Anguilliformes) of Bago River and Adjacent Waters in Negros

The Anguilliform Eels (Pisces: Anguilliformes) of Bago River and Adjacent Waters in Negros Occidental, Philippines

Abner A. Bucol

Silliman University Angelo King Center for Research and Environmental Management, Dumaguete City

Carmen C. Menes Joji D. Linaugo

La Consolacion College-Bacolod, Bacolod City

Jessica O. Pacalioga

Biology Department, University of St. La Salle, Bacolod City

The Eels (Order Anguilliformes) occurring in Bago River and BEKBDFOU XBUFST JO /FHSPT 0DDJEFOUBM 1IJMJQQJOFT BSF CSJFnZ annotated. A total of 21 species in six families are included in this paper. Snake-eels (Ophichthidae) consist of 11 species, moray Eels (Muraenidae) are represented by four species, Conger Eels (Congridae) are represented by three species, while only two species belong to the family Muraenesocidae and the Freshwater Eels (Anguillidae) and Spaghetti Eels (Moringuidae) are each represented by a single species.

KEYWORDS: eels, river, brackish water, freshwater, Negros Occidental

INTRODUCTION

A

—ž’••’˜›–ȱŽŽ•œȱŠ›ŽȱŒ‘Š›ŠŒŽ›’£Žȱ‹¢ȱ‘ŠŸ’—ȱŽ•˜—ŠŽǰȱœ—Š”ŽȬ

•’”Žȱ˜›ȱ ˜›–Ȭ•’”Žȱ‹˜’Žœȱǻ–’‘ȱǭȱŒ˜œ”Ž›ǰȱŗşşşǼȱŠ—ȱŠ••ȱ

‘ŠŸŽȱ•Ž™˜ŒŽ™‘Š•’Œȱ•Š›ŸŠ•ȱœŠŽȱǻ–’‘ǰȱŗşŝşDzȱ–’‘ȱǭȱŠœ•Žǰȱ ŗşŝŘDzȱŠœ•ŽǰȱŗşŜŜǰȱŗşŜŞDzȱ’••Ž›ȱǭȱœž”Š–˜˜ǰȱŘŖŖŚǼǯȱ

