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Buketov
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Buketov
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Buketov
University
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Buketov
University
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Buketov
University
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1 Smelyansky I. Saigas are returning to the Russian Trans-Volga region / I. Smelyansky, V. Kirilyuk, S. Titova // Saiga news.
— 2022/23. — Iss.28. — P. 23-25.
2 A#>C2 &29@2%Y? ? f252a&12"$: 1Y @Y?Y7B1 %Y/Y@b, A$"4$ b Y $3 52B?/B=1 p$73$74. — [/$%17Y""49 7$&A7&]. — {$;b3 #Y&1AV2: https://www.inform.kz/ru/sud-ba-saygakov-v-kazahstane-chto-govoryat-ekologi-uchenye-i-o-chem-zayavlyayut- fermery_a3952746
3 sY/$$ 300 14&. VY&$?Y? VY172?/$"Y &29@2%23b. — [/$%17Y""49 7$&A7&]. — {$;b3 #Y&1AV2:
https://kz.kursiv.media/2022-07-07/bolee-300-tys-posevov-potravleno-sajgakami/.
4 o2#$$? `.j. 29@2% ? f252a&12"$ / `.j. o2#$$?, j.j. /At%b9. — j/32-j12: ~2A%2 f25{, 1982. — 160 &.
5 _b7"Y? z.`. `Y5?72$"b$ % ;b5"b: %Y/Y@bB, Ya72"2 b b&VY/>5Y?2"b$ &29@2%Y? / z.`. _b7"Y?. — y., z$&"2B V7Y- 34W/$""Y&1>, 1982. — 224 &.
6 /At%b9 j.j. 29@2% ? f252a&12"$ b V$7&V$%1b?4 $@Y V7Y34&/2 / j.j. /At%b9, `.j. o2#$$? // `$&1". j~ f25{,
— 1977. — 3. — . 28–32.
7 o2#$$? `.j. n7Y34&$/ &29@2%2 ? f252a&12"$ / `.j. o2#$$?, j.{. .22# // aY12 b YaY1"b>$ aY5B9&1?Y. — 1980. —
10. — .10–12.
8 Y%Y/Y? `.w. 29@2%. ob/Y@$"bB, &b&1$321b%2, %Y/Y@bB, Ya72"2 b b&VY/>5Y?2"b$ / `.w. Y%Y/Y?, z.`. _b7"Y?. — y.,1998. — 356 &.
9 ja3$#$"Y? f.y. z2"#W2p14 2V2#"Y@Y f252a&12"2: Y$7%b YC YC$%12a V7b7Y#"Y@Y "2&/$#bB / f.y. ja3$#$"Y?,
.f. {23252"Y?, q.j. fb"#$7C2$?2. — y.: n$7Y, 2015. — 250 &.
10 ~2tbY"2/>"49 #Y%/2# Y &Y&1YB"bb Y%7A;2=$9 &7$#4 b YC b&VY/>5Y?2"bb V7b7Y#"4a 7$&A7&Y? {$&VAC/b%b f252a-
&12" 52 2021 @Y#. — [/$%17Y""49 7$&A7&]. — {$;b3 #Y&1AV2: https:
//www.gov.kz/memleket/entities/ecogeo/documents/details/383692?lang=ru — 517 &.
Buketov
University
11 ~2&b$? s.~. t$"%2 &VY&YCY? b&VY/>5Y?2"bB V2&1Cb VY/AVA&14""Y9 5Y"4 2V2#"Y@Y f252a&12"2 / s.~. ~2&b$?,
~._. _2"212/2VY?, j.f. s$%%2/b$?, j.f. s$%%2/b$?2 // j@727"49 ?$&1"b% 72/2. — 2021. — 11 (214). — . 20–26. DOI:
10.32417/1997-4868-2021-214-11-20-26.
12 Nasiev B.N. The Impact of Pasturing Technology on the Current State of Pastures / B.N. Nasiev, j.K. Bekkaliev // Annals of Agri-Bio Research. — 2019. — Vol. 24. — No. 2. — P. 246–254.
13 2AC2$? s.q. {$&A7&"Y-%Y/Y@b$&%2B Yt$"%2 &Y&1YB"bB V$&2"4a V2&1Cb ~274"&%ba V$&%Y? / s.q. 2AC2$? // {$@b- Y". ?4V. «`$&1"b% yj~s». — 2008. — . 13, 5. — . 43–46.
