• Tidak ada hasil yang ditemukan

The Effect of Mycorrhizal Fungi, the Combined Ratio of Auxin/Gibberellin and Ascorbic acid on the Yield, Yield Components and Germination of Corn

N/A
N/A
Protected

Academic year: 2024

Membagikan "The Effect of Mycorrhizal Fungi, the Combined Ratio of Auxin/Gibberellin and Ascorbic acid on the Yield, Yield Components and Germination of Corn"

Copied!
18
0
0

Teks penuh

(1)

Online ISSN: 2345-6957 Homepage: https://jci.ut.ac.ir/

University of Tehran Press

The Effect of Mycorrhizal Fungi, the Combined Ratio of

Auxin/Gibberellin and Ascorbic acid on the Yield, Yield Components and Germination of Corn

Tayebeh Jamshidnia1 | Tayab Sakinejad2 | Ali Reza Shakofar3 | Mohammad Reza Dadnia4 | Seyed Kivan Marashi5

1. Department of Agriculture, Ahvaz Branch, Islamic Azad University, Ahvaz, Iran.E-mail: [email protected] 2. Corresponding Author, Department of Agriculture, Ahvaz Branch, Islamic Azad University, Ahvaz, Iran. E-mail:

[email protected]

3. Department of Agriculture, Ahvaz Branch, Islamic Azad University, Ahvaz, Iran. E-mail: [email protected] 4. Department of Agriculture, Ahvaz Branch, Islamic Azad University, Ahvaz, Iran. E-mail: [email protected] 5. Department of Agriculture, Ahvaz Branch, Islamic Azad University, Ahvaz, Iran. E-mail: [email protected]

Article Info ABSTRACT

Article type:

Research Article

Article history:

Received: 2 February 2022 Received in revised form:

19 November 2022

Accepted: 30 November 2022 Published online: 24 June 2023

Keywords:

Corn,

Fungi growth regulator.

Harvest index, Optimized growth.

The present two-year research (1397-1398) investigates the effect of Mycorrhizal inoculation. For this purpose, it takes the combined ratio of Auxin/Gibirlin and Ascorbic acid on the yield and yield components of corn, in the form of split factorial design in the form of randomized complete blocks in three replicates in Shahid Salemi farm. It has been done north of Ahvaz city. The main factor of the treatments includeMycorrhizalfungus (Glomus intraradices species) at two levels (inoculation and non-inoculation) before planting and the secondary factor includes the ratio of Auxin hormone to gibberellin at three levels (0, 300/600, and 600/300) mg/liter and ascorbic acid at three levels (0, 100, and 200) mg/liter are applied as foliar spraying. The results show that Mycorrhiza has a significant effect on the dry weight of leaves, stems, and cobs, seed weight and yield, as well as biological traits, respectively, the highest seed yield amounting to (7410.72) kg is related to the treatment levels of Mycorrhizal inoculation, the ratio of Auxin to gibberellin is 600 mg to 300 mg and in 200 mg of ascorbic acid and the highest yield of stem dry weight (100 grams). In the triple interaction of the experimental factors in the treatment levels, there has not been any Mycorrhiza inoculation. For not using the hormonal compound and using 100 mg of ascorbic acid, the highest seed yield in the row (45.46%) has been obtained in the ratio of Auxin to gibberellin of 600 mg to 300 mg and using 200 mg of ascorbic acid. According to the positive effect of the use of experimental factors, the use of Mycorrhiza and the mentioned hormones, and ascorbic acid are recommended.

Cite this article: Jamshidnia, T., Sakinejad, T., Shakofar, A. R., Dadnia, M. R., & Marashi, S. K. (2023). The Effect of Mycorrhizal Fungi, the Combined Ratio of Auxin/Gibberellin and Ascorbic acid on the Yield, Yield Components and Germination of Corn. Journal of Crops Improvement, 25 (2), 419-436. DOI: https://doi.org/10.22059/jci.2022.338452.2674

© The Authors. Publisher: University of Tehran Press.

DOI: https://doi.org/10.22059/jci.2022.338452.2674

(2)

نا هﺎﮕﺸ اد تارﺎﺸ ا

یزروﺎﺸ ﯽ ارز

Homepage: https://jci.ut.ac.ir/

:ﯽﯿوﺮ ﻟا ﺎﺷ ۶۹۵۷ - ۲۳۴۵

/

!

" #$

% &

' ( ) *

& +

|1

2 3 |

!|

|4

#

5 !

1 .

. : [email protected]

2 . ! "# $#

.

: [email protected]

3 .

. : [email protected]

4 .

. : [email protected]

5 .

. : [email protected]

% &! ' (

)

: % &! + () *+ ,

, - . - / : 13 / 11 / 1400

:0 1 2 3 .- , 28

/ 08 / 1401

, - . 45 - 6 : 09 / 09 / 1401

, - . 7 : 03 / 04 / 1402

8 9 : :

1$2 3 4 5 6 7 $$8

9 :

;< 7 3$

7 < => 7 .

<

1$

<

3?@2 ABC < 3 D3< A E3+ 3FG /E3#8 F38 2 F# H A I3,B2

HG J ABC 9 : + K () E )

1398 - 1397 ( <

9 P Q R S 3 T B 8 U V W+ 6

B<

X J 8 T Y2 V 2

A H 2 ,3,Z2 ;7 ! C7 [6 C+ 3;7 \ ]

. T H A 5 6 V 7 C32 BP U8 T * )

^ B U$

H#

( I3,B2) I_

K3) (I3,B2 \ 8

+ F3G < E3#8 F# V 7 T U8 T 7 ) I_ E3

`P

P 600 / 300 300 / 600 ( B3 U3+ < \ D3< A

3 ) I_

`P 100 200 (

B3 U3+ < \

<

9 P

! BZ 7 ) b < D > < H A 8 c U . 7 \ ]

ABC ! < 6 4

E3$d A e + 3< 9 `P 3?@2

f$

. J

<

W32 2 E 2g <

) ABC , 72

/ 7410 ( B38

\ E3+ F3G < E3#8 F# H A I3,B2 J C32 R _

600 B3

\ <

300 B3 \ 200 B3 \ D3< A 3 ABC E 2g <

D >

) 6 100

\ ( 4 <

K$8 V

J K 2 R _ 3 C J ,B2 \ 3 I A H \

8 2 h Y 3 W h Y

100 B3 \ D3< A 3 2g <

E ABC 3 H

i ) 46 / 45 P

#8 F# ( 3

E G <

3 + F 3 E 600 B3

\ <

300 B3 \ h Y 200

B3 \ D3< A 3 <

. < G 2 <

3?@2 U8 T < 8 Fj J

`U

3 A H J

< A 8k 3

D 3 P 2 3 . 7

: ;7 CG 3 3 S l F3 8 ( S lW3 A7 T B l m 3 m CZ 38 3

) 1402 .(

<

?@2 3 5 6 A H F#

8 2 3F

#8 3 E / FG 3 +3 E

< A 3 D 3 <

P Y>

3 9 T 3 H e + J 9 :

< .

J 8

25 ) 2 ( 419 - 436 .

DOI:https://doi.org/10.22059/jci.2022.338452.2674

:

;2 9 U # n

. . $# ©

https://doi.org/10.22059/jci.2022.338452.2674 DOI:

(3)

1 . . !/

4; 9 :

< 6 9g YZ E 2 ( V3+

< > 3#< J

< ( < J 9 6 V3+

C3B6 o 7 > < 8 I_ 1 c < \ $ f< \ A UT p U# 3 \ C2 3#<

q YU>

)

Mahender et al., 2018

.(

< D H 25

- 20

< 9 : ;G 9 3+ 2 P 43,U# 9 P

VA7 \ kr h Y < ( 3r B3 $ 37 9 : ) iBUs J

< . c$)

H3 9 : 3+ 2 P

T ;G 6 `U Uf$P J

3 $< >) 1387 .(

3 9 : 8 I_

140 UA 3B3

60 7 p H UA H .

3 3+ 2 E3 3 3500

3+ 2 E3 3 UA \ B38

T B 36

7 p H UA E2 AC AB )

1399 .(

E <

D (2 U 4; ! YZ D 9 : 8

7 <

i3fm ! YZ E 3+ 2 8

<

3+ 2 <

E3<

8 / 6 - 2 / 7 UA E2 F2 ! 9 :

62 3

. 6

! m K () <

! F + 2 3 3F J 9 : <

( t `2 2 8

$U#

8 48 2 k) J ABC

K3<

2 J 6 3

^ <

3F J G + ) > <

1393 (.

p J <

! 9 : \ 6 R P

f 2 7 <

4 9 :

# U) $d 3

V J <

J 7 < u ) 3

J <

HT h < R P K

3 H ABC

< . > < ! YZ v2 t 6 B

33 9 2 $T 3Q v2 < w< _ 33

9 7 o Z 3 _ T f D 3F < 9 : J

3 [ 7 <

o x iBUs 8 >

7 AC J CZ )

1392 .(

, 3, x sU 3F W $ A

V x #Z

7 S 8 w

# U) 3 V U e 3 A G 3

3?@2 ! YZ ABC <

k ?

4 <

2 e K$8 3

y Z 3 o

2 A 6

s 2 3

= 3F J 2 <

3 G < 9 : AC J CZ )

1392 l

2016 et al., Dayal

.(

2 12 34 .5 6 4 .

8 <

J 3 J

>

3 p U# < G 2 <

8 J ) UT 6 G 2

E

<

`B2 3, 3 6 G 2

3 C 7 . 3

C] o 2 J

+ A

! J 8 d D

<

\ J

3 1$2 3 4 E . 7 > < 3

K ()

< S 8 $U7 ;z > J

! U$8 C; K, E3U$38 E3+ F3G

$ T ` 3 9 3 > d ! S iBUs J

$$8 )

Khan et al.,

2007; Javid et al., 2022

.(

D3+ F3G ! T 3

h f E U;< A GA

E E 2 . 7 <

8

# 3 E 3 Z K, J

1$2 3 4

$ T J T iBUs 3 H e + D BCG

<

( J

T

<

; )

Marschner, 2011

( .