‘ŽȱŽŽ•œȱ’—ȱ‘Žȱ‘’•’™™’—Žœȱ‘ŠŸŽȱ‹ŽŽ—ȱœž’Žȱ‹¢ȱœŽŸŽ›Š•ȱŠž‘˜›œȱ

’—Œ•ž’—ȱ •ŽŽ”Ž›ȱ ǻŗŞŜŚǼǰȱ Š—ȱ Ž››Žȱ ǻŗşŘřǰȱ ŗşŘŚǼǯȱ –’‘ȱ ǻŘŖŖŚǼȱ

›Ž™˜›Žȱ ˜—ȱ ‘Žȱ ¢™Žȱ –ŠŽ›’Š•œȱ ˜ȱ ŽŽ•œȱ Ž›’ŸŽȱ ›˜–ȱ ‘Žȱ Ž¡Ž—œ’ŸŽȱ

Œ˜••ŽŒ’˜—ȱ–ŠŽȱ‹¢ȱ‘Žȱ•‹Š›˜œœȱ¡™Ž’’˜—ȱ›˜–ȱŗşŖŝȱ˜ȱŗşŗŖȱǻ–’‘ȱ ǭȱ’••’Š–œǰȱŗşşşǼǯȱ

ȱŽ’——’—ȱ’—ȱ‘Žȱ•ŠŽȱŗşşŖœǰȱœŽŸŽ›Š•ȱ—Ž ȱœ™ŽŒ’Žœȱ˜ȱŽŽ•œȱ‘ŠŸŽȱ‹ŽŽ—ȱ

ŠŽȱ ˜ȱ ‘Žȱ ꜑ȱ Šž—Šȱ ˜ȱ ‘Žȱ ‘’•’™™’—Žœȱ ǻŒ˜œ”Ž›ǰȱ ŗşşŞǰȱ ŘŖŗŖDzȱ

Šœ•Žȱ ǭȱ –’‘ǰȱ ŗşşşDzȱ 㑕”Žǰȱ ŘŖŖŖDzȱ 㑕”Žȱ ǭȱ –’‘ǰȱ ŘŖŖŘDzȱ –’‘ȱ ǭȱ Š›–˜Ÿœ”Š¢Šǰȱ ŘŖŖřǼǯȱ ŽŸŽ›Š•ȱ ž—”—˜ —ȱ œ™ŽŒ’Žœȱ ‘Šȱ ‘ŠŸŽȱ ‹ŽŽ—ȱ

’œŒ˜ŸŽ›Žȱ›ŽŒŽ—•¢ȱœ’••ȱŠ Š’ȱ˜›–Š•ȱŽœŒ›’™’˜—ȱ ‘’•ŽȱŒŽ›Š’—ȱ›˜ž™œȱ

œžŒ‘ȱ Šœȱ –˜›’—ž’œȱ Š—ȱ ˜™‘’Œ‘‘’œȱ —ŽŽȱ Šȱ ‘˜›˜ž‘ȱ Š¡˜—˜–’Œȱ

›ŽŸ’œ’˜—ȱǻǯ ǯȱ–’‘ȱǭȱǯǯȱŒ˜œ”Ž›ǰȱ™Ž›œǯȱŒ˜––ǯǼǯȱ˜›ȱŽ¡Š–™•Žǰȱ‘Žȱ

›ŽŒŽ—•¢ȱ ŽœŒ›’‹ŽȱAnguilla huangiȱ ǻA. luzonensisǼȱ ›˜–ȱ Š’ Š—ȱ Š—ȱ

‘Žȱ—˜›‘Ž›—ȱ‘’•’™™’—Žœȱ ŠœȱŒ˜—žœŽȱ ’‘ȱA. celebesensisȱ˜›ȱŠȱ•˜—ȱ

™Ž›’˜ȱ˜ȱ’–ŽȱǻŽ—ǰȱ’—ȱǭȱ£Ž—ǰȱŘŖŖşǼǯȱ

ȱ’œȱ‘Žȱ™ž›™˜œŽȱ˜ȱ‘’œȱ™Š™Ž›ȱ˜ȱ™›ŽœŽ—ȱ‘Žȱœ™ŽŒ’Žœȱ˜ȱŽŽ•œȱ‘žœȱ

Š›ȱ”—˜ —ȱ›˜–ȱŠ˜ȱ’ŸŽ›ǰȱŽ›˜œȱŒŒ’Ž—Š•ȱŠ—ȱŠ“ŠŒŽ—ȱ ŠŽ›œǯȱ

MATERIALS AND METHODS

ŽȱŒ˜••ŽŒŽȱœŠ–™•Žœȱ˜ȱŽŽ•œȱ–Š’—•¢ȱ›˜–ȱ‘Žȱ˜ž›ȱŒ˜••ŽŒ’—ȱœŠ’˜—œȱ

’—ȱŠ˜ȱ’ŸŽ›ǰȱŽ›˜œȱŒŒ’Ž—Š•ȱǻ’ž›ŽȱŗǼȱžœ’—ȱŠȱŸŠ›’Ž¢ȱ˜ȱŽŠ›ȱ

œžŒ‘ȱ Šœȱgarabȱ ǻŠ—ȱ ’—’Ž—˜žœȱ ŽŠ›Ǽǰȱ œ™ŽŠ›œǰȱ ꗎȬ—Žœȱ Š—ȱ ‹Š–‹˜˜ȱ

Figure 1. Map showing the Bago River in Negros Occidental.