14 y2a23C$1Y?Õy._. t$"%2 V7Yt$&&Y? ?Y&&12"Y?/$"bB #$@72#b7Y?2""4a %Y&b&1$3 j1472A&%Y9 YC/2&1b: #b&. … #-72 pb/Y&. "2A%: 6D060800 — %Y/Y@bB. — j/3214: f~j, 2016. — 152Õ&.
15 fA/b% f.~. !53$"$"b$ pb1Yt$"Y5Y? `Y/@Y-72/>&%Y@Y 3$;#A7$>B VY# ?/bB"b$3 V2&1Cb"4a "2@7A5Y%
/ f.~. fA/b%, s._. w&32@A/Y?2, .ä. fYW$/$?2, f.s. yAW2$?2, .. .b"%27$"%Y // `$&1". `r. $7. r$Y@72pbB, @$Y%Y/Y-
@bB. — 2016. — 4. — . 25–32.
16 .b"%27$"%Y .. $5Y""2B #b"23b%2 NDVI V2&1Cb"4a /2"#W2p1Y? $?$7"Y@Y n7b%2&VbB VY #2""43 MODIS / .. .b"%27$"%Y, .j. s2712/$? // Y?7$3$""4$ V7YC/$34 #b&12"tbY""Y@Y 5Y"#b7Y?2"bB $3/b b5 %Y&3Y&2. — 2020. —
. 17, 4. — . 179–194. DOI: 10.21046/2070-7401-2020-17-4-179-194.
17 fA/b% f.~. r$Yb"pY732tbY""49 2"2/b5 YVA&14"b?2"bB $?$7Y-2V2#"Y@Y n7b%2&VbB / f.~. fA/b%, `.!. n$17Y?,
`.r. äp$7$?, ~.j. %2$"%Y, .. .b"%27$"%Y // j7b#"4$ %Y&b&1$34. — 2020. — . 26, 2(83). — . 16–24. DOI:
10.24411/1993-3916-2020-10091
18 b1%Y?2 .s. f/b321b$&%b9 7AC$; ? &1$V"Y9 5Y"$ `Y&1Y"Y-w?7YV$9&%Y9 72?"b"4: b"#b%21Y74 b 7253$$"b$
/ .s. b1%Y?2, j.~. Y/Y1Y%74/b", `.`. `b"Y@72#Y?2 // !5?. {j~. $7. @$Y@72p. — 2020. — . 84, 6. — . 864–873. DOI:
10.31857/S2587556620050167
19 Robinson S. Pastoralists as optimal foragers? reoccupation and site selection in the deserts of post-Soviet Kazakhstan / S. Robinson, C. Kerven, R. Behnke, K. Kushenov, E.J. Milner-Gulland // Hum. Ecol. — 2017. — No. 45. — P. 5–21. DOI:
10.1007/s10745-016-9870-5
20 Kerven C. Pastoralism at Scale on the Kazakh Rangelands: From Clans to Workers to Ranchers / C. Kerven, S. Robinson, R. Behnke // Front. Sustain. Food Syst. — 2021. — 4:590401. DOI: 10.3389/fsufs.2020.590401
21 Dara A. Post-soviet land-use change affected fire regimes on the Eurasian steppes / A. Dara, M. Baumann, M. Freitag, N. Hölzel, P. Hostert, J. Kamp, D. Müller, B. Ullrich, T. Kuemmerle // Ecosystems — 2019. — No 23. — P. 943–956. DOI:
10.1007/s10021-019-00447-w
22 Dara A. Annual Landsat time series reveal post-soviet changes in grazing pressure / A. Dara, M. Baumann, M. Freitag, N. Hölzel, P. Hostert, J. Kamp, D. Müller, j. Prishchepov, T. Kuemmerle // Remote Sensing of Environment. — 2020. — Vol. 239.
— 111667 DOI: 10.1016/j.rse.2020. 111667
23 Kamp J. Persistent and novel threats to the biodiversity of Kazakhstan’s steppes and semi-deserts / J. Kamp, M.A. Koshkin, T.M. Bragina, T.E. Katzner, E.J. Milner-Gulland, D. Schreiber, R. Sheldon, A. Shmalenko, I. Smelansky, J. Terraube, R. Urazaliev // Biodivers. Conserv. — 2016. — Vol. 25. — {. 2521–2541. DOI:10.1007/s10531-016-1083-0
24 $/>&%Y$, /$&"Y$ b 74C"Y$ aY5B9&1?Y ? 2V2#"Y-f252a&12"&%Y9 YC/2&1b 2017–2021: &121. &C. — [/$%17Y""49 7$-
&A7&]. — {$;b3 #Y&1AV2: https://stat.gov.kz/region/253160/statistical_information/publication. — 117 &.