<

\ ] ? 7 J

8

# 3 Y$ E J

<

7

Ks<

3 < ? 8 5 6 .

J A3 H J <

BP {HG $ K3<

2 8 3 4U#

UFj 9 ? <

P Y>

3 9 8 C 38

` 3 HC

#< 9 f+ _ . # 3

J 5 6 K, <

J A3 H J HT K

2 3 xkG K3<

2 f P $ 7 7 x

)

Tao & Zhiwei, 2005

(.

2 P 8 Fr < C2 K3) 8 JH3d < < E $d $ s< 8

$3<

8 9g YZ

;< K > J 3+ 2 $

7

| CU # <

< 2 $7 < U7 $7 < G 3 7 X Z D <

3 7 X Z < F# 8 U> $7

<

W#8 J U;< o 7 $ U<

4 T > J 8

.

4;

V E 2 Z

X 7 3 HT BC K A HT ! YZ , K

3 2

<

< V 3 C J J

8 C D <

8 K$2 K J

Z 3 _ 7 BCG J

>

A 4; C 2

8 h Y K 8

J 7 3 C 3

(4)

C 7 )

Rai & Varma, 2005; Chen et al., 2018; Ronga et al., 2019; Hosseinifard et al., 2022

.(

4 E3$d 5 6 J A3 H J 4 T <

I_ C

xkG 8 $$

3 [

< 2 J kr P $ ! ,U

> G X

<

3 7 F;< WF 3

. H

J

` } F2 E 3

+ 2 < 2 3

J 5 6

A < 6 H 8

1Br K D H#<

3 HT K 8

# 3 FG E 3 +3 . $U# E

.

Sun et al

) ( 2018

! BZ ! C ~) 7 )

D3+ F3G 3

1Br J 20 30 40 50 60 \

UA 3

9 : K3<

2 ! < ! S 3 ! < x d f2 + 8 f2 3 t `2 9 `P K HT E

! C < u ) ! < f2 D3+ F3G

3 1Br 8 8

60 UA \ K3<

2 E 3?@2

7 • T 9 `P H3 F;<

.

f+ _ J

<

1$

<

3?@2

! BZ 7 ) D3< A 3

P Y> <

3 e + $T 9 T D

3 H e + 7 9 : D 3

E

4 e Z2

< J 3 J <

9 8 9 P X B< R S W+ 6 7 > J

8 J T Y2 V A2 <

! U 1393

U . 7 \ ] A > K$2 8 c

! < f2 2 < t `2 K 8 WG

H T B8 ABC 3

UZ V8 V J

F# x HT b <

+ ) K 3 + fT E 3 4 H

#8 ) J 3

2 C#

< 7 g 2 8 J S

8 7 Z2 7 K$2 o

A >

! < f2 H

ABC

<

W32 2

03 / 43 11 / 23 49 ) K 8 P 3

V< , . 8

! BZ 7 ) D3< A 3

HT WG 2 < t `2 K

H T B8 ABC 3

UZ V8 V J

F# x + ) K 8 b <

3 + fT E 3 4 H

#8 ) J 3

2 C#

7 g 2 8 K3<

2 E 3?@2 2 Fj 3 C 120 3B + \ 3 U D3< A 3

. 7 4 E3$d

! BZ 7 ) <

120 3B + \ 3 U D3< A 3

HT WG K 1 / 32 P J ABC 7

)

J T

AC 1395

.(

f+ _ c U

J J

E3+ F3G K$2

A >

<

9 : 8

D3+ F3G 3

w S 3?@2 Fj

<

K HT F;<

HG J ABC 2 ABC 9 :

K HT AC AB )

1399 .(

h f+ _ m

<

1$

€ B

;G F;<

3C8 9 3P Y>

G 9 : 3 k<

V $3

^ 8 704 5 6 8 7 <

A H 8 2 F#

3F

#8 3 E FG / 3 +3 E D3< A 3

w S 7 UT ;<

~ J J

=> 7 J

1 > ) V3BZ2 H]2 <

. 7

3 . 8 12 34 59

E H K 2 ,3,Z2

C+ 3;7 3T vG ! S H E . G < ;7 ! C7 [6

48

G 40 3T vG • 6 7 ,36 31

G 30 t `2 < + C7 ,36 5

/ 22 E . [6 I_ U

43#,2 wFS 3

> 43B6 J E) 8 J $<

D 3 C

>

D <

Jg < g E3 3 J 18

U G

m ) 2017 .(

K () E ) !

1397 1398 J 18

2 20 U G (

<

R S 9 P

Q3 T B 8 U B< W+ 6 V X

J 8 T Y2 V 2

A G I_ H A 5 6 V 7 C32 . 7 (i+

\ I3,B2

<

$ I3,B2 (x 7 k<

<

5 6 A H

^ B H# U$

>) J 8 7 7

# HF $T

, <

100 (\

$UT 6 BP 9 8 E3+ F3G < E3#8 F# V 7 T U8 T

J >) 7

q B> , < U 8 8 7 10

2 25 J P ( ) I_

P `P 600

/ 300 300 / 600 ( I_

D3< A 3

) V 7

`P 100 200 U3+ \ B38 (

<

9 P 9 8 V U8 T . $UT 6 T J

5 6 J

A H

`U E

K () m

\ $ 4 8 G J U

V <

3 \

# n

(5)

,Z2 3 9 ,

>

X x 8 2 A j3 + 2 3 7

<

3 H

\ V 7)

<

$ V +

3

\ 3 i d 6

< ( Q J

8

<

J k<

150 - 200 7 7 UT 1 5 6 ! fT \ 3

J

q Ys 8

7 k<

wC

;d U U J 6 7 E3#8 .

G F3 +3 E D3< A 3

<

3 H P

+3 U

<

1Br H

<

9 P J k< h Y 23

C G F Z o 2

x T

<

k<

8 |

U r 7 8 .

# 3 B E c$)

<

! BZ 7 ) 7 G F3 +3 E ~)

8

# 3 T m 9 : < E 7

.

D3< A 3

J >) 7

# n C37 AUF q B> , <

90 P

<

9 P (x ! BZ

) I_

`P 100 200 B ( c$)

<

<

9 P

! BZ 7 )

`U T 6 .

<

1$

8 3`38

xkG ;G J 8 ‚` 3 D f6 7 < w< _ K3<

2

`U J 3 o 2

K3) .

\ ] K D W8 C

>

X H 3;2

<

>

X x 3

. 7 ! ;7

t

>

X VZ K UB3 - + <

6pH

/ 7 7 EC

/ 1

~$C

<

U

<

.

k<

\ 6 f+ _

H8 9 ,3,Z2 8 J [< $ ;7 f3FS 3;2

7 F3 9 : k< .

V $3

^ 8

704 i E3< BP T <

75 U 2 < BP T U 17

U

. 8 U

<

J I3,B2 k<

`

\ I3,B2 8

\ J 107 5 6

A H

! fT

<

`U 7 3B8.

9 3BC o3Z

x UT

\ ] T k<

~) D >

7

7 8

E3 .

1

! VF6 K 3;2 E3 V 7 w3C 4s7

D#

f<

7 E3

~Q I3_#2

<

B3 + +

<

9 8 D .

< J K7

7 8 o>

< < ! S K7

U

`U 7 8 . T 6 ]$)

4 . \ ] <

`U f+ _ E

8 J U#

<

1$

I3,B2 k<

ƒ <

7 > 3+ 2 J

V S 2 2 <

J 46

9 : .

\ J 3< 7 8 f< BP T < ! J 3<

;d J 3< . T 9 P 7 8 f<

J f< J

^ <

75

B3 ^ 8 3sF2 DU 2 3sF2 U . 7 431$2A

EUT 1 < 9 `P J 3 D

J ;U U< U

< 9 8 37 $ 4

E3$d x sU 7 8 } _>

D K7 <

$ . 7 \ ] 37

J <

! BZ C8 1Br H3 J 3 B ~$d p 3#8 )

)

Chance and Maehly

( 8 <

7 `U 9 33v2 .

J 3

^ <

! A 7 3#8 H3 \ ] 4 H E o 21

p E . 3 33

B3 D < „ sU Y U3+ A3 V 7 8 3#8 ) ! BZ U3+

13 B3 g ! A 5 B3 g

e 3 3#8 ) )

H2O2

( 50 B3 g 43 U) 9 `#T T <

8

= C } Bs pH

<

9 < ,36 D

P T V 10

! S 3 ?

„ 470nm

EU> J < . 7 > xkG 100

B3 9 `#T T < U3+

43 U) 39 B3 U3+

9 `#T D < 43 U) 50

B3 g <

61 B3 9 `#T U3+

J 43 U) D <

50 B3 g 7 W38 2 .

3 . 1 . & :"

; <=) ; <=)

=s < ;G J C32 b < I_ 9 33v2 C

<

BP 3 J < 7 B

C J <

~) 14

K t 7

b < U< . 7 7 < 9 8 2 <

7 G 6 2 < J

b H< • b < ! S

;) E 2

$ b < J

<

B3 U 3

< ]2 p < 2 < I_ ~Q 7 J

F Z )

A=L*W*0.75

( . ! T E b < # A

b H< W

b < J $;) E 2 b < ! S L

m _< b < I_ => 7 ~Q . 7 <

) 1 (

<

)

Amanullah et al., 2007

(:

1. Gaiacul

(6)

*_<

1 (

SA LAILA

) b < I_ F#

(LAR

< . 3 V8 < 3 b < I_ F# $2 F 1

I_ D$;) I_ F# 2 F

T <

J T < < $$8HU$ UT

~`$2 J . 3 $$8 H3 3 D < ) $ LAR

7 <

H3 b < 3+ 2 3 J k

.

$$8HU$ UT U F# n LAR

) \ < [< U

m2/gr

( 43#,2 w S 7 <

< F# 7 =+ > xkG

<

.

<

J E33f2 H

10 C 100 k< 2 J

< 9 8 7 x sU T Y2 S

^ <

14 x sU ~) H3 ! < f2 i f2 ! < i f2 9 `P . 7 F Z < S P

T Y2 10 ! <

7 J 3 7) AC $3#

1392 .(

<

i ; ABC F Z 1$

o J

! < f2 2 < f2 U< 1$ E < . 7 7 < (J 37 B ) ; 7 < 37 < 9 8 ABC F Z ! < ABC ~Q p C7

<

^ 14

< S U `U < < S P c$

9 r

! )

Farmex MT-PRO

>

8

$T 7 E33f2 ( .

7 < => 7 )

(HI

43#,2 < H3 < ABC 8

3< ABC

< P W# < D e +

7) AC $3#

1392 .(

_<

* 2 (

100

× J YU6 ABC =

7 < => 7 )

(HI

A e + 3< ABC

3 . 2 .

>* !

?$ 9 ( @

!9 ( 2

! YZ 7 )

(CGR

`U < [< U < \ W# < 2

_<

) 3 ( 7 F Z )

Karimi &

Siddique, 1990

(:

_<

* 3 (

1 2

1 2

T T

w CGR w

 

W1

+ U 7 < J 3+ 2 D > :W2

T1

T :T2

7 < E3< BP

4 E3$d

=+ > HU$ UT =+ > xkG )

(NAR

< b < I_ [< U < \ W# < 3

`U _<

) 4 ( 7 F Z )

Wilson, 1981

(.

_<

* 4 (

1 2

1 2 1

2 1 2

LA LA

A Ln A Ln T T

w NAR w

3B8 ;

\ `U < BP J J HT

)SAS

(9.4

E3 3 . $UT 6 ~ H]2

$f 9 `P 7

$ $d ^ <

EA J I_ 4

! CU c$) J < . 7 # , P H]2

V3BZ2

\ HT

26

\ C 4 ;G `U SPSS

HT

Excel

) (2016

7 < ;<

.

1. Lowest Area Ratio 2.Crop Growth Rate 3. Net Assimilation Rate 4.Duncan's multiple range test

(7)

4 . .B%

12 34 ( 2

c U H]2 ! G

~ ! ? 8

<

HG 7 < => 7 BC 9 `P

A HG J BC A

$f BC F A

HG J BC A Z2 3?@2 h Y 8 <

A . T 6 H 3?@2

F#

J

8

# 3 FG E 3 +3 9 `P \ C2 < E

<

$f

. 7

3?@2 A

<

D3 3 <

HG < b < F# < S P

9 `P

<

$f V< ,U ? . 7 A

f2 < o,T H

$f i ? . 7

V< ,U A F# H 8

# 3 FG E 3 +3 E

<

HG BC <

A <

9 `P

<

$f

. 7

4 E3$d

F# ? 8

# 3 FG E 3 +3 E D3< A

< 3 J UZ F#

x b <

) (RWC

i f2 1

! <

$f ? .

A H

D3< A 3

9 `P \ C2 <

<

$f

. 7

T ! ;d V< ,U ? 8

U

<

$f .

D3< A E3+ F3G/E3#8 C32 H A BP ? W8 H]2 c U 3

4 <

$8 K

H A - H A

- D3< A 3

/E3#8 E3+ F3G

- D3< A 3

4 E3$d 4 <

K$8

! CU I_

D

$f P ! G) <

1 ( . ) b < D > E 2g <

74 / 61 ( P 4 <

K$8

V

K J

I3,B2 J C32 R _ E3+ F3G < E3#8 F# H A

600 B3 \ <

300 B3 \ \

h Y D3< A 3

VA7) 7 VP 1

- (i+

.

4 . 1 . . 6$ C*

E3 3 # , c U I3,B2 8 H3

H A

$f K 8 ƒ <

6 D >

53 / 97 <

70 / 82 P

4 . 7 W38 2 h Y E3$d 600

< E3#8 300 B3 \

$f K HT ƒ < E3+ F3G 7 < F# 6 D >

D3< A BP ? < _< . 7 D3< A C32 7 H3 3

$f K HT ƒ < 3 F# 6 D >

K3< 8 7 7 <

D > , E 2 1Br 6

100 B3 \ D3< A

< 3 .

E 2g <

) 6 D >

52 / 116 ( P 4 <

K$8 V

K J

I3,B2 \ J C32 R _

h Y W38 2 h Y \ H A

100 B3 \ D3< A 3

7 VP VA7) 1 - x .(

T CU )

1399 ( V< 6 x H3 K 8 <

T B D > X > `U

<

S

$f J

K 8

<

K 8 V f2 ƒ < H A 5 6 < I3,B2 7

<

J S 8 K 8 32 < W# P

7 .

f+ _ UT E 7 p H H3 < 9

AC J 8)

1393 .(

3

G 3 + F 3

< D 1$2 $ 3 4 7 $$8 3

#,2 <g < <

3 + B 4 G f 2

;U J F G

HT ƒ <

K

kr xkG

#,2 8 d . 3

+ B 4 3

K3<

J 2 kr <

< G K3<

2

xkG 8

<

U ) F;< ƒ J

7 J 8 . 7 > V 2 7 BCG 7 K HT E

K3<

2 C#6 3

; WG K3<

2

\ D > 7 C#6

J 7

.

4 . 2 . D ; ; . ; ; !E

/E3#8 C32 H A ? H3 W8 H]2 c U G

3 + F 3 E D3< A 3

4 <

K$8 A

H -

A H - D3< A 3

- D3< A 3

4 <

K$8

! CU I_

D P

1.Relative Water Content

(8)

$f ! G) <

1 ) i H3 E 2g < .(

46 / 45 P ( 4 <

K$8 V

K J

E3+ F3G < E3#8 F# H A I3,B2 J C32 R _ 600

<

300 B3 \ h Y 200 B3 \ D3< A 3

VA7) 7 VP 2

( .

4 . 3 . FG ; D ; ; !E

/E3#8 C32 H A BP ? H3 W8 H]2 c U ! CU I_ E3+ F3G

D P

D3< A 3

! CU I_

c$)

$f P . <

4 E3$d ;$2 c U 4 <

K$8 H A

-

! CU I_

c$)

$f P ! G) <

1 i f2 E 2g < .(

4 <

K$8 J C32

`U E3+ F3G < E3#8 F# H A I3,B2 J C32 R _ 600

<

300 B3 \ C32 I_

D3< A 3

VA7) 7 VP 3

(.

4 . 4 . 9 ; ?$ 9

/E3#8 C32 H A BP ? H3 W8 H]2 c U 3 D3< A E3+ F3G

4 <

K$8

A H - H A

- D3< A 3

E3+ F3G /E3#8 -

D3< A 3

4 <

K$8 I_

! CU D

$f P ! G) <

1 .(

4 E3$d

$f 9 `2 7 < => 7 ! 3vU . J

) 7 < => 7 , E 2g <

45 / 49 ( P 4 <

K$8 J C32

`U I3,B2 J C32 R _

E3+ F3G < E3#8 F# H A 600

<

300 B3

\ 100 B3

\ D3< A 3

J C32 < 8 7 VP

E3+ F3G < E3#8 F# H A I3,B2 300

<

600 B3

\ 7 D3< A 3

$f 9 `2 7 J

<

$f 9 `2 C32

VA7) J

4 ( .

< G 2 <

E 8 H _< 7 < => 7 A

ABC <

3 f< 2 HT k+ . ABC

K

ABC HT ƒ < 2

7 < => 7 `P K 3

. 7

<

1 HT E

7 K 7 < =>

3 <

V3+

7 F;<

kv2 o J 3 2 ! C ? 3

5 6 . 7 < • T J C 3

A U# AF7 < H 3

# 3B 3 < \

,S $ U X >

8 J

3 C : ` wS $ 2 8

. $ HC J ) J <

1393 ( 3 H

p H 8 5 6 3 A H HT ƒ <

K 3 9 : 7 < => 7 H .

4 . 5 .

I J ( BK

/E3#8 C32 H A BP ? H3 W8 H]2 c U E3+ F3G

4 <

K$8 H A

-

4 <

K$8 ! CU I_

D

$f P ! G) <

1 ( . ) F# < S , E 2g <

39 / 75 (

P 4 <

K$8 V

K J

E3+ F3G < E3#8 F# H A I3,B2 J C32 R _ 600

<

300 B3

\ 100 B3

\ D3< A 3

VA7) 7 VP 5

.(

H A3 3 b < x F# J UZ K HT V3+

i3 K, < 2 xkG d 6 J

K3< xkG < H A3 3 . F# < x x 2

. $7 < U7 J 2g < x F# J UZ $ 2

4 Y2 E3$d H3 U# HC o 7 A3+ 3 < H A3 3 x xkG K HT 7

7 < oF2 .

Aliasgharzad et al.

) ( 2006

3 7 B b < x F# J UZ 8 )

B

(9)

‡ B<

( 48 <

K 8 X > < S 7 h P . <

x F# J UZ H A3 3 X > < S 1

< b <

g 3r 3 < F# J 2

$U7 H A3 f+ _ .

K ()

! BZ H3 7 )

D3< A b < x F# J UZ K HT WG 3

) 7

J T

AC 1395

.(

4 . 6 . ' ; <=)

U , c # 3 3 E

<

3 E <

8

$f K HT ƒ < H A I3,B2 . 7 D e + 3< ABC

4 E3$d

8 2 h Y W3

600 < E3#8 300

$f K HT ƒ < E3+ F3G < _< . 7 7 < F# D e + 3< ABC

BP ? D3< A

3 D3< A 1Br K HT 7 H3

$f K HT ƒ < 3 7 D e + 3< ABC

! G) 1 ( . ) D e + 3< ABC , E 2g <

19 / 15356 ( P 4 <

K$8 V

J R _

E3+ F3G < E3#8 F# H A I3,B2 J C32 600

<

300 B3

\ 200 D3< A 3

B3

\ 7 VP

VA7) 6 - (i+

.

. 9 : 3 < H A3 5 6 < 8 J C32 W38 2 Fj 9 ? D e + 3< ABC K HT V3+

H A3 5 6

e U3 `#T H3 K HT \ 3B3#3 3+ 2 w S 3 3 kr P $ x xkG V3# U) J 48 P $

< X > h Y 3 `U 1$

48 Je H A3 3 [6 . 7 <

V3A 2 J < J 2

h P

$$8 3 E k+

3 < F# J 2g < \ F#

J Y ) 1392

.(

4 E3$d

D e + 3< ABC K HT V3+ 8 T ]3U 2 4; C HU$ UT E3#8 K, < 2

$ HF 2

b < EU7 9 : 7 . 7 < 2HF J

< 8 \ J C32 8 < 8 < F#

b < pH K3< 9 : J

2 )

Ashraf et al., 2006

.(

K HT + S Z K HT < H3 E3+ F3G

3 BP T 9 : + S 7 K HT < ƒ <

7 ) D e + 3< ABC K HT ;

) H$3A) 1383

J 8 .(

AC )

1393 ( ? < <

T B 9 : 3 7 < E3#8 E3+ F3G J J

K HT 8 3 ]3U E < E3+ F3G iBUs R _ 3?@2 Z2 T B V8 ABC 3 2

K HT E K, 3 HU$ UT 3# E3+ F3G 8 E $

K HT < ]$ HU$ UT F;< 8

9 3C3

; 7 > D e + 3< ABC .

4 . 7 . . ; ; <=)

/E3#8 C32 H A BP ? H3 W8 H]2 c U D3< A E3+ F3G

3 4 <

K$8

/E3#8 E3+ F3G - D3< A 4 3

4 < E3$d K$8

! CU I_

D

$f P ! G) <

1 ( .

) ABC , E 2g <

72 / 7410 ( P 4 <

K$8 J C32

`U I3,B2 J C32 R _

E3+ F3G < E3#8 F# H A 600

<

300 B3 \ 200 B3 \ D3< A 3

J C32 < 8 7 VP

E3+ F3G < E3#8 F# H A I3,B2 600

<

300 B3

\ 100 B3

\ D3< A 3

$f 9 `2 J

7

$f 9 `2 C32 <

VA7) J

6 - (x .

BF6 9 f+ _

! BZ 7 ) <

120 B3 \ D3< A 3

K HT WG 1

/ 32 p H ABC P

) < 7

J T

AC 1395

.(

K HT V3+

2 Fj 9 ? ABC 38

2 W 3 J C 8 5 6 <

3 A < H

3 [6 . 9 : 3

3 A H Je 48 2 A 2 J < J 3

V h P 8

$$

k+

E 3 F#

(10)

\ < F# J 2g <

3 J Y )

1391 .(

G 3 + F 3 HT < H3 E + S Z K

HT K

3 BP T < ƒ <

9 : + S 7 K HT 7

) D e + 3< ABC K HT ; )

H$3A)

1383 .(

3 7 J <

;

< 3>:

3>: Hs ;$2 $

> `U 3 J 7

C; K, 8

) ABC K HT AC J 8)

1393 .(

431$2 Fj ? < w< _2

$$8 c U ABC < 7 J

f+ _ ) AC

1396 ( U3 h Y

H3 < ABC K HT ƒ < E3#8 E38 3

/ 34 5 / 43 P

7 3> f+ _ c U . H3

8 D3+ F3G 3

w S Fj 3?@2

<

K HT F;<

ABC J HG ;

2

ABC 9 :

K HT AC AB )

1399 .(

4 . 8 . 4 &

* !

D3< A W38 2 H A BP ? H3 W8 H]2 c U 3

4 E3$d 4 <

K$8

W38 2 -

D3< A 3

4 <

K$8 I_ C32

D

$f P ! G) <

1 E 2g < .(

, A ) 9 <

8 3#

4 <

K$8 V

K J

W38 2 H A I3,B2 J C32 R _

600 E3#8 - 300 E3+ F3G B3

U3+ < \ h Y \

D3< A 3

VA7) 7 VP 7

(.

K9 1 . 9 : ABC J HG ABC 9 `P W8 H]2 c U

' L3 ! C 1 !

02E

M, N3 ;!

'32B5 RWC F-8%3 O-L,K(3 /3 PQ O-L, R, FQ #29 ? FQ #29 S3 FQ #29

0,01ns

13,08ns 10214,40ns 1039,70ns 11,3ns 1,001 ** 7,54ns 0,28ns 64,49ns 20,80ns 1 (Y) ! 3,19

206,27 313012,13 315313,30 38,75 7,39 81,65 15,26 891,00 775,33 4 (! ) X B<

8,71**

2998,65 ** 60923526059 ** 224546990,50** 122,34 ** 7934,30 ** 2824,12 ** 0,99ns 5934,54** 96,85** 1 (M)H A 0,22* 6,52ns 376,80ns 14180,30ns 0,46ns 0,69ns 56,73 ** 0,14ns 23,22ns 5,86ns 1 Y×M

0,12

8,99 3316,56 27954,60 1,60 3,02 0,33 0,56 2,40 8,06 4 BP J _>

1,90**

729,49 ** 28689656,06 ** 17346705,10** 2,11 ** 341,79 ** 382,61 ** 1,46 ** 573,87** 349,31 ** 2 (H) E3+ F3G/E3#8 0,07ns

2,05ns 8555,67ns 10413,30ns 0,002ns 0,65ns 6,18* 1,44 ** 0,02ns 0,04ns 2 Y×O 0,04ns

121,69 ** 4009185,93 ** 364861,20ns 5,65** 7,25 ** 66,90 ** 2,99 ** 1082,26** 335,59** 2 M×O

0,01ns

13,61ns 166,55ns 17845,00ns 0,67ns 0,92ns 26,08** 0,69ns 20,82ns 19,94ns 2 Y×M×O 0,66**

12,94ns 17043203,5 ** 752245,30** 2,63 * 23,22 ** 56,84 ** 2,65 ** 1010,04** 90,71** 2 (A) 3 D3< A 0,01ns

0,86ns 4034,32ns 5110,50ns 0,92ns 0,12ns 3,13ns 0,02ns 1,31ns 27,80ns 2 Y×A 0,08ns

1,84ns 206796,39 ** 15385,80ns 1,66ns 3,45 ** 5,96* 2,26** 929,26** 964,44** 2 M×A 0,02ns 3,63ns 4780,40 ** 5476,50ns 0,22ns 0,16ns 2,66 ** 0,05ns 46,91ns 10,20ns 2 Y×M×A

0,30**

8,87ns 550884,80 ** 82552,50** 1,49ns 3,41** 7,18ns ** 7,33 ** 1586,39** 221,73** 4 O×A 0,08ns

3,68ns 15604,29ns 1187,00ns 0,18ns 0,31ns 6,94** 0,12ns 6,44ns 16,26ns 4 Y×O×A 0,21**

31,05ns 141202,34 ** 52076,40** 0,18ns 6,82 ** 8,87 ** 1,52** 1235,62** 73,89** 4 M×O×A 0,004ns

1,13ns 3811,58ns 7067,00ns 0,43ns 027ns 1,62ns 0,14ns 73,77* 15,41ns 4 Y×M×O×A

0,05 5,34 6893,5 9864,3 0,72 0,51 1,82 0,27 27,34 11,72 65 _>

13,26

3,49 6,06 19,22 7,12 19,27 4,01 8,68 5,80 8,52 (%) 9 33v2 W m

* ns

< :**

$f 3r W32 2

$f ! CU I_

5 1 . P

(11)

5 . OK

I3,B2 J k<

\ $

<

5 6 A H F#

#8 3 E FG / 3 +3 E D3< A 3

B

\ 6 9 : ?

V< 6

;G 2

<

ABC ABC J HG 3`38

G J k<

9 : J 3+ 2 7 < H A 8 c U .

4 ABC ! < 6 b < D >

$f 3?@2 A e + 3< 9 `P E3$d . J

l(635 : U J _>)9 : 3 D > < K J V K$84 < ? ( i+ . 1 R 4 < ? (x K$8 V

K J

9 : 3 6 D > <

: U J _>) 606

.(

h U> G \ 3< U X U h

$f ^ <

EA 7 <

. bcd

efg hi

efg efg ghi

a

efg bc

i b

cde hi

b

f-i fgh

cde def

0 10 20 30 40 50 60 70

Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Auxin to Gibberellin

(0%)

Auxin to Gibberellin (600/300)

Auxin to Gibberellin (300/600)

Auxin to Gibberellin (0%)

Auxin to Gibberellin (600/300)

Auxin to Gibberellin (300/600)

Mycorrhiza (inoculation) Mycorrhiza (inoculation)

S3 FQ #29)N3!7! VW(

) O%

k

hi i

b

j d-h

i c-f

ghi c-g a

cd c

ab e-h

i

cde fgh

0 20 40 60 80 100 120 140

Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Ascorbic acid (0) Ascorbic acid (100) Ascorbic acid (200) Auxin to Gibberellin

(0%)

Auxin to Gibberellin (600/300)

Auxin to Gibberellin (300/600)

Auxin to Gibberellin (0%)

Auxin to Gibberellin (600/300)

Auxin to Gibberellin (300/600)

Mycorrhiza (inoculation) Mycorrhiza (inoculation)

? FQ #29)N3!7! VW (

) Y

Referensi

Dokumen terkait