DIAGNOSIS.-Face (prefrons) generally projecting and broadly arched transversely; facial stiffness, especially along the oral margin; at least 1 pair of lateroclinate fronto-orbital bristles; pulvilli well developed;. One pair of large lateroclinate fronto-orbital bristles 7 Two pairs of large lateroclinate fronto-orbital bristles 9 7. DIAGNOSIS.-Specimens of Austrocoenia do not closely resemble those of any other genus of the tribe Scatellini and are easily distinguished by the following combination of character states: Setae generally reduced ; mesophrons uniformly setose, completely pollinose, dull; 2 pairs of large lateroclinate fronto-orbital bristles; inner and outer vertical bristles both well developed; paravertical bristles either reduced or absent; antenna short, third segment equal to length of second segment seen dorsally; Arista short, slightly longer than length of third segment, basal two-thirds thickened; antennal fovea deeply impressed, face long, distinctly projecting forward;.
I have tentatively placed it near the Paracoenia-Calocoenia line because adults have five pairs of dorsocentral bristles. DIAGNOSIS.—Specimens of Calocoenia closely resemble those of Paracoenia, but may be distinguished from the latter and other genera of Scatellini by the following combination of character states: Generally dark in color; mesophron subglabrous to glabrous, inconspicuously setulose; 2 pairs of large frontoclinate fronto-orbital hairs; internal and external vertical hairs well developed; paravertical seta well developed, about half the length of vertical cross hairs; arista pectinate, rays with dorsal branching subequal to half width of third antennal segment; face with interfoveal carina, but dorsal crease not as distinct as in specimens of Paracoenia; eye sub-spherical to sub-elliptical, oriented at a slight oblique angle to the general plane of the head; acrostical setae uniformly small, hair-like, arranged in 2 rows extending to base of scutellum; 5 pairs of dorsocentral bristles (2 + 3); humeral callus with 2-3 larger, hair-like setae; supra-alar hairs well developed, subequal to anterolateral postal hairs; scutellum setose dorsally, with 2 pairs of lateral hairs; with dorsum subplanes; costal margin with equal spine-shaped setae distinct from remaining setae; radial running vein bare; male fifth abdominal tergum slightly wider than long; terga generally unicolorous; surstylus as angular processes extending from ventral margin of epandrium, attached medially; aedeagus well developed, with acute tip; Female ventral receptacle with small operculum, subtrapezoid conformation in lateral view, extending process much longer than width of operculum. DIAGNOSIS.—Specimens of Paracoenia are similar to those of Coenia and Notiocoenia, but may be distinguished from either by the following combination of character states: Generally dark-colored, subgloss to glossy dorsum; mesophrons subglabrous to shining, in marked contrast to paler paraphrons; 2 pairs of large frontoclinate fronto-orbital hairs; internal and external vertical hairs well developed; paravertical seta well developed, about half the length of vertical transverse punctures; arista with pectinate rays, dorsally branched; eye subspherical to subelliptical, oriented at an oblique angle to general plane of head, acrostical setae uniformly small, hair-like without rows, 4-5 irregular rows; 4-5 pairs or dorsocentral punctures J- 2); humeral callus with 2-3 larger setae; supra-alar hairs variously developed; scutellum setose dorsally, with 2 pairs of lateral hairs; arm developed normally; radial running vein with setae inserted in anterior part of transverse septum; fifth abdominal tergum longer than third or fourth; surstylus visible as ventral lobelike extension separated by a triangular median process; aedeagus well developed, with acute tip; female ventral receptacle with helmet-shaped operculum, extending short process.
Chaetotaxy of the thorax as follows: Acrostichal bristles of equal size, without series, 4-6 irregular rows; 4-5 pairs of dorsocentral bristles, the posterior pair, laterally displaced from the others; 2-4 pairs of humeral bristles; 1 pair of interalar bristles; 5 pairs of dorsocentral setae (2 -f- 3); supra-alar bristles well developed, less long than the length of the anterolateral postal bristles; sternopleural bristles well developed; wing generally hyaline to very slightly waterlogged; femora more swollen, almost twice as wide.
Paracoenia (Paracoenia) wirthi Mathis
Both supra-alar and sternopleural bristles are absent or greatly reduced; middle femur of male without row of closely spaced setae along posteroventral surface; apically 4 tarsomeres explanatory and short; pulvilli are absent; third antenna segment short, less than the length of the second segment; DIAGNOSIS. General body color dark black-brown to black; The setting is generally weakly developed, especially on the abdomen; frowns extending forward, almost to the face; antenna short, third segment as wide as long; length of dorsal rays of arista short, approximately equal to half the width of the third antennal segment; face short, not equal to the length of longer facial brushes along the oral margin; face with much longer hair along the edge, the rest of the face at most setulose; antennal fovea well developed, deeply impressed; 4-5 pairs of dorsocentral setae f-3); supra-alar bristles absent or hair-like, sterno-. Frown usually monochromatic, uniform, dull, dusted, without distinctly colored mesophrons; 2 pairs of large lateroclinate fronto-orbital bristles and 1 pair of much smaller proclinate bristles anterior to larger bristles; paravertical bristles small, inconspicuous, not half the length of vertical bristles; eye shape subspherical, slightly higher than wide; gena short, eye-to-cheek ratio 1:0.20 or less; acrostic brushes arranged in 2 rows; 4 larger dorsocentral setae (1 + 3), rarely with a fifth (2 + 3) setae; 1 pair of interalar bristles inserted just behind the transverse suture; wing normally developed, hyaline to slightly saturated; male terminalia as in species group diagnoses.
Chaetotaxia of the thorax as follows: Acrostical bristles arranged in 2 rows, larger bristles usually anterior to the transverse suture; 4-5 pairs of large dorso-central bristles in front, 1-2 pairs of smaller bristles, several smaller bristles in front or larger bristles; 1 pair of interalar bristles; 1 pair of interalar bristles inserted just behind the transverse suture; supra-alar setae variable; postal bristles well developed; 2 pairs of side bristles; 2 pairs of notopleural setae, inserted near each ventral corner; 1 pair of mesopleural setae, inserted near the center of the posterior margin, numerous smaller setae, especially towards the dorsal and posterior margins; 1 pair of larger sternopleural bristles and 1-2 smaller bristles around larger bristles; middle coxa with. The outer vertical bristles are less similar to the inner vertical bristles; supra-alar bristles absent or greatly reduced; arista at most macropubesccnt; humeral callus setosis; wing appears dull, slightly drenched, 2 white spots on either side of the posterior transverse vein (the pollinosa group) 5. 1 pair of smaller proclinate fronto-orbital setae inserted anterior to larger lateroclinate setae, about one-third larger setae; arista pectinate with dorsal branches along basal two-thirds, longest rays less than half the width of the third antennal segment;
Notiocoenia acutella, new species
Although an occasional specimen was encountered in other habitats, most were collected at one site near Aguas Calientes, Parque National Puyehue. A densely vegetated, steep embankment ran its length to the south; the path marked its northern boundary. A dense overhang of brush from the adjacent bank shaded the area at all times and probably provided most of the organic litter.
Most samples were collected by sweeping under the overhang and just over the muddy seepage area. The holotype is in the National Museum of Natural History, Smithsonian Institution, Washington, D.C., type no. The holotype specimen is double mounted (minute needle) and is in good condition. Figure 18).—This species is largely sympatric with N .
ETYMOLOGY.—The species epithet, acutella, is derived from the Latin acutus ("sharp"), which refers to the sharp-pointed aedeagus.
Notiocoenia paniculata, new species
Wing (Figure 68) mostly hyaline, shiny; slight anterior infusion and especially around the posterior cross vein; costal vein index on average 1:0.14; Vein index Mi+2 on average 1:0.66. The legs are pale orange, often with a little infusion; tarsi becoming darker, black, towards apical tarsomere. Abdomen (Figure 22): Coloration almost monochromatic, brownish black with some bluish tints, subgloss to glossy.
The holotype, allotype, and paratypes from the type locality are in the National Museum of Natural History, Smithsonian Institution, Washington, D.C., type 76068. The holotype specimen is double mounted (minute mount) and is in good condition. Fig. 28).—This species occurs along the western slope of the Andes, mostly in the foothills, between 24° and 41° S. latitude. FIGURES 23-27.—Notiocoenia paniculata: 23, epandrium and cerci, posterior side; 24, male terminals, lateral view; 25, internal male genitalia, lateral view; 26, female ventral receptacle, anterior; 27, female ventral receptacle, lateral view.
ETYMOLOGY.-The type ephite, paniculata, is the diminutive form of pannus ("ear of millet"), referring to the tuft of setae on the fourth sternum of males. DIAGNOSIS. - Specimens of this species group are similar to those of the paniculata group, but can be distinguished by the following combination of character states: Thrones narrower, width at level of median ocellus about 2 times distance between ptilinal suture and median ocellus. ; arrangement of ocelli clearly forming isocellular triangle, distance between posterior pair much greater than between posterior and median eyes; outer vertical hairs well developed, subequal to inner vertical hairs; smaller fronto-orbital setae much less than one fourth latero- larger.
Notiocoenia pollinosa, new species FIGURES 29-35, 69
ETYMOLOGY.—The species epithet, pollinosa, is derived from the Latin pollen ("fine flour or flour"), referring to the pollinous coating of the fungi on specimens of this species. Becker (1905b) seemed to prefer Scatophila mainly because the costal vein appeared to terminate close to the third vein (R 4+ 5), which is one of the main identifying character states of that genus. What replacement event resulted in the speciation of the two known species is difficult to determine.
TYPE MATERIAL. I have not studied the type or type series of the senior or junior synonyms. Length of Arista: In general, the arista is as long as or longer than the combined length of the antenna segments. Surstyli: Paired surstyli are absent or indistinguishable from the ventral edge of the epan-.
In males of these two genera, the apodeme has lost its attachment (articulation) with the hypandrium and is only loosely attached to the base of the aedeagus. My study of the subgenera of Scatella is incomplete, as I have not examined specimens of all described subgenera in detail. Vestiture and color of frons: In general, the mesophrons is differentiated from the rest of the frons by its clothing.
In specimens of Synhoplos the wings are micropterous, no longer than the basitarsus of the hind leg. Throughout most of the family, the apical pair of shoulder hairs project along the same plane as the body. Strength of setae of male midfemur: In general, there is a row of short setae along the posteroventral margin of midfemur in males of Ephydrinae.
The monophyletic relationship and sister group of the Coenia-Notiocoenia lineage (Figure 76) is indicated by character state number nine as mentioned earlier. In specimens of the pollinosa group, such as specimens of Scatella, the supra-alar is reduced. As in Scatella and related genera, the wing of pollinosa-group specimens is infuscate and has white spots (see species-group description for details).
In specimens of the subgenus Paracoenia, the acrostichal appendages are in four to six unarranged irregular rows. Males of the subgenus Paracoenia are unique in having a dense row of longer setae.
