Vol. XIXT ^^^y,,sT-EV.,
An
UndescribedForm
of Black Duck. I83
1Q02 J
Near
the anchored steamer weremany
Terns {Sterna hirundo),
butI could not tellwhetheror not they liveon the cliff.
The
lengthof the cliffinhabitedby
thebirds isabout one mile.By
counting the birds on a section of thecliff,we
estimated thenumber
of birds at not less than 60, 000.The
photographs (PI.VII) were taken undervery unfavorable circumstances, owingto
foggy weather, but
may
helpto givesome
idea oftheabundance
of the birds.
AN UNDESCRIBED FORM OF THE BLACK DUCK
{AJVAS OBSCURA).
BY
WILLIAM
BREWSTER.Itis amatterof
common
beliefamong
ourmore
intelligent and observingwild-fowlgunnersthattwokindsofBlackDucks
arefoundin
New
England, and certain of the charactersby which they are thought to be separable have been mentioned, as well as briefly discussed, by writers on ornithology or field sports.To
one, a comparatively small, darkracehaving a dusky or olivaceous billand brownish legs, all our local or breeding birds are supposed to belong; the other, a larger, lighter-coloredformwith clear yel- lowbill
and
bright red legs, isknown
to occur only duringmigra- tion or in winter. I have often been struckby these and certain other differenceswhich willbe mentioned later,and
for the pur- poseof testing theirvalueand significanceIhave broughttogether, with the kindassistance of several ofmy
friends, a large series of specimens mostof which wereobtainedinNew England
inautumn,winter or early spring.
Among them
are a dozen ormore
col-lected in late August
and
early September atLake Umbagog,
which almost certainly represent the formresident insummer
throughoutNew England
although I havenone
from anylocalitysouth of the Gulf of St.
Lawrence
which weretaken attheheightof the breeding season.
Some
ofmy
specimens were weighed beforebeing skinnedand many
ofthem
areaccompanied
bynotes184
Brewster, Afi UndescribedForm
of Black Duck. [AukApr.
relating to the original coloringof their bills,legs andfeet.
As
the
more
brilliant tintsof these 'soft parts' fade soon after death, andeventually nearl)' or quite disappear, they are not available incomparisons of dried skins unless recorded by the collectorsoon
after his birds arekilled.
A
carefulstudyof this material has convincedme
thatthelarge, red-legged bird differs sufficientlyfrom true ohsciira to stand as a distinct subspecies,whichmay
be brieflycharacterized asfollows:Anas obscura
rubripes,new
subspecies.Black Duck.
Red-legged
Suhspecific characters.
—
Similarto^. obscura but larger; the feathers ofthepileumconspicuously edged with grayish or fulvous; the dark markings onthe foreneckandthesides of the head coarser, blackerand moresharply defined;theentire throat usually streaked orspottedwith blackish; thetarsiandtoes bright red; the billyellow.Tj/Je,No.30252,
$
ad. Collection ofWilliam Brewster,LakeUnibagog (New Hampshireshore), October8,1889;W.
Brewster.Habitat.
—
Occurring during migration or in winter on or near the Atlantic Coast from Newfoundland to Virginia (Cobbs Island); in the interior as far to the south and westasArkansas.Summer
range not definitelyknown
butbreeding specimens examined from NorthernLabra- dor,JamesBayandthewest shoreofHudsonBay.Measurements.
Vol. XIX
1902 I
Brewster,
Ah
Utide.scribcdForm
of Black Duck. l8'^chrome
to canary or sulphur yellow, the legs and toesbright red, var}'mg from light scarlet to deep orange, the dark feathers of thepileum and nape conspicuouslymargined with grayor fulvous, and the throat (as well as sometimes the chin, also) profusely spotted orstreaked with blackish. All the dark markings on the cheeks, throat and neck are broader, blacker andmore
sharply definedthan in true ohscura andthey often take theformofcoarse, rounded spots which are seldom if ever presenton the head or neck of the smallerbird.Intypical examples ofobscura the bill is greenish black, dusky
olive, or olive green, the legs are olivaceous
brown
with, at most,only a tinge ofreddish, the pileum and nape nearly orquite uniformly dark, the throat and chin immaculate, the markings on the neck andsides of the head fine, linear, and duskyrather than blackish. In respect to these characteristics obscura does notseem
to vary with age or season formy
series includes severalyoung
notsufficiently large and fully feathered to have been able to fly which are colored andmarked
precisely like specimens killed in late autumn, while breeding birds are distinguishable from the latter onlyby themore worn and
faded appearance of their plumage.The
males of both forms, however, are almost invariably larger than the females aswell asmore
richly coloredand
heavily marked, especially on the head and neck; a fact which should be bornecarefully inmind when
specimens of the two are compared.Both races are evidentlysubject to a
good
deal of individual or geographical variation Avhich tendsto connectthem
by a series of intergrading specimens.Thus
I have small birds with grayish crowns orstreaked throatsand
even one or two which, in life,apparently
had
yellow bills andred legs, while several ofthelarge ones have plain black crowns orimmaculate throats. I haveyet to see a specimen of obscura,however,whichpossesses thecoarse, rounded, deepblack spots that are usuallypresent in greater or lessnumbers
on the neck, as well as often on the throat, of rubripes.The
existence of a small percentage of non-typical examples, likethosejust mentioned, does not necessarilyaffect the diagnos- tic value of the characters to which I have called attention.1
86
Brewster,An
UndescribedForm
of Black Duck. [a"''Indeed it would be possible to contend that these aberrant or intermediate specimens are really hybrids,for in the series before
me
theydo not exceed innumber
the birds (no less than nine) whichshow
unmistakable traces ofan infusion ofMallard
blood.Since two species so obviously distinct as are the Mallard and Black
Duck
are connectedby
intergrades k?iowti to be hybrids,why
shouldwe
notassume that the scarcelymore numerous
inter- grades between the red-leggedand
brown-legged BlackDucks
are also hybrids?Not
thatIam
disposedtoseriouslypressthisargu-ment
for,however
plausible itmay
seem,my
present impressionisthat the forms of the Black
Duck
here considered are only sub- specifically distinct.There
can be no reasonable doubtthat the smaller of the twoistheoriginal
Anas
obscura. Thisname
has remainedunchanged
inform
and
uncoupled with anys3monym
ever since itwas
insti- tuted,more
than onehundred
years ago, byGmelin
(Syst. Nat.I,partii, 1788, 541), vvho based it on the
"Dusky Duck"
of Pen- nant. This isdescribed (Arct. Zool., II, 564) ascoming
"fromthe province ofNew York" and
having"a
long and narrow duskybill, tinged with blue: chin white: neck pale brown, streaked
downwards
withdusky lines." Pennant adds that the legsin one of hisbirdswere"dusky,inanotheryellow"; butasthespecimens which he examined were evidently dried skins (in the BlasiusMuseum)
this statement, as well as that relatingtothecolor ofthebill, loses
much
of itsapparent importance.At Lake Umbagog,
wherethe BlackDuck
breedsrather plenti- fully, I have not cared to incur theodium
of breaking thegame
laws
and
the reproaches ofmy own
conscience by killing birds which were sittingon theireggs or in chargeof broods of tender young, but I have shot afew specimens in late August and ver}?^many
during themonth
ofSeptember.Among
theseIhave found onlyone example of rubripes, a nearly typical female taken onSeptember
28, 1S89.With
thissingleexception Ihave nevermet
with the red-legged form at thislocality before October8.Soon
afterthat date it
becomes common,
remaining until the waters of the lake are closedby
ice.In Massachusetts, also, the locally bred birds or earlymigrants from the north, which
we
kill duringSeptember and
the first halfVol.XIX
iqo2 1 B-RM.\vs.T)E.K,
An
UtidescribedForm
of Black Duck. 187
of October, are, as far as I have observed, invariably obscura.
Most
of the representatives of this race evidently pass further southward to spend the winter, but Ihavethreetypicalspecimens which were shoton ourseacoast (at Ipswich andChathamj
during the latter half of February, 1901.Until very recently I had supposedthat the Black
Ducks
which breed about the Gulf of St.Lawrence
and to the northward along the eastern coast of Labradorwould
prove to belong to the red- legged form, but Mr. C. F. Batchelderhasshown me
seven speci-mens
(all but one adult) which were collected forhim
inNew-
foundlandinJune andJulyandwhich, althoughslightly largerthan our New-England-breedingbirds, are preciselysimilar tothe latter in color and markings.A
female in Mr. O. Bang's collection taken in the Straits of Belle Isleon April 25, 1900, must also be referred to obscura. Another, belonging to Mr. J. D. Sornborger, which, with her brood of ducklings,wascaptured on July 8, 1896, atOkak,
on the northeastern coast of Labrador,isintermediatein certain respects between ^/w/<;rrt;and
rnbripes, but on the whole perhaps nearer the former.To
the red-legged race I can unhesitatingly refer only four of thebreeding BlackDucks
which I have examined.One
of these (a female)was
taken by Mr. L.M.
Turner on July i, 1884, atUngava,
Northern Labrador; another (unsexed andwithout date)by
Mr.John McKensie
atMoose
Factory onJames Bay
; a third (representedby
only thehead and
wing and bearing no sexmark
but evidently a female, for itwas
"with young")by
Mr. C.Drexler, on June 19, i860, at
Cape Hope,
Severn River; the fourth (a maie) by Mr. E. A. Preble, on July 28, 1900, at FortChurchill;—
the two localities lastnamed
being on the western shores ofHudson
Bay. Mr. Preble'sspecimen isinthecollection of the Biological Survey while the others belong to the NationalMuseum.
All four of these birds are in poor condition for comparison.Two
ofthem
were moulting, and theplumage
of the other two isworn and
faded, while that of theMoose
Factory skin is also strongly tinged with rusty chestnut—
amere
super- ficial stain, apparently. Nevertheless theyshow
satisfactorily mostof the essential characters of rnbripes. In respect to sizeand
the character and distribution of the black markings onthe1
88
Brewster,An
U7idescribedForm
of Black Duck. LApr.head
andneck
they are quite typical of that form.The
original coloringof thesoft partsis not noted on anyofthe labels,butthat of the legs in the three specimens which still possess these appendageswas
apparently bright red.As would
be expected, the light edging on the feathers of the pileum ismuch
narrower and less conspicuous than in birds infresh winterplumage.From
this evidence it seemsreasonably safe toassume
thatthe breeding rangeof true ohscura extends, coastwise, to the northand east atleastas far asNewfoundland and
SouthernLabrador and
thatthroughoutthis maritime belt, as well as in
New England and
to the southward, rubripes occurs only during migration or in winter.
The summer
distribution of the latter remains to be definitely ascertained, but besides occupyingthe shores ofHud-
son
Bay and
those of northern Labrador it probably frequentsmore
or less of thevast interior region lying between the points just mentionedand
the St.Lawrence
River.The
frequency of its occurrence in lateautumn
atLake Umbagog
indicates thatagood many
of the birds which breed aboutHudson Bay
or tothe southward take the shortest possibleroute to theirwinter quarters onthe coast ofNew
England. Others,no
doubt,move
directly southward forthere isa typical red-legged birdin theMuseum
of Comparative Zoology whichwas
taken in Mississippi County, Arkansas, on Nov. 5, 1887.^Those
which pass theirsummers
in northern Labrador probablyfollowthe Atlantic coast line during migration for Mr. Batchelder has a specimen of rubripes whichwas
shotat Custlett, Newfoundland, onNovember
6, 1890.It is interesting to note that in respectto one of the
more
es- sential of its distinguishing characters—
viz.,theimmaculatebuffythroat
—
themore
southern ofthe forms just consideredshows
aslightbut significantapproachto theBlack
Duck
of Florida{Anas fiilviguld) which has not only the entire throat, but also the jugu-lum
and the greaterpart of the cheeks, entirely free from mark- ings.'It is probablethata large proportionof the birdswhich occurinautumn or winterinthe Mississippi ValleyandabouttheGreatLakesbelongto theform rubripes but the only Black Duck of any kind that Ihave seen fromthis regionistheoneabove mentioned.