›Š™œǯȱ’’˜—Š•ȱœŠ–™•Žœȱ™ž›Œ‘ŠœŽȱ’›ŽŒ•¢ȱ›˜–ȱ•˜ŒŠ•ȱ꜑Ž›œȱŠ—ȱ

•˜ŒŠ•ȱ꜑ȱ–Š›”Žœȱ’—ȱ—Ž’‘‹˜›’—ȱ•˜ŒŠ•’’Žœȱ Ž›ŽȱŠ•œ˜ȱ’—Œ•žŽȱ’—ȱ‘’œȱ

œž¢ǯ

Š–™•Žœȱ  Ž›Žȱ ’––Ž’ŠŽ•¢ȱ ę¡Žȱ ’—ȱ ŗŖƖȱ ˜›–Š•’—ȱ Š—ȱ ‘Ž—ȱ ’—ȱ ŝŖƖȱ Ž‘Š—˜•ȱ ŠŽ›ȱ Šȱ •ŽŠœȱ Šȱ  ŽŽ”ȱ ’—ȱ ˜›–Š•’—ȱ Š—ȱŽ™˜œ’Žȱ Šȱ ‘Žȱ ǯǯȱ ˜—£Š•Žœȱ žœŽž–ȱ ˜ȱ Šž›Š•ȱ ’œ˜›¢ȱ Šȱ ’••’–Š—ȱ —’ŸŽ›œ’¢ǰȱ

ž–ŠžŽŽȱ’¢ǯȱ

—ȱ Š’’˜—ǰȱ œ™ŽŒ’–Ž—œȱ ›˜–ȱ ‘Žȱ Š“ŠŒŽ—ȱ  ŠŽ›œȱ œžŒ‘ȱ Šœȱ ‘˜œŽȱ

Œ˜••ŽŒŽȱ’—ȱ ž’–Š›Šœȱ›Š’ȱŠ—ȱ’—ȱ•˜’•˜ȱ™›˜Ÿ’—ŒŽȱ‘ŠȱŠ›Žȱ™›ŽœŽ—•¢ȱ

‘˜žœŽȱ’—ȱ‘Žȱ‘’•’™™’—ŽȱŠ’˜—Š•ȱžœŽž–ȱǻǼȱŠ—ȱ Ž›ŽȱŽ¡Š–’—Žȱ

‹¢ȱ‘ŽȱœŽ—’˜›ȱŠž‘˜›ȱ˜—ȱřŖȱŽ™Ž–‹Ž›ȱŘŖŖşȱ Ž›ŽȱŠ•œ˜ȱ’—Œ•žŽǯȱŽ›Š’—ȱ

œ™ŽŒ’Žœȱ ”—˜ —ȱ ’—ȱ ‘Žȱ Š“ŠŒŽ—ȱ  ŠŽ›œȱ •’œŽȱ ’—ȱ ™ž‹•’œ‘Žȱ •’Ž›Šž›Žȱ  Ž›ŽȱŠ•œ˜ȱ’—Œ•žŽȱ’—ȱ‘Žȱœ™ŽŒ’Žœȱ•’œǯ

˜›ȱ ŸŽ›’ęŒŠ’˜—ǰȱ ™‘˜˜›Š™‘œȱ  Ž›Žȱ ˜‹Š’—Žȱ ˜›ȱ ŽŠŒ‘ȱ œ™ŽŒ’Žœǰȱ

Š—ȱ‘Ž—ȱœŽ—ȱ˜ȱŽŽ•ȱœ™ŽŒ’Š•’œœȱǻǯ ǯȱ–’‘ȱ’—ȱ‘Žȱ’Ÿ’œ’˜—ȱ˜ȱ’œ‘Žœǰȱ

—’Žȱ ŠŽœȱ Š’˜—Š•ȱ žœŽž–ȱ ˜ȱ Šž›Š•ȱ ’œ˜›¢ǰȱ –’‘œ˜—’Š—ȱ

—œ’ž’˜—ȱŠ—ȱŒ˜œ”Ž›ǰȱǯǯȱ˜ȱ‘ŽȱŠ•’˜›—’ŠȱŒŠŽ–¢ȱ˜ȱŒ’Ž—ŒŽœǰȱ Ǽǯȱž™•’ŒŠŽȱœŠ–™•Žœȱ Ž›Žȱ•ŠŽ›ȱ˜—ŠŽȱ˜ȱ‘Žȱȱ‘›˜ž‘ȱ

