenter.
Plate 37, fig. 3.
Cerithiopsis ? tuberculoidcsalbonodosaCarpenter, Cat. Maz. Shells, 1857, p. 443.
Of
thisDoctor
Carpentersays:
Shell similar totuberculoideswiththe tuberclesmoredistantandwhite. Nuclear whorlsfour, lesselevatedandsubcarinate.
Tablet 2029, Liverpool collection, British
Museum,
contains ayoung
shelland
a fragment ofan
adult shell takenfrom
Spondylus atMazatlan, Mexico.Our
figure is after acamera
lucida sketch of the typeby
Doctor Carpenter.CERITHIOPSIS (CERITHIOPSIS)PUPIFORMISCarpenter.
Plate 38, figs. 1and5.
CerithiopsispupiformisCarpenter, Cat. Maz. Shells, 1857, pp. 443-444.
Shellminute,elongate-conic,brown. Nuclear whorlsfour,smooth, separated b}T a constricted sutureforming a
mucronate
apex. Post- nuclear whorlsmarked by
three spiral cords ofwhich
the first is at80796°—Proc.N.M.vol.40—11 22
338 PROCEEDINGS OF THE NATIONAL MUSEUM.
vol.40.the
summit,
the third immediately posterior to the suture, while the second is halfway between
these two.These
cords are a littlewider than the spaces that separate them. In addition to the spiral cords, the whorls are
marked by somewhat
retractive, axial ribs alittle less strong than the spiral cords.
Of
these 20 occurupon
the second, 22upon
the thirdand
fourth,and
24upon
the penultimate turn.The
junctions of the axial ribsand
spiral cordsform
wellrounded
tubercles, while the spacesbetween them
are smallrounded
pits. Sutures well impressed. Periphery of the last whorl
marked by
a deep spiral sulcus.Base
moderately long withtwo
tumid, broad, rounded, spiral cords. Aperture irregularly oval, decidedly channeled anteriorly; posterior angle obtuse; outer lip rendered sinuousby
the external sculpture; columella curvedand
twisted, with the edge reflected; parietal wall covered with a thick callus.Tablet2031,Liverpoolcollection, British
Museum,
containsDoctor
Carpenter'stwo
cotypeswhich
werecollectedon
SpondylusatMazat-
lan,Mexico.
The
largest ofthesecotypesmeasures:Length
1.9mm.,
diameter 0.3mm. Our
figure is after acamera
lucida drawingby Doctor
Carpenter of the type.CERITHIOPSIS (CERITHIOPSIS) ABREOJOSENSIS,newspecies.
Plate37,fig.4.
Shell minute, elongate-conic, brown, excepting the nuclear whorls
which
are white. Nuclear whorls fourand
one-half, well rounded, smooth, separatedby
a moderately constricted suture, formingan
elevated, cylindric spirewhich
givestheapex
oftheshellamucronate
appearance. Post-nuclearwhorls veryslightlyrounded,ornamented by
three spiral cords, the firstofwhich
is at thesummit,
thesecond at the anteriorterminationofthe posteriorthirdbetween
thesutures, while the last is considerably posterior to the suture.The median
cord, therefore, is
much
nearer the cord at thesummit
than that at thesuture. In addition to these spiral cords, thewhorls aremarked
by
very slightly retractive, axial ribswhich
are almost as strong as thespiral cords.Of
theseribs 16occurupon
allbut thepenultimate turn,winch
has 18.The
junctions of the axial ribsand
the spiral cordsform
wellrounded
tubercles,while the spaces enclosedbetween
them
are elongated slitsbetween
the firstand
second spiral cordsand
elongated, almost squarish pitsbetween
themedian and
the peripheralcord. Suturesstronglyconstricted. Peripheryofthelast whorlmarked by
a spiral sulcus as wide as thatwinch
separates the supraperipheralcordfrom
themedian
cordbetween
the sutures,and
crossedby
the continuations of the axial ribs.Base
moderately long,marked by two
strong, spiralcords, oneofwhich
isimmediately belowthe periphery, while the otherisalittleposterior tothemiddle of thebase, thetwo
beingseparatedby
a channelequaling the periph-no. 1823.
MOLLUSKS OF THE GENUS CERITHIOPSIS— BARTSCH. 339
eral one in width. In addition to the
above
sculpture, the base ismarked by weak
incremental lines. Aperture irregularly ovate, very strongly channeled anteriorly; posterior angle obtuse; outer lip thin, renderedsomewhat
sinuousby
the external sculpture;columella stout, curved,and somewhat
twisted, the edge reflected; parietal wall glazed with a thick callus.The
type (Cat. No. 106506, U.S.N.M.)comes from
Point Abreojos,Lower
California. It has six post-nuclear whorlsand
measures:Length
2.3mm.,
diameter0.8mm.
CERITHIOPSIS (CERITHIOPSIS) BERRYI, new species.
Plate39,fig.8.
Shell small, elongate-come, brown. Nuclear whorls four, well rounded, separated
by
constrictedsutures, smooth, formingamucro-
nateapex
to the shell. Post-nuclear whorls strongly sculptured, having three spiral cordsbetween
the sutures, ofwhich
the one at thesummit
ismuch
smaller than the reston
the early whorls; the othertwo
aresubequal instrength, thesecond one beingclose to the one at thesummit,
while the third is almost halfwaybetween
the secondand
the suture.On
the lastwhorl, the cordat thesummit
isalmost equal to the other two. In addition to the spiral sculpture, thewhorls are
marked by
verticalaxial ribswhich
are stronger than thespiralcords.Of
theseribs, 12 occurupon
the second, 16upon
the third to fifth, 18upon
the sixth,and
20upon
the penultimate turn.The
junctions of the axial ribsand
spiralcordsform
strongtubercles,which
are truncated posteriorlyand
slope gently anteriorly.The
spaces inclosed
between them
are elongated,narrow
pitsbetween
thefirstand
secondspiralcordson
the early whorls, whileon
thelasttheyaresquarishpitsin this region.
The
spacesbetween
themedian and
supraperipheral cord are stronglyimpressed, large, squarish pitson
all the whorls. Sutures strongly channeled. Periphery of the last whorlmarked by
a broad sulcus, equaling thatwhich
separates the supraperipheralfrom
themedian
cord, crossedby
the continua- tions of the axial ribs.Base marked by
a strong, broad,,rounded
cordimmediately below theperipheryand
asecondless strongon
its middle, while a slenderthread encirclestheinsertion of the columella.In addition to this sculpture, the base is
marked by
strong, incre- mental lines. Aperture irregularly oval, decidedly channeled ante- riorly; posterior angle obtuse; outer lip thin,showing
the external sculpture within, rendered sinuous at theedgeby
the external sculp- ture;columellastout,curved, stronglytwisted,withtheedgereflected:parietal wall glazed with a thick callus.
The
typeand two
additionalspecimens (Cat.No. 195182,U.S.N.M.) were dredgedby
Mr. S. S. Berryin 12 fathomsoffDel Monte,Monte-
rey, California.
The
type haslostthenucleus, the eight post-nuclear340 PROCEEDINGS OF THE NATIONAL MUSEUM.
vol.40.whorls measuring:
Length
3.4mm.,
diameter 1.3mm. The
nucleuswas
describedfrom
one of four specimens (Cat. No. 195198a, U.S.N.M.)from Whites
Point,San
Pedro, California. Cat. No.195183, U.S.N.M., contains seven specimens
from
Monterey,and
Cat. No. 195184, U.S.N.M., contains four specimensfrom San Pedro
Bay, California.CERITHIOPSIS (CERITHIOPSIS) GALAPAGENSIS. newspecies.
Plate39,fig. 4.
Shell minute, regularly elongate-conic, golden brown, with the
row
of tubercles at thesummit
of the shell darker. Nuclear whorlsfive, well rounded, smooth, separated
by
a well-impressed suture.Post-nuclear whorls moderately rounded,
ornamented
with three tuberculate, spiral cords, ofwhich
the first (which is at thesummit)
is considerably less strong than the other two; the third is
imme-
diately
above
the suture, while the second is considerably nearer thefirst thanthe third.
The
spacebetween
the firstand
secondis about half the width of the middle cord, while thatbetween
the secondand
third is equal to it. In addition to the spiral cords, the whorls aremarked by
axial ribs,which
are about equal to the spiral cords in strength; of these ribs, 16 occurupon
all but the penultimate whorl,which
has 20.The
intersections of the axial ribsand
spiral cordsform
prominent tubercles,which
are truncated posteriorlyand
slope gently anteriorly, while the spaces inclosedbetween them
are elongated pitsbetween
the firstand
second spiral cords, having their long axes parallel with the spiral sculpture,and
squarish pitsbetween
the secondand
third cords. Sutures strongly channeled.Periphery of the last whorl
marked by
a channel almost as wide as thatwhich
separates the supraperipheral cordfrom
themedian
and, like it, crossedby
the continuations of the axial ribs.Base moder-
ately long, concave,marked by
a strong spiral cord immediatelybelow
the peripheral sulcus,and
another almost as strongwhich
encircles the columella at its insertion. In addition to the spiral sculpture, the base is
marked by
fine, incremental lines. Aperture irregularly triangular, strongly channeled anteriorly, very effuseand
angulated at the junction of the basaland
outer lip; posterior angle very obtuse; outer lip thin,showing
the external sculpture within, rendered sinuous at the edgeby
the external sculpture; columella moderately long, stout, curved, with the edge reflected; parietal wall coveredwith a thick callus.The
typeand two
additionalspecimens (Cat.No. 195185,U.S.N.M.)were
dredgedby
the U. S.Bureau
of Fisheries steamer Albatross at station 2813 in 40 fathoms,on
coral sand bottom,bottom
tempera- ture 81°, off Galapagos Islands..The
type is a perfect specimen, having six post-nuclear whorlsand
measures:Length
2.4mm.,
no. 1823.
MOLLVSKS OF THE GENUS CERITIIIOPSIS—BARTSCn. 341
diameter 0.8mm.
Cat. No. 195187, U.S.N.M., contains three speci-mens
from thesame
locality. Cat. No. 122128, U.S.N.M., containstwo
specimens dredged off Indefatigable Island, Galapagos Islands.CERITHIOPSIS (CERITHIOPSIS) CESTA, newspecies.
Plate39, fig.5.
Shell elongate-conic, chestnut brown. Nuclear whorls, three, well
rounded and smooth
forming a slender elongate spire. Post-nuclear whorls strongly rounded,marked
with three feebly tuberculated spiral keelsbetween
the sutures ofwhich
the posterior is at thesummit and
the anterior atsome
little distance posterior to the suture, themedian
one being a little nearer the anterior than itsother neighbor.
The
spacesbetween
these keels are not quite as wide as the keels. In addition to the spiral keels, the whorls aremarked
with feeble,somewhat
irregular axial ribletswhich
are lessthan half as strong as the spiral keels; of these ribs, 14 occur
upon
the thirdand
32upon
the remaining whorls. These ribs render their junction with the spiral cords feebly nodulose.The
spaces inclosedbetween
the spiral cordsand
the axial ribs are feebly im- pressed, squarish pits. In addition to theabove
sculpture, the entire surface of the shell is crossedby
fine incremental linesand
very fine spiral striations. Sutures strongly impressed, with the peripheral cordshowing
as a slender,smooth band
in all but thefirst
two
whorls. Periphery of the last whorlmarked by
a broad, depressed spiral cordwhich
is separatedfrom
the first suprasutural cordofthe spireby
a sulcusaswide as that separatingthis cordfrom
its posterior neighbor. This groove is crossed
by
the continuations of the axial ribs.Base
large, moderately rounded,marked
with a slender spiral cordwhich
is a little nearer the peripheral cord than the slender fasciole at the insertion of the columella. Aperture irregularlyovate; outer lip thin, rendered sinuous at theedgeby
the external sculptureshowing
the external sculpture within; columella stout, rather long, twistedand
curved^parietal wall glazed with a thin callus.The above
description is basedon two
individuals, onean
adult (Cat. No. 153057, U.S.N.M.) collected at theGovernment
Jetty,
San
Diego, California. This has lost all but the last nuclear whorl, having eight post-nuclear whorlsand
measuring:Length
5.8
mm.,
diameter 2.1mm. The
other is one of three specimens (Cat.No.
160079 U.S.N.M.) collected atSan
Diego. This has the nucleusand
six post-nuclearwhorlsand
measures:Length
3.2mm.,
diameter 1.6mm.
342 PROCEEDINGS OF THE NATIONAL MUSEUM.
vol.40.CERITHIOPSIS (CERITHIOPSIS) STEJNEGERI Dall.
Plate40,fig.3.
Cerithiopsis stejnegeriDall,Proc.U.S.Nat. Mus.,vol.7,1884,pp. 345-346,pi. 2,fig.4.
Shell broadly conic, light chestnut brown, excepting the nucleus
and
the columella. Nuclear whorls two, strongly rounded, smooth.Post-nuclear whorls well rounded,
marked
with three nodulose spiral keels, ofwhich
oneis at theshoulderedsummit,
anothersome- what
posterior to the suture, while the third ismedian between
the two.The
anterior of the three is the strongerand
the one at thesummit
the weakest. These cords are about as wide as the spaces that separate them. In addition to the spiral cords the whorls aremarked by
very poorly developed, almost vertical axial riblets, the junctions ofwhich
with thespiral cordsform
the nodules.Of
these ribs 16 occurupon
the second, 18upon
thethird,22upon
thefourth, 24upon
the fifth,and
26upon
the penultimate whorl.The
spaces inclosedbetween
thespiralcordsand
axial ribs are irregular,shallow, impressed,rounded
pits. Sutures strongly constricted,showing
the posterior edge of the base. Periphery of the last whorlmarked by
a sulcuswhich
is crossedby
the feeble axial riblets; base well rounded, smooth, excepting incremental lines. Aperture broadly ovate; decidedly channeled anteriorly; posterior angle obtuse; outer lipthin,renderedwavy by
the external sculptureshowing
theexternal sculpture within; columellashort, stout,twisted,and
curved;parietal wall glazed with a thin calluswhich
extendsfrom
the inner edge of thecolumellato the posterior angle ofthe aperture.The
type (Cat. No. 40932, U.S.N.M.)was
collectedby
Dr. L.Stejneger atBering Island. Ithas eightpost-nuclear whorls,having lost the nuclear turns
and
measures:Length
5.5mm.,
diameter 2.2mm. The
nuclear whorlswere
describedfrom
a specimen of Cat.No. 195172,
U.S.N.M.
Specimensexamined.
U.S.N.M.
Cat.No. Locality.
Numberof specimens.
i40932 161049 195172 130625 195174 161108 161036 195173
BeringIsland
KyskaUarbor, AleutianIslands,Alaska AmchitkaIslands,AleutianIslands,Alaska
AmchitkaIslands,AleutianIslands,Alaska(lowwater) ConstantineHarbor, AleutianIslands,Alaska ...do
CaptainsHarbor, Unalaska, AleutianIslands,Alaska...
EastsideSimeonoffIsland,ShumaginIslands,Alaska...
Type.
no. 1823.
MOLLUSKS OF THE GENUS CERITHIOPSIS—BARTSCII. 343
CERITHIOPSIS (CERITHIOPSIS) STEJNEGERI DINA,newsubspecies.
Plate40,fig.7.
This is the southern representative of 0. stejnegeri, differing
from
the species in having the sculpturemuch more
stronglypronounced and
in being larger. Three specimens areknown
(Cat. No. 195175, U.S.N.M.)which come from
Sitka, Alaska.The
type has three nuclearwhorlsand
seven post-nuclearwhorlsand
measures:Length
7
mm.,
diameter2.6mm.
CERITHIOPSIS (CERITHIOPSIS) NEGLECTA C.B.Adams.
Plate40,fig.5.
CerithiumneglectumC. B.Adams, Ann. N. Y. LyceumNat. Hist., vol. 5, 1852, p. 378.
Shell minute, elongate-conic, dark
brown
excepting the nuclear whorls,which
arewhite. Nuclear whorlsatleast four, wellrounded, smooth, forming amucronate
apex. Post-nuclear whorls orna-mented
with three spiral cords, the first ofwhich
is very feebleon
the early whorlsand even
less strong than the othertwo on
the lastvolution; the second cord is at the posterior termination of the anterior thirdbetween
the sutures, while the third is a little nearer the suture than the median.The
firstand
second are only about half as far apart as themedian and
third. In addition to the spiral sculpture, the whorls aremarked by
retractive axial ribs,which
are about as strong as the spiral cords.Of
these ribs, 16 occurupon
thefirst, 14upon
thesecond tofourth, 16upon
the fifthand
sixth,and
18upon
the penultimate turn.The
intersections of the axial ribsand
the spiral cordsform
strong, cusped tubercles.Those on
the firstspiral cord are theweakest and
are wellrounded
;
those
on
thesecondand
third are truncatedposteriorlyand
slope lessabruptly anteriorly.
The
spaces inclosedbetween
the axial ribsand
spiral cords are elongated pitsbetween
the firstand
second spiral cords, having their long axes parallel to the spiral sculpture, while thosebetween
the secondand
third are strongly impressed, squarish pits. Suture strongly constricted. Periphery of the last whorlmarked by
a strong, spiral cord, separatedfrom
the supra- sutural cordby
a sulcus as wide as thatwhich
separates the supra- suturalfrom
themedian
and, like this, crossedby
the continuations ofthe axialribs,which
terminate atthe posterioredgeofthe periph- eralcord.Base
moderately long,ornamented
withtwo
spiral cords.Aperture irregularly subquadrate, decidedly channeled anteriorly;
posterior angle acute; outer lip rendered
wavy by
the external sculpture,which
is apparent within the aperture; columella short, stout, curved, with the free edge reflected; parietal wall covered344 PROCEEDINGS OF THE NATIONAL MUSEUM.
vol. 40."with a thick callus
which
connects the columella with the posterior angle of the aperture.The
specimen describedand
figured (Cat.No.
211566,U.S.N
.M.) isone collectedby
Prof. C. B.Adams
atPanama.
Ithaseight post- nuclear whorlsand
measures:Length
3.2mm.,
diameter1.1mm.
CERITHIOPSIS (CERITHIOPSIS) HALIA, newspecies.
Plate40, fig.8.
Shell small, broadly elongate-conic, chocolate brown. (Early nuclear whorls decollated.)
Only
the last volution remains,which
is smooth. Post-nuclear whorls strongly shouldered at the
summit on
the early whorls,ornamented
with threespiral cords, of whichthefirst, at the
summit,
is very poorly developed; the second issituated at theposteriortermination ofthe anteriorthirdbetween
the sutures;while the third,
which
equals the second, is about halfwaybetween
this
and
the suture. In addition to the spiral cords, the whorls aremarked by
decidedly retractive, axial ribswhich
are almost equal to thespiral cordsinstrength.Of
theseribs, 12 occurupon
thefirst,14
upon
thesecond, 16upon
the thirdand
fourth, 18upon
the fifth,and
20upon
the penultimate turn.The
intersections of the axial ribsand
spiral cordsform
well-rounded tubercles, while the spaces inclosedby them
are elongated pitsbetween
the firstand
second spiral cord,and
large, squarish pitsbetween
the secondand
third.Periphery of the last whorl
marked by
a sulcus not quite as broad as thatwhich
separates the supraperipheralfrom
themedian
cord,and
crossedby
thecontinuations of the axial ribs.Base
short, con- cave,marked by two
spiral cords immediately below the peripheral sulcusand by numerous
fine, incremental lines. Aperturesubquad-
rate, decidedly channeled anteriorly; posterior angle obtuse; outer lip rendered sinuous at the edge
by
the external sculpturewhich
is apparent in the aperture; columella short, curved,
and
twisted, with thefree edge reflected; parietal wallglazed with a thin callus.We have
seen three specimens of this species,none
ofthem
quite mature. Cat. No. 32399, U.S.N.M., contains the typeand
another specimenfrom Todos
Santos Bay,Lower
California.The
type has eight post-nuclear whorlsand
measures:Length
4.5mm.,
diameter2.1
mm. The
thirdspecimen (Cat. No. 16196,U.S.N.M.)comes from Cape San
Lucas,Lower
California.CERITHIOPSIS (CERITHIOPSIS) AUREA,newspecies.
Plate40,fig.1.
Shell elongate-conic, bright, light chestnut brown, excepting the extreme apex,
which
iswax
yellow. Early nuclear whorls decol- lated; the last one only remainsand
is wellrounded and
smooth.Post-nuclear whorls well rounded, appressed at the