25 2V2"Y? y.f. %Y/Y@b$&%b$ VY&/$#&1?bB VY1$V/$"bB %/b3212 ? $?$7"Y3 n7b%2&Vbb / y.f.2V2"Y? // j7b#"4$
%Y&b&1$34. — 2018. — . 24, 1 (74). — . 20–31.
26 ~A7AW$? y._. n7YC/$34 b 3$1Y#4 &V2&$"bB &29@2%2 (Saiga tatarica L.) ? f252a&12"$ / y._. ~A7AW$?,
.j. s2912"2$? // s=//$1$"> 7$"CA7@&%Y@Y "2A"Y@Y t$"172 7 {j~. — 2018. — 1. — 19 c. — [/$%17Y""49 7$&A7&.
— {$;b3 #Y&1AV2: http://elmag.uran.ru:9673/magazine/Numbers/2018-1/Articles/ZNM-2018-1.pdf). DOI: 10.24411/2304-9081- 2018-11005
27 yb/"$7-rA//2"# .q;. t$"%2 #2""4a #b"23b%b tb%/Y? ? VYVA/Btbb &29@2%Y? / .q;. yb/"$7-rA//2"# // Saiga news. — 2009. — `4V. 9. — . 9, 10.
28 jC21A7Y? s.q. $5Y""2B #b"23b%2 %Y73Y?4a 7$&A7&Y? b Vb12"b$ &29@2%2 "2 $&1$&1?$""Y3 V2&1Cb$ ? VY/AVA&14"$
/ s.q. jC21A7Y?, s.!. n$17b$?, y.n. fY/$&"b%Y?, j.w. ACCY1b" // &V$ab &Y?7$3$""Y9 CbY/Y@bb. — 1998. — . 118, 5.
— . 564–584.
29 Buuveibaatar B. Factors affecting survival and cause-specific mortality of saiga calves in Mongolia / B. Buuveibaatar, J. Young, J. Berger, A. Fine, B. Lkhagvasuren, P. Zahler, T. Fuller // J. of Mammalogy. — 2013. — Vol. 94. — Iss. 1. — P. 127–
136.
30 White T.C.R. Experimental and observational evidence reveals that predators in natural environments do not regulate their prey: they are passengers, not drivers / T.C.R. White // Acta Oecologica. — 2013. — Vol. 53. — P. 73–87.
31 r72$? ä.j. 2b&/$""Y&1>, &17A%1A72 b ?Y&V7Yb5?Y#&1?Y VYVA/Btb9 &29@2%2 ? f252a&12"$ / ä.j. r72$?, j.y. y$/#$C$%Y?, j.s. s$%$"Y? // 1$V"Y9 C=//$1$">. — 2009. — 27. — . 47–50.
32 2V2"Y? y.f. `/bB"b$ V7b7Y#"Y-%/b321b$&%ba p2%1Y7Y? "2 b&/$""Y&1> &29@2%Y? (Saiga tatarica Pall.) ? `Y/@Y-
72/>&%Y3 3$;#A7$>$ / y.f. 2V2"Y? // nY?Y/;. %Y/Y@. ;A7". — 2016. — 4. — . 445–454.
33 Kock R. Mass Mortality Events: Publication of the Kazakh — British research team studies on saiga deaths / R. Kock, E.J. Milner-Gulland, S. Robinson, S. Zuther, M. Orynbayev // Saiga News. — 2018. — Iss. 23. — {. 24-26.
Buketov
University
34 Kock R.j. Saigas on the brink: multi-disciplinary analysis of the factors influencing a mass die-off event / R.A. Kock, y. Orynbayev, S. Robinson, S. Zuther, N.J. Singh, W. Beauvais, E.R. Morgan, A. Kerimbayev, S. Khomenko, H.M. Martineau, R. Rystaeva, Z. Omarova, S. Wolfs, F. Hawotte, J. Radoux, E.J. yilner-Gulland // Science Advances. — 2018. — Vol. 4. — Iss. 1.