›ǯȱ Žěȱ ’••’Š–œǯȱ œ’Žȱ ›˜–ȱ ‘Žȱ ’Ž—’ęŒŠ’˜—ȱ ™›˜Ÿ’Žȱ ˜ȱ žœȱ ‹¢ȱ

œ™ŽŒ’Š•’œœȱ˜—ȱŽŽ•œǰȱ Žȱ‘ŠŸŽȱŠ•œ˜ȱž’•’£Žȱ‘Žȱ˜••˜ ’—ȱœ˜ž›ŒŽœDZȱ–’‘ȱ

Š—ȱŒ˜œ”Ž›ȱǻŗşşşǼȱŠ—ȱŒ˜œ”Ž›ȱǻŘŖŗŖǼȱ˜›ȱ˜™‘’Œ‘‘’œDzȱ㑕”Žǰȱ

Œ˜œ”Ž›ȱ ǭȱ –’‘ȱ ǻŗşşşǼȱ ˜›ȱ –ž›ŠŽ—’œǰȱ Š—ȱ –’‘ȱ ǻŗşşşǼȱ ˜›ȱ

–˜›’—ž’œȱŠ—ȱŒ˜—›’œǯ

RESULTS AND DISCUSSION List of species

‘Žȱ›ŽœŽŠ›Œ‘ȱŽ—Ž›ŠŽȱŠ—ȱŠ——˜ŠŽȱ•’œȱ˜ȱŘŗȱŠ—ž’••’˜›–ȱŽŽ•ȱœ™ŽŒ’Žœȱ

Œ˜••ŽŒŽȱ›˜–ȱŠ˜ȱ’ŸŽ›ȱŠ—ȱ‘ŽȱŠ“ŠŒŽ—ȱ ŠŽ›œȱ˜ȱ•˜ȱ’ŸŽ›ȱŠ—ȱ ž’–Š›Šœȱ›Š’ǯȱŠ“˜›’¢ȱŠ›Žȱ”—˜ —ȱ‹ž››˜ Ž›œȱ’—‘Š‹’’—ȱŽœžŠ›’—Žȱ

Š›ŽŠœȱ‘Šȱ‹Ž•˜—ȱ˜ȱ™‘’Œ‘‘’ŠŽȱǻŗŖȱœ™ŽŒ’ŽœǼȱŠ—ȱ˜›’—ž’ŠŽȱǻ˜ž›ȱ

œ™ŽŒ’ŽœǼȱ Š–’•’Žœǯȱ ˜–Žȱ ˜ȱ ‘Žȱ ŽŽ•ȱ œ™ŽŒ’Žœȱ —ŽŽȱ ž›‘Ž›ȱ Š¡˜—˜–’Œȱ

ŠĴŽ—’˜—Dzȱœ˜–Žȱ˜ȱ‘ŽœŽȱŠœ™ŽŒœȱŠ›Žȱ‹›’ŽĚ¢ȱ’œŒžœœŽǯ

ORDER ANGUILLIFORMES

FAMILY ANGUILLIDAE (Freshwater Eels)

