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بررسی اثر متقابل نماتد ریشه گرهی Meloidogyne javanica و قارچ عامل پژمردگی ورتیسیلیومی Verticillium dahliae روی نهال‌های برخی از ارقام زیتون

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REFERENCES

1. Afshari Azad, H. & Khozeini, F. (2000). Isolated fungi from seedlings of olive in Iran. In: Proceedings of the 14th Iranian Plant Protection Congress, Esfehan University, Isfehan, Iran, Vol II, p.333.

2. Afshari Azad, H. & Alizadeh, P. (2004). Isolation of Verticillium dahliae from olive trees in Kohkiloei

& Boyerahmad, Iran. In: Proceedings of the 16th Iranian Plant Protection Congress, Tabriz University, Tabriz, Iran, Vol II, p.348. (In Farsi).

3. Akhiani, A., Mojtahed, H. & Naderi, A. (1984). Species and physiological races of root-knot nematodes in Iran. Iranian Journal of Plant Pathology, 20, 1-4. (In Farsi).

4. Al-Ahmad, M. A. (1993). The solar chamber: an innovative technique for controlling Verticillium wilt of olive. Bulletin OEPP. (Organisation Europenne et Mediterraneenne pour la Protection de Plantes), 23, 531-532.

5. Barthels, N., Lee, F. M., Lap, J. K., Goddijn, O. J. M., Karimi, M., Puzio, P., Grundler, F. M. W. & Ohi, S. A. (1997). Regulatory sequences of Arabidopsis drive reporter gene expression in nematode feeding structures. Plant Cell, 9, 2119- 2134.

6. Bellini, E. (2002). Miglioramento Genetico. In Arsia (Ed.), La Toscana Nella Storia Dell'olivo e Dell'olio. (pp. 229-260). Florence, Italy.

7. Bhat, R. G. & Subbarao, K. V. (1999). Host range specificity in Verticillium dahliae.Phytopathology, 89, 1218-1225.

8. Bowers, J. H., Nameth, S. T. & Rowe, R. C. (1996). Infection and colonization of potato roots by Verticillium dahliae as affected by Pratylenchus penetrans and P.crenatus.Phytopathology, 86, 614- 621.

9. Caballero, J. M., Perez-Hernandez, J., Blanco-Lopez, M. A. & Jimenez-Diaz, R. M. (1980). Olive, a new host of Verticillum dahliae in Spain. In: Proceedings of the 5th Congress of the Mediterranean Phytopathological Union, Patras, Greece, P.50.

10. De Grisse, A. (1969). Redescription ou modifications de quelques techniques utilisees dans letude des nematodes phytoparasitaires. Mededelingen Rijksfaculteit der Landbouwwetenschappen, 34, 351-369.

11. Devay J. E., Garber, L. L. & Butterfield, E. J. (1973). Characteristics and concentration of propagules of Verticillium dahliae in air-dried field soils in relation to the prevalence of Verticillium wilt in cotton.

Phytopathology, 64,22-29.

12. Dhingra, O. D. & Sinclair, J. B. (1986). Basic plant pathology methods. C. R. C. Press. Inc. 355 pp.

13. Eisenback, J. & Triantaphyllou, H. H. (1991). Root-knot nematodes: Meloidogyne species and races. In W. R. Nickle, (Ed). Manual of agriculture nematology. (pp. 191-274). Marcel Dekker. New York.

14. Erwin, D. C., Tsai, S. D. & Khan, R. A. (1976). Reduction of the severity of verticillium wilt of cotton by the growth retardant, tributyl (5-chloro-2-thienyl methyl) phosphonium chloride. Phytopathology, 66, 106-110.

15. Fattah, F. & Webster, J. M. (1983). Ultrastructural changes caused by Fusarium oxysporum f.sp.

lycopersici in Meloidogyne javanica induced giant cells in Fusarium: resistance and susceptible tomato cultivars. Journal of Nematology, 15, 128-135.

16. Gerik, J. S. & Huisman, O. C. (1988). Study of field–grown cotton roots infected with Verticillium

(11)

dahliae using an immunoenzymatic staining technique. Phytopathology, 78, 1174-1178.

17. Hadizadeh, I. & Banihashemi, Z. (2005). Reaction of Pistacia vera cultivars to Verticillium dahliae the causal agent of vascular wilt. Iranian Journal of Plant Pathology, 41, 561-583. (In Farsi).

18. Hall, R. & Ly, H. (1972). Development and quantitative measurement of Verticillium dahliae.Canadian Journal of Botany, 50, 2097-2102.

19. Hassan, A. (1993). The role of fungi in fungus–nematode interactions. In M. Wajid Khan (Ed.), Nematode Interactions, (pp. 273-288). Chapman and Hall.

