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REFERENCES
1. Bouhache, M. & David, E. B. (1993). Photosynthetic response of flooded rice (Oryza sativa) and three Echinochloa species to Changes in environmental factors. Weed science, 41, 611-614.
2. Brown, R. H. (1985). Growth of C3and C4grasses under low N levels. Crop Science, 25, 954-957.
3. Delmar, V. & Robert, A. (1981). Development of photosynthetic activity following anaerobic germination in Rice-mimic grass (Echinochloa crus galli var oryzicola). Ameican Jounal of Botany, 42, 1269-1277.
4. Delmar, V. & Robert, A. (1983). Germination and seedling growth in Echinochloa crussgalli var oryzicola under anoxic conditions. Ameican Jounal of Botany, 68, 1269-1277.
5. FAO. (1996). Weed management in rice. Pp. 272.
6. Griffin, R. M., Webster, E. P., Leon, C. T., Zhang, W. & Mudge, C. R. (2004). Comparison of rice graminicides in Louisiana rice. Proc. South. Weed Science Soc, 57, 58.
7. Griffin, R. M. (2006). Echinochloa polystachya management in Louisiana rice. The Department of Agronomy and Environmental management. Pp. 91.
8. Kennedy, R. A., Barrett, S. C. H., Vanderzee, D. & Muzik, M. E. (1980). Germination and seedling growth under anaerobic conditions in Echinochloa crus galli (barnyardgrass). Plant Cell Environ, 3, 243-248.
9. Kent, R. J. & Johnson, D. E. (2001). Influence of flood depth and duration on growth of lowland rice weeds. Crop protection, 20, 691-694.
10. Koo, S. J., Kwon, Y. W., Chin, D. V. & Cung, H. A. (2000). Common weeds invietnam. Agricultural Publishing House, Ho chi Minh city.
11. Modaraye Mashhoud, M. Esfahani, M. & Nahvi, M. (2006). Effect of preharvest chemical desiccation on harvest time and grain quality in Rice (Oryza sativa L.). Journal of Science and Technology Agricalture and Natural Resource, 11(42), 80-91. (In Farsi).
12. Mosavi, M. (2001). Integrated weed management. Principles and methods. Miaad press. Pp.460. (In Farsi)
13. Omrani, M. (2008). Study the management of barnyardgrass (Echinochloa crusgalli(L.) Beauv) weed on rice cultivars of Shafagh and Tarom. M.Sc thesis. Islamic Azad University. Science and research branch. Tehran. (In Farsi)
14. Shad, R. A., Hussain, M., Khan, R. U. & Ziauddin, M. (1986). Socioeconomic aspects of loses in rice due to weeds. Potato Agriculture Research, 7, 257-263. (In Farsi).
15. Son, N. H. (2002). Study on weeds in transplanted rice, their control method in Red River data. Ph.D.
thesis, Agriculture Hural University, No.1, Hanoi (in Vietnamese).
16. Wayne, S. (2003). Rice, origin, history, technology, and production. Rice weed control. Texas University. Handbook. Pp: 831.
17. William, L. (2003). Echinochloa Beauv. spp. Status of Introduced Plants in Southern Arizona Parks.
University of Arizona. Pp 52.
18. Yaghoubi, B., Zand, E. & Joharali, A. (2006). New specie of Echinochloa a serious problem for Iran paddy. In: Proceedings of the 17th Iranian plant pathology congress. Karaj. (In Farsi).
19. Yaghoubi, B., Zand, E. Mohammadsharifi, M., Valialapour, R., Shekari, A. & Jamali, M. (2003). Study the possibility resistance of Echinochloa crus galli to ordinary herbicides in Iran paddy fields.
Agricaltural Research Organ. Pp: 42. (In Farsi)
20. Yamasue, Y. (2001). Strategy of Echinochloa oryzicola Vasing. for survival in flooded rice. Weed Biology, 1, 28-36.
21. Yamasue, Y., Asai, Y., Ueki, K. & Kusanagi, T. (1989). Anaerobic seed germination for the habitat segregation in Echinochloa weeds. Japan Journal Breed, 39, 159-168.