Anguilla marmorataȱž˜¢ȱŠ—ȱ Š’–Š›ȱŗŞŘŚȱǻ’ž›ŽȱŘǼ

Specimens:ȱ ȱŗŖŖśŗȬŗŖŖśŘ

Remarks:ȱ’œ’—ž’œ‘ŽȱŽŠœ’•¢ȱ›˜–ȱ˜‘Ž›ȱAnguillaȱ‹¢ȱ’œȱ–˜Ĵ•ŽȱŒ˜•˜›ȱ

Š—ȱ•˜—ȱ˜›œŠ•ȱę—ȱ ‘’Œ‘ȱ˜›’’—ŠŽœȱŒ•˜œŽ›ȱ˜ȱ‘Žȱ’••ȱ˜™Ž—’—ȱ‘Š—ȱ‘Žȱ

Š—žœȱǻ–’‘ȱŗşşşŠǼǯȱ‘Žȱ•Š›Žœȱ’—’Ÿ’žŠ•ȱ‘Šȱ ŽȱŒŠž‘ȱǻŠ’˜—ȱŗȱ’—ȱ

˜—ȱŠ•ŸŠ˜›ȱŽ—Ž’Œ˜Ǽȱ–ŽŠœž›ŽœȱŒŠȱؖȱŠ—ȱ Ž’‘œȱŗŚȱ”ǯȱ

Figure 2. Giant Mottled Eel Anguilla marmorata from Bago River (weight = 14 Kg).

Figure 3. The worm eel Moringua raitaborua from Bago River estuary.

ȱ ȱǻ™Š‘ŽĴ’ȱŽ•œǼ Moringua raitaboruaȱŠ–’•˜—ȱŗŞŘŘȱǻ’ž›ŽȱřǼ

Specimens:ȱ ȱŗŖŖřŖȬśŖDzȱŠ˜ȱ’ŸŽ›ȱœžŠ›¢Dzȱ˜••DZȱǯȱžŒ˜•ǰȱǯȱ

Ž—Žœǰȱǯȱ’—Šž˜ǰȱŠ—ȱǯȱŠ•ŽœDzȱŒǯȱŗśǰȱŘŖŖşǯ

Remarks:ȱ—˜Ž—Ž’ŒȱŠ—ȱœŽ¡žŠ•ȱŸŠ›’Š’˜—œȱǻ–’‘ȱǭȱŠœ•ŽǰȱŗşŝŘDzȱ

–’‘ǰȱ ŗşşşǼȱ ‘ŠŸŽȱ ŒŠžœŽȱ Š¡˜—˜–’Œȱ Œ˜—žœ’˜—ȱ  ’‘’—ȱ ‘Žȱ Ž—žœȱ Moringuaǯȱ ˜–Žȱ ˜ȱ ‘Žȱ œ™ŽŒ’Žœȱ ‘ŠŸŽȱ ‹ŽŽ—ȱ Šœœ’—Žȱ Ž››˜—Ž˜žœ•¢ȱ ˜ȱ

’쎛Ž—ȱŠ–’•’ŽœȱŠ—ȱŽ—Ž›Šǯȱ‘Žȱ’––Šž›Žȱ˜›–œȱ˜ȱMoringua were

ŽœŒ›’‹Žȱ–Š—¢ȱ ’–Žœȱ’—ȱ ‘Žȱ •’Ž›Šž›Žȱ ŠœȱApthalmichthys, Stilbiscus,

Š—ȱAnguillichthysȱǻ ˜œ•’—Žȱǭȱ›Šœ‹ž›ǰȱŗşśŜDzȱ ˜›˜—ǰȱŗşśŚDzȱŠœ•Žǰȱ ŗşŜŞDzȱŠœ•Žȱǭȱ–’‘ǰȱŗşŝŘǼǯȱ

‘Žȱ ŠŒžŠ•ȱ —ž–‹Ž›ȱ ˜ȱ œ™ŽŒ’Žœȱ ˜ȱMoringuaȱ ‘Šœȱ —˜ȱ ¢Žȱ ‹ŽŽ—ȱ

ŽŽ›–’—Žǯȱ ž–Ž›˜žœȱ —Š–Žœȱ ǻŠ‹˜žȱ řŞǼȱ ‘ŠŸŽȱ ‹ŽŽ—ȱ ™ž‹•’œ‘Žȱ ’—ȱ

‘Žȱ•’Ž›Šž›ŽȱǻœŽŽȱŠœ•ŽǰȱŗşŜŞǼǰȱ‹žȱ‘ŽȱŠŒžŠ•ȱ—ž–‹Ž›ȱ˜ȱœ™ŽŒ’Žœȱ’œȱ

™›˜‹Š‹•¢ȱœ–Š••ȱǻ–’‘ǰȱŗşşşǰȱŘŖŗŖȱ™Ž›œǯȱŒ˜––ǼǯȱŽ››ŽȱǻŗşŘŚǼȱ™ž‹•’œ‘Žȱ

—˜–’—Š•ȱ—Š–ŽœȱœžŒ‘ȱŠœȱMoringua cagayanaȱŠ—ȱM. robustaǯȱȱŠ™™ŽŠ›œȱ that M. robustaȱ–’‘ȱ‹ŽȱŠȱ–Šž›’—ȱŽ–Š•Žȱ˜ȱM. raitaboruaǯȱ˜ ŽŸŽ›ǰȱ

‘ŽȱŽ—’›ŽȱŽ—žœȱ—ŽŽœȱ›ŽŸ’œ’˜—ȱŠ—ȱŸŠ›’Š’˜—œȱ ’‘’—ȱ‘ŽȱŽ—žœȱ ’••ȱ

‹Žȱ ’œŒžœœŽȱ •ŠŽ›ǯȱ Š›Žž•ȱ Ž¡Š–’—Š’˜—ȱ ˜ȱ ‘Žȱ ‹˜¢ȱ ™›˜™˜›’˜—œǰȱ

ŸŽ›Ž‹›Š•ȱ Œ˜ž—œǰȱ ŽŸŽ•˜™–Ž—Š•ȱ œŠŽœȱ Š—ȱ œŽ¡žŠ•ȱ ’–˜›™‘’œ–ȱ

œžŽœȱ‘ŽȱœŠ–™•Žœȱ Žȱ˜‹Š’—ŽȱŠ›ŽȱMoringua raitaborua.

ŽŸŽ›Š•ȱ œ™ŽŒ’–Ž—œȱ ˜ȱMoringuaȱ Š›Žȱ ™›ŽœŽ—•¢ȱ Ž™˜œ’Žȱ ’—ȱ ‘Žȱ ǰȱ ‹žȱ ‘Žȱ ›ŽœŽŠ›Œ‘Ž›œȱ  Ž›Žȱ ž—Š‹•Žȱ ˜ȱ Ž¡Š–’—Žȱ ‘Ž–ǯȱ–˜—ȱ

‘ŽœŽȱŠ›Žȱœ™ŽŒ’–Ž—œȱŒ˜••ŽŒŽȱ‹¢ȱǯȱ’••’Š–œȱŽȱŠ•ǯȱ’—ȱŗşşśȱŠȱ ž’–Š›Šœȱ ǻȱ řŚřşŞŞǼȱ Š—ȱ Š—ȱ ˜Ššž’—ǰȱ •˜’•˜ȱ ™›˜Ÿ’—ŒŽȱ ǻȱ řŚřşŞŝǼǰȱ

‹¢ȱ ǯȱ Ž••Ž›œȱ ’—ȱ ŗşŘşȱ ž›’—ȱ ‘Žȱ ǯǯȱ ŠŸ¢ȱ Œ•’™œŽȱ ¡™Ž’’˜—ȱ Šȱ

’Œ˜›’Šœǰȱ Ž›˜œȱ ŒŒ’Ž—Š•ȱ ǻȱ ŗŗŘŝŝřǼǰȱ ‹¢ȱ ‘Žȱ •‹Š›˜œœȱ

‘’•’™™’—Žȱ¡™Ž’’˜—ȱǻŗşŖŝȬŗşŗŖǼȱ’—ȱŗşŖŞȱŠȱŠœ˜ȱǯǰȱŠ—Š¢ȱǻȱ ŗřśŗŖŚǼȱŠ—ȱ’—ȱ•˜’•˜ȱŠ›”Žȱ’—ȱŗşŖşȱǻȱŗřśŗŖŞǼǯȱ—ȱ’––Šž›Žȱ Moringuaȱ Ž™˜œ’Žȱ ’—ȱ ȱ –ŽŠœž›’—ȱ ŗŝŖȱ ––ȱ ǻǼȱ Œ˜••ŽŒŽȱ ‹¢ȱ

’••˜œ˜ȱŽȱŠ•ǯȱ’—ȱŽ‹›žŠ›¢ȱŗşŝśȱ›˜–ȱ˜Šœȱœ•Š—ǰȱ—’—’Ȭ¢ǰȱ—’šžŽǰȱ  Šœȱ˜ž—ȱŠ–˜—ȱGymnothoraxȱœ™ŽŒ’–Ž—œȱǻȱŗŖŝśŖǼǯ

Ž››Žȱ ǻŗşŘřǼȱ ›Ž™˜›Žȱ Apthalmichthys macrocephalusȱ •ŽŽ”Ž›ǰȱ

™›˜‹Š‹•¢ȱŠ—ȱ’––Šž›Žȱœ™ŽŒ’–Ž—ȱ˜ȱMoringua raitaboruaǰȱ›˜–ȱž‹˜Œȱ

ŽŠŒ‘ǰȱ•˜’•˜ǯȱȱȱȱ

FAMILY CONGRIDAE (Conger Eels) Bathymyrus œ™ǯ

Specimen:ȱ ȱŗŖŖŜśDzȱŠ‹Š—”Š•Š—ȱ’œ‘ȱŠ›”ŽDzȱǯȱŠŒŠ•’˜Š

Uroconger lepturusȱǻ’Œ‘Š›œ˜—ǰȱŗŞŚśǼ

Specimens:ȱ ȱ ŗŖŖśřDzȱ Š‹Š—”Š•Š—ȱ ’œ‘ȱ Š›”Žǰȱ Ž›˜œȱ

ŒŒ’Ž—Š•DzȱŒ˜••DZȱǯȱŠŒŠ•’˜Šǯȱ

Remarks:ȱǯȱžŒ˜•ȱ˜‹Š’—ŽȱŠȱœ’—•Žȱœ™ŽŒ’–Ž—ȱ›˜–ȱ˜˜ȱ’œ‘ȱŠ›”Žȱ

’—ȱ Œǯȱ ŘŖŖşǯȱ Ž››Žȱ ǻŗşŘřǼȱ ›Ž™˜›Žȱ œŠ–™•Žœȱ Œ˜••ŽŒŽȱ ›˜–ȱ ž‹˜Œȱ

ŽŠŒ‘ǰȱ•˜’•˜ǯȱ

˜—Ž›ȱŒ’—Ž›Žžœȱû™™Ž••ǰȱŗŞŘŞ

Specimens:ȱ˜ž›ȱž—•Š‹Ž•Žȱœ™ŽŒ’–Ž—œȱŠȱȱǻŗŖŜŜŜǼȱŒ˜••ŽŒŽȱ›˜–ȱ

˜Šœȱœ•Š—ǰȱ—’—’Ȭ¢ǰȱ—’šžŽǰȱŠ—Š¢ȱ’—ȱŽ‹›žŠ›¢ȱŗşŝśǯȱ Remarks:ȱ›˜‹Š‹•¢ȱŒ˜––˜—ȱ’—ȱ ž’–Š›Šœȱ›Š’ǯ

FAMILY MURAENESOCIDAE (Pike Congers) Muraenesox cinereusȱǻ˜›œœ”ª•ǰȱŗŝŝśǼȱǻ’ž›ŽȱŚǼ

Specimens:ȱ˜––˜—•¢ȱœ˜•ȱŠœȱ‹¢ȬŒŠŒ‘ȱ’—ȱ꜑ȱ–Š›”Žœǯ

Remarks:ȱ˜–Ž’–ŽœȱŒŠž‘ȱ‹¢ȱ꜑Ž›œȱ—ŽŠ›ȱ‘Žȱ–˜ž‘œȱ˜ȱ•˜ȱŠ—ȱ

Š˜ȱ›’ŸŽ›œǯ

Muraenesox bagioȱǻŠ–’•˜—ǰȱŗŞŘŘǼȱǻ’ž›ŽȱśǼ

Specimens:ȱ ȱŗŖŖśŚȱǻ ˜ȱœŠ–™•ŽœǼDzȱ•˜ȱ’ŸŽ›ȱœžŠ›¢

Figure 4. Head region of the Pike-conger eel Muraenesox cinereus.

FAMILY OPHICHTHIDAE (Snake or Worm Eels) Subfamily Myrophinae

Neenchelys œ™ǯȱ

Specimens:ȱ ȱŗŖŖśś

Remarks:ȱ™ŽŒ’–Ž—œȱ Ž›Žȱ˜ž—ȱ’—ȱ‘ŽȱŽœžŠ›’—ŽȱŠ›ŽŠȱ˜ȱŠ˜ȱ’ŸŽ›ǯȱ

ŽŒŠžœŽȱ‘ŽȱŽ—žœȱ’œȱŠ¡˜—˜–’ŒŠ••¢ȱŒ˜—žœ’—ǰȱ’Ž—’ęŒŠ’˜—ȱ’œȱ•’–’Žȱ

˜ȱ‘ŽȱŽ—Ž›’Œȱ•ŽŸŽ•ǯȱȱ

Muraenichthys thompsoniȱ˜›Š—ȱŠ—ȱ’Œ‘Š›œ˜—ǰȱŗşŖŞȱǻ’ž›ŽȱŜǼ

Figure 5. Muraenesox bagio (dorso-lateral view).

Figure 6. Muraenichthys cf thompsoni.

Specimens:ȱ ȱŗŖŖśŜ

Remarks:ȱȱŽ ȱœŠ–™•Žœȱ Ž›Žȱ˜‹Š’—Žȱ›˜–ȱ‘ŽȱŽœžŠ›’—ŽȱŠ›ŽŠȱ ’‘ȱ

‘ŽȱžœŽȱ˜ȱŠȱ•˜ŒŠ•ȱŽŠ›ȱŒŠ••ŽȱgarabǯȱŒ˜œ”Ž›ȱǻŗşŝŖǼȱŒ˜—œ’Ž›ŽȱM.

malabonensisȱŠœȱŠȱœ¢—˜—¢–ȱ˜ȱM. thompsoni.

Benthenchelys cartieriȱ˜ •Ž›ǰȱŗşřŚ

Remarks:ȱ‘Žȱ˜••˜ ’—ȱŠŠȱ Ž›ŽȱŠ”Ž—ȱ›˜–ȱ–’‘ȱǻŗşşŚǼǯȱ˜•˜¢™ŽDZȱ ȱ şŘřśŜȱ ǻŗŗśȱ ––ǰȱ ŸŽ›Ž‹›ŠŽȱ ŚŜȦśŖȦŗŜŝǼǯȱ ŠŒ’ęŒǰȱ ‘’•’™™’—Žȱ œǯDzȱ

‹Ž ŽŽ—ȱŠ—Š¢ȱŠ—ȱŽ›˜œǰȱžœŠ›Š–ȱǰȱȱŘřǚȱǰȱŘśǯśŖȱ–’ȱǻŚŗȱ”–ǼDzȱ ŗŖǚŖśȇŚśȈǰȱŗŘŘǚŗŞȇřŖȈDzȱŜřŞȱ–ȱǻŗŗŜŞȱ–ǼDzȱřŖȱŠ›ȱŗşŖŞDzȱAlbatrossȱśŗŞśDzȱ ŗŝŘŜȱ‘›ǯȱŠ›Š¢™ŽœDZȱȱşřