iSSs.]
Ridgway
on Helminthophila leucobronchialis.3S9
Mr. Ridway
concurs,and
says "it differsfrom
T.vinaceo-rufa
in
much deeper
general coloration, thevery different colorof thecrissum
(rufous instead of purplish vinaceous),and much
larger size, especially thebill."T.
ruficauda seems
tohe
a rare species, as there isno
speci-men
intheNational Museum
or in thatof theAmerican Museum
of
Natural
History,New York.
It is described asbeing
quitedark above ("brunneo-ardesiaca"), with
thehind neck plumbe- ous
;below and on
theneck
rosywine
color; a spotunder
the eyeand
a linebehind
theeye,violet-black,bordered with white
; sides of theneck
golden-violet;wing-coverts "
ardesiacis "; quills blackish, tail short, outer tail-feathers "fusco-ardesiacis,"
with
a black spot across themiddle,
theend
broadly rufous.New York,
September15, 1885.HELMINTHOPHILA LEUCOBRONCHIALIS.
BY ROBERT RIDGWAY.
Without having any new evidence
to presentconcerning
thisperplexing
bird intheway
of additionalspecimens,
Iwould
like to offer forconsiderationahypothesiswhich
itseems
tome
affords amore
plausible explanation of certain aspectsof
the casethan
thatwhich
suggestshybridism between simply H. chrysoptera and H. pinns
as sufficient toaccount
for the variable 'aberrant' serieswhich has been
referred toU.
leucobronchialis, togetherwith
thosewhich
represent the so-calledH. lawrcncei
'.It has
been
stated thatH.
Icucobroiichialisand H. lawrencei
possessno important
original characters.This
is certainly true of the latter,which
isvery obviouslyahybrid
of theabove-named
species;but
it isnot true of the former,which
initspure white
throat, invery
strikingcontradistinction tothedeep gray
orblack throatofone and
brightyellow
throat of the other alleged parent, certainlydoes
possessone
veryimportant
original character,which
it is impossible toconceive
can
be derivedfrom
the crossing of thetwo
species in question.H)0 Ridgway
on Helminthophilaleucobronchialis. [OctoberA supposed
strong pointwhich
h;isbeen urged
against the validityof //. leucobronchialis&% adistinct species, isthecircum-
stance that typicalspecimens
constitute a smallproportion
of allthose obtained,
and
the suspiciouscircumstance
thateach
in- dividual of the aberrant majority inclined, inone
respect or another,toward
//. f^iiius.Only
seven of thetwenty-two exam-
ples
which have been
recorded (including those of'//. lawrencei,*which must
be considered in this connection), oi" not quite one- third, are true //. leucobronchialis; butwhen,
instead of takingtwo elements
into consideration (i. e., //.piiutsand
//.chrysop-
/</</)
we add
a third (//.leucobronchialis
itself), thedispro- portionbecomes
far less significant.In
no
otherway than by assuming
that //.leucobronchialis
\§ a distinct species,which
itself hybridizeswith
its allies,can we
account
tor the origin of thepuzzling
seriesofspecimens which have
so prejudiced the claims of the birdunder
consideration to specificrank
;and
1 feelconvinced
thatMr. Brewster expressed
the exact truthwhen
he stated {cf. Nuttall Bulletin, 111, p. 99) that"the
validity of this distinctly characterized speciesmust now
beregarded
as established,"and
I feel equally satisfied that the hypothesis ofhybridism which
hesubsequently advanced
tails to settle the case as affecting //.
leucobronchialis
itself.however
well itmay answer
for the 'aberrant'specimens which have been wrongly
referred to it.Regarding
the very closeresemblance between
certain speci-mensof
//. /<,//<t>/>/-o//c///<r//.s\Mr. Brewster observes
:"Indeed,
itwould
be difficult toselect three individuals ofany
specieswhich
vary so little interse." Ihave myself
seenonly
thetype and
theVirgina specimen
in the NationalMuseum
collection ; but Ican
affirm that thetwo
are as nearly counterparts ofone
another, so far as details ofplumage
areconcerned,
asany two specimens,
ofany
species, thathave
evercome under my
observation.'The following classification of all the
specimens*
hitherto recorded ofboth
//.leucobronchialis
(in its widest sense)and
•//. lawrencei,' expresses
my views
as to their natureand
origin.The arrangement
is of course purely hypothetical, but at thesame
timeadmits
of amuch more
satisfactory solution of theproblem which Mr. Brewster
has so carefully discussed than*
A
fewmaypossiblyhave beenoverlooked.18S5.]
Ridgway
on Helminthophilaleucobrorichialis.^61
the theory
which supposes
the entire scries to be hybrids ofH. chrys
ipteraand
//.plnus
alone, <>r of theirprogeny
interse.I.
Typical Specimens
of //• leucobrorichialis.Characters. —
Throat and cheeks pure white; postocular black or duskystreak verynarrow, not involving the auriculars; breast white, or but veryfaintlytinged vvitli yellow; wing-patch, or bands, yellow.(1)
The
type,an adult$
from Newtonville, Mass.,May
iS, 1870. | Cf.Brewster,Bull. Null. Orn. Club, I, p. 1, pi. i.)
(2)
An
adult$
shot byChristopher I).Wood,
nearClifton, Delware Co., Penn.,May
12, 1S77.(C/
Trotter, Bull. Nutt.Orn. Club, [I,pp. 79, So.)(3)
An
adidt$
in the collection ofthe PhiladelphiaAcademy
of Natural Sciences,supposedto have hern shot byj. G. Bell atRockland, N.Y.. in thespring of 1832. (Cf. Trotter,Bull.Nutt. Orn. Club, 111, p. 44; IV, p. 59.)(4)
An
adult$
shot atWauregan,
Conn.,May
25, 1875, by Charles M. Carpenter. {Cf. Brewster, Bull, Nutt. Orn.Club, 111. p. 99.|(5)
An
adultJ
obtainedatSull'ield,Conn.,July3,1875,byK.1.Shores,{Cf
Brewster,Bull. Nutt. Orn. Club, III, p. 199.)(6)
An
adultJ
incollectionofWilliamsCollege,Williamstown,Mass., obtainedatHudson,
Mass.,inMay
or June, 1858, bySamuel
Jillson.{Cf
Purdie. Bull. Nutt. Orn. Club. IV, p. [84.)
(7)
An
adultJ
shot, near Arlington, Va.,May
15, 1S85, by William Palmer, and bv him presented tothe NationalMuseum (museum
register No. 104,684). {Cf. Palmer, Auk,II, p. 304.)II.
Presumed
hybrids or- //. leucobronchialisand
//. pinus, <>uof THE FORMER
WilliTHE
(RossBETWEEN
//. pitlUSAND
• //. chrysoftera
(=
'H. latvrexcei Merrick.').Characters —
Throatwhite, sometimes faintly tinged with yellow;breastyellow; grayof upperpartstinged with olive-green.
a. Wing-band, or fatch,yellow.
(1)
An
adult$
taken at Portland,Conn., by Williarn-W.Coe,May22,
1S75. Ilas"a broad bandor blotch"of yellowon the breast, while"Ihe wholedorsal plumage, from thecrown, is faintly washed with the
same
tint."{Cf. Purdie, Bull. Nutt. Orn. Club, IV, p. 184.)
(2)
An
adult cf takenatSaybrook,Conn.,May
30,1879,hyJ-N
-Clark.Has
"a patch of bright yellow across the breast from the bendof the wings." {Cf. Purdie, 1. c.)(3)
Ayoungbird
(sexunknown)
taken by Dr. E.A.Mearns
at High- land Falls, N. V*.,July7, 1879 'Coll.W.
Brewster,No. 4,668). Specimennumber
4 under//, lawrencei(No. 4,667, Coll.W.
Brewster),said to362 Ridgway
on Helminthophilaleucobronchialis. [Octoberbe the parent of this specimen.
{Cf
Brewster, Bull. Nutt. Orn. Club,VI, p. j J 1.)
(4)
An
adult(?),sexnotstated, obtainedatSingSing, N.Y.,August
3, 1SS1,byDr.A. K. Fisher. Resembles specimen No. 8 (No.605, Coll.
A. K. Fisher), buthas the wing-bands "yellow, not white."
{Cf
Fisher,Bull.Nutt. Orn. Club, VI, p. 245.)
(5)
An
adult$
(?) obtained inOttawa
Co., Mich..May
25, 1879, byW.
A.Gunn. Whole
breastbrightgamboge
yellow,withasharplydefined, truncated anterior outline, but posteriorlyextending farthest along themedian
line, to the upperpart of theabdomen;
cheeks and chin tinged with lightyellow;upperpartwashed
with olive-green.Type
of"//.gunni
Gibbs." {Cf. Gibbs, 'Daily Democrat,' a newspaper ofGrand
Rapids, Mich..June 1, 1S79.—
Purdie,Bull.Nutt.Orn.Club. IV,p. 1S5.—
Ridgw.ib., p. 233.)
(6)
An
adult$
taken atSingSing, N. Y.,August
3, 1SS1, by Dr. A.K. Fisher (collector's No. 1235).
Has
"a broad patch of pale yellow on the breast."{Cf
Brewster, Bull. Nutt. Orn.Club,VI
p.219.)(7)
An
adult $ takenatSingSing,N.Y., July24, 18S1, byDr. A. K.Fisher (collector's No. 1210). Entirely pale greenish yellow beneath;
the backissimilar tothat ofpinusbut the napeisdecidedly ashy, and the wing-bands as clearlyyellowas inchrysoptera; the dusky
brown
eye-stripeisrestricted totheloresandpost-orbitalspot."
{Cf
Brewster,Bull. Nutt.Orn.Club, VI,p. 220.)
b. Wing-bands-white.
(8)
An
adult$
collected atSingSing, N. Y.,August
24, 1S79,'5yDr.A. K. Fisher(collector'sNo. 605). "Exhibitsa faint
wash
oflemon
yellowon
the throat, while a broad space across the breast is deepgamboge
yellow,and the wing-bandsare pure white."{Cf
Brewster, Bull. Nutt.Orn.Club, VI, p. 219.) »
(9)
An
adult $ (?) collected atNyack, N.J.,in
May,
1S7S,byEugene
P. Bicknell;No. 2620, Coll.
W.
Brewster."Has
thechin decidedly yellow;thethroat, cheeks,andasmall space on the
abdomen
white; the remain- der of the lower partsgamboge
yellow; thewing-bandswhite; thenape ashy tingedwith green;the occiput, back,andwingsaspurely green as inH.
pinus."{Cf
Brewster/.c.)c. Wing-bands mixedwhite
and
yellow.(10)
An
adult$
killed atNew
Haven, Conn.,May
19, 1SS5, D}'Harry W.
Flint. It"shows
a slightsuffusionofyellowunder each eye and on thechin, aswellas a light bar of thesame
color across the breast ....the wing-barsarevery
much
restricted,and thewhite is tinged with yel- low, andthere isaspotofthesame
colorontheback.{Cf
Sage,Auk, II,P- 304- )
'S35-J
Ridgway
onJuncocinereusand
itsGeographicalRaces. '\6'lIII.
Presumed hybrids of H.
leucobronchialiswith
//. chrysoptera.Characters. — Entirely white beneath (except on sides), as in //
leucobronchialis,but with black auriculars of
H.
chrysopti-ra.(i)
An
adult $ (?) obtainedatSingSing, N. Y.,July24, 1SS1,byDr.A. K. Fisher(collector'sNo. 1208). Cf.Brewster, Bull. Nutt.
On.
Club, VI,p. 219.)(2)
An
adult $ (?) obtainedatSingSing, N.Y., July24, 1SS1,by Dr.A. K. Fisher.
{Cf
Fisher, Bull.Nutt. Orn. Club, VI, p. 245.)IV.
Presumed hybirds between
H.chrysopteraand H.
pinus(=
'//. lazvrencciHerrick').Characters
: Blackorgraythroatandauricularsof chrysoptera. with restof headandthe lowerpartsyellow,as inpinus; upperpartsolive-green asinpinus.Wing-bands
yellowor white.a. Wing-bands-white.
(1)
The
type, an adult J, obtained at Passaic, N.J.,May,
1S74, by HaroldHerrick.{Cf
Herrick,Pr. Phil.Ac. Nat. Sci. 1874,p. 220, pi. 15.) (2)An
adult$
fromHoboken,
N.J., taken in the spring of 1876.{Cf
Herrick,Bull. Nutt.Orn. Club, II,p. 19.)b. Wing-bandsyellozv.
(3)
An
adult $ taken atHighland Falls, N. Y.,July7, 1879, by Dr.Edgar
A.Mearns
(No.4667, Coll.W.
Brewster). This specimenpresents nearlythecharacters that wouldbelookedfor inthefemale oflaivrencei;the throatand cheek-markings are those ofchrysoptera (female), while theremainderof the