20. Hilloks, R. J. (1992). Cotton Diseases. CABI Publishing, Walling ford.

21. Hosseininejad, S. A. & Khan, M. W. (2001). Interaction of root-knot nematode, Meloidogyne incognita (race 1), on chick–pea cultivars. Applied Entomology and Phytopathology, 68, 1-11.

22. Hosseininejad, S. A. & Ramezani Malekrodi, M. (2005). Reaction of olive cultivars to Meloidogyne javanica.Integrated Protection of Olive Crops, IOBC, WPRS Bull. 28(9), 141-145.

23. Hussey, R. S. & Barker, K. R. (1973). A comparison of methods of collecting inocula of Meloidogyne spp. including a new technique. Plant Disease Reporter, 57, 1025-1028.

24. Hussey, R. S. & Janssen, G. J. W. (2002). Root-Knot. nematodes: Meloidogyne species. In J. L. Starr, J.

Bridge and R.Cook, (Eds). Plant resistance to parasitic nematodes. (pp. 43-70). CABI Publishing, Walling ford.

25. Jepson, S. B. (1987). Identification of root-knot nematodes (Meloidogyne species). CABI Publishing, Walling ford. 265pp.

26. Khan, A., Atibalentja, N. & Eastburn, D. M. (2000). Influence of inoculum density of Verticillium dahliae on root discoloration of horseradish. Plant Disease, 84, 309-315.

27. Letani, S., Taheri, A., Tanha Moafi, Z. & Razavi, S. A. (2002). Study of interaction of root-lesion nematode, Pratylenchus thornei, and wilt fungus, Verticillium dahliae on potato plants. In: Proceedings of the 17thIranian Plant Protection Congress, Tehran University, Karaj, Iran, Vol II, p. 227. (In Farsi).

28. Mai, W. F. & Abawi, G. S. (1987). Interaction among Root-knot nematodes and Fusarium wilt fungi on host plant. Annual Review of Phytopathology, 25, 317-338.

29. McDonald, J. D., Goldhamer, D., Bolkan, L. & Beede, R. (1992). Influence of irrigation method on distribution of water, root and Verticillium dahliae pistachio industry. Annual Report Crop Year, 1991- 1992.

30. Nickle, W. R. (1991). Mannual of Agricultural Nematology. Marcel DeKKer, 1035pp.

31. Rahnama, K., Razavi, A. S., Latifi, N. & Zarei, H. (1998). Occurrence of die-back and decline of olive trees in Gorgan and Gondad. In: Proceedings of the 13th Iranian Plant Protection Congress, Tehran University, Karaj, Iran, Vol II, p. 222. (In Farsi).

32. Razavi, S. A., Sanei, S. J. & Okhovvat, S. M. (2003). Olive Verticillium wilt in Golestan Province, Iran.

In: Proceeding of the workshop of Olive. Institute of Agriculture in Golestan Province, Iran, p. 4.

(In Farsi).

33. Rowe, R. C., Riedel, R. M. & Martin, M. J. (1985). Synergistic interactions between Verticillium dahliae and Pratylenchus penetrans in potato early dying disease. Phytopathology, 75, 412-418.

34. Ruggieri, G. (1946). A new disease of olive. Italian Agricola, 83, 369-372.

35. Sadegh Moosavi, S., Karegar, A. & Deljoo, A. (2006). Responses of some common cucumber cultivars in Iran to root-knot nematode, Meloidogyne incognita, under greenhouse condition. Iranian Journal of Plant Pathology, 42(2), 241-252.

36. Saeedizadeh A., Kheiri, A., Okhovvat, S. M. & Hoseininejad, A. (2003). Study on interaction between root-knot nematode, Meloidogyne javanica, and wilt fungus, Verticillium dahliae, on olive seedlings in greenhouse. Communication Applied of Biology Science, 68(4a), 139-143.

37. Sarejanni, J. A., Demetriades, S. D. & Zachos, D. G. (1952). Rapport sommaire sur les principles maladies des plantes observees en Grece au cours de L’ anee 1951. Annales de L Institut Phytopathologique Benaki, 6, 5-9.

38. Sasanelli, N., Fontanazza, G., Lamberti, F., D’Addabbo, T., Patumi, M. & Vergari, G. (1997). Reaction of olive cultivars to Meloidogyne species. Nematologia Mediterranea, 25, 183-190.

39. Saydam, C. & Copcu, M. (1972). Verticillium wilt of olives in Turkey. Journal of Turkish Phytopathology, 1, 45-48.

40. Tous, J. & Ferguson, L. (1997). La Colltura Dell'olivo in California. Olivae, 67, 18-26.

41. Walters, S. A., Wehner, T. C. & Barker, K. R. (1999). Greenhouse and field resistance in cucumber to root-knot nematodes. Journal of Nematology, 1, 279-284.

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