— P.1-10. DOI: 10.1126/sciadv. aao2314
35 j9%b3C2$? y.j. &/A2Ba ?4#$/$"bB V2&1$7$//$52 Y1 &29@2%Y? ? p$?72/$–3271$ 1984 @Y#2 ? 72/>&%Y9 YC/2&1b / y.j. j9%b3C2$?, !.z. y271b"$?&%b9, j.j. j/1AaY?, .!. !?2"Y?, `.o. A7Y? // !5?. j~ f25{. — 1985. — 4. — .
39–41.
36 jC&21b7Y? r.r. {$5A/>1214 %Y3V/$%&"Y@Y %Y/Y@Y-Vb5YY1Y/Y@b$&%Y@Y 3Y"b1Y7b"@2 V7bb" 32&&Y?Y9 @bC$/b &29@2-
%Y? / r.r. jC&21b7Y?, j.j. b#Y7A%, .s. 2AC2$?, f._. fAW2/b$?, f.w. yA752C2$?, y.r. f2%bW$?, o.³. ~A7;2"Y?2,
~.. rb"2B1Y? // {Y&. ?$1$7b"27. ;A7". y$/%b$ #Y32W"b$ b #b%b$ ;b?Y1"4$. — 2013. — 5. — . 22–25.
37 yb$"%Y j.`. n7YC/$32 32&&Y?Y9 @bC$/b &29@2%Y? / j.`. yb$"%Y, `.j. yb$"%Y, j.f. f272A/Y?, j.`. nY1$ab", j.n. y$;"$? // `$1$7b"27bB &$@Y#"B. — 2016. — 4 (19). — . 40–45.
38 Orynbayev M.B. Seroprevalence of infectious diseases in saiga antelope (Saiga tatarica tatarica) in Kazakhstan 2012–2014 / M.B. Orynbayev, W. Beauvais, A.R. Sansyzbaya, R.A. Rystaevaa, K.T. Sultankulovaa, A.A. Kerimbaeva, M.N. Kospanovaa, R.A. Kock // Preventive Veterinary Medicine. — 2016. — Vol. 127. — P. 100-104. DOI: 10.1016/j.prevetmed.2016.03.016
39 Orynbayev M.B. Biological characterization of Pasteurella multocida present in the saiga population / M.B. Orynbayev, K.T. Sultankulova // BMC Microbiology — 2019. — 19:37. DOI:10.1186/s12866-019-1407-9
40 Robinson S. Opportunistic bacteria and mass mortality in ungulates: lessons from an extreme event / S. Robinson, E.J. Milner-Gulland, Y. Grachev, A. Salemgareyev, M. Orynbayev, A. Lushchekina, E. Morgan, W. Beauvais, N. Singh, S. Khomenko, R. Cammack, R. Kock // Ecosphere. — 2019. — Vol. 10. — Iss. 6. DOI: 10.1002/ecs2.2671
41 2V2"Y? y.f. Y&1YB"b$ VY@Y/Y?>B b V7bb"4 @bC$/b &29@2%Y? ? $?$7"Y3 n7b%2&Vbb / y.f.2V2"Y? // Selevinia.
YY/. $;$@Y#"b% f252a&12"2 b $"172/>"Y9 j5bb. — 2015. — . 23. — . 194–197.
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Buketov
University
M.K. Sapanov, K.M. Akhmedenov
Natural and historical prerequisites for saiga hunting in Kazakhstan
The number of saigas in Kazakhstan's Ural group, which inhabits the Volga-Ural interfluve, increased to a record 801,000 in 2022. Local farmers claim that these animals severely devastate agricultural fields and pas- tures and demand artificial reduction of the population. At the same time, nature conservation organizations suggest their further protection. In this study, we analyzed the literature and our own results on the dynamics of saiga numbers and their spring mass mortality in conjunction with changes in the natural and climatic con- ditions of the region. Industrial shooting of saigas, the Ural saiga group, began in 1954 and continued inter- mittently until 1994. In other words, saiga hunting was a routine regulatory activity at that time. Then there was a long-term period of depression in their numbers, which was attributed to excessive shooting and poach- ing. It has been shown that the wave-like dynamics of saiga population numbers depends entirely on the productivity of the grass stand and the watering of their habitat. A deep and prolonged depression in the num- ber of animals in the 2000s resulted from a severe reduction in watering places, as during this period, due to a 14-year absence of surface runoff of spring meltwater into the local hydrographic network, ponds and sor- rows dried up everywhere and lakes and rivers shallowed. It was only in 2010-2011 that new recharge of open reservoirs with melt water began, and since then they have never dried out until 2022. It was at that time that the saiga population began to increase rapidly. The reasons for the periodic mass selective spring mortali- ty of saigas were analyzed. It is shown that this phenomenon could be caused by a noncontagious disease, timpania, which is identified by the rapidity of the disease, which explains its selective nature (mainly seden- tary females giving birth perish). Timpania, apparently, could serve as a trigger mechanism for triggering other secondary diseases in animals, including contagious pasteurellosis, which is constantly registered in dead animals. The number of animals after the mass saiga mass mortality in spring recovers in a few years to the previous level due to high fecundity, both in periods of depression and high numbers. Therefore, there is no reason to cancel the industrial shooting of such a valuable self-renewable resource, taking into account the dynamics of its abundance under the influence of natural and climatic condition.
Keywords: saigas, Ural grouping, population dynamics, fodder base, area watering, spring mortality, timpani, animal fishery.
References
1 Smelyansky, I., Kirilyuk, V. & Titova, S. (2022/23). Saigas are returning to the Russian Trans-Volga region. Saiga news, 28, 23-25.
2 Sudba saigakov v Kazakhstane: chto govoriat ekologi, uchenye i o chem zaiavliauit fermery [The fate of saigas in Kazakh- stan: what ecologists, scientists say and what farmers say]. Retrieved from https://www.inform.kz/ru/sud-ba-saygakov-v-kazahstane- chto-govoryat-ekologi-uchenye-i-o-chem-zayavlyayut-fermery_a3952746 [in Russian].
3 Bolee 300 tysiach posevov potravleno saigakami [Over 300,000 crops were damaged by saigas]. Retrieved from https://kz.kursiv.media/2022-07-07/bolee-300-tys-posevov-potravleno-sajgakami/ [in Russian].
4 Fadeev, V.A. & Slutsky, A.A. (1982). Saigak v Kazakhstane [Saiga in Kazakhstan]. Almaty: Izdatelstvo «Nauka»
Kaz2khskoi SSSR [in Russian].
5 Zhirnov, L.V. (1982). Vozvrashchenie k zhizni: Ekologiia, okhrana i ispolzovanie saigakov [Return to Life: Ecology, Con- servation and Use of the Saiga]. Moscow: Lesnaia promyshlennost [in Russian].
6 Slutsky, A.A., & Fadeev, V.A. (1977). Saigak v Kazakhstane i perspektivy ego promysla [Saiga antelope in Kazakhstan and prospects for its fishery]. Vestnik Akademii nauk Kazakhskoi SSR — Bulletin of the Academy of Sciences of Kazakh SSR, 3, 28–32 [in Russian].
7 Fadeev, V.A., & Schaad, A.R. (1980). Promysel saigaka v Kazakhstane [Saiga hunting in Kazakhstan]. Okhota i okhotniche khoziaistvo —Hunting and hunting economy, 10, 10–12 [in Russian].
8 Sokolov, V.E., & Zhirnov, L.V. (1998). Saigak. Filogeniia, sistematika, ekologiia, okhrana i ispolzovanie [Saiga. Phylogeny, taxonomy, ecology, protection and use]. Moscow [in Russian].
9 Akhmedenov, K.M., Ramazanov, S.K., & Kinderbaeva, D.A. (2015). Landshafty Zapadnogo Kazakhstana: ocherki ob obektakh prirodnogo naslediia [Landscapes of Western Kazakhstan: essays on natural heritage sites]. Moscow: Pero [in Russian].
10 Natsionalnyi doklad o sostoianii okruzhaiushchei sredy i ob ispolzovanii prirodnykh resursov Respubliki Kazakhstan za 2021 god [National report on the state of the environment and on the use of natural resources of the Republic of Kazakhstan for 2021].
Retrieved from https://www.gov.kz/memleket/entities/ecogeo/documents/details/383692?lang=ru [in Russian].
11 Nasiev, B.N., Zhanatalapov, N.Zh., Bekkaliev, A.K. & Bekkalieva, A.K. (2021). Otsenka sposobov ispolzovaniia pastbishch polupustynno·y zony Zapadnogo Kazakhstana [Assessment of ways to use pastures in semi-desert zone of West Kazakhstan].
Agrarnyi vestnik Urala — Agrarian Bulletin of the Urals, 11(214), 20-26. DOI: 10.32417/1997-4868-2021-214-11-20-26 [in Rus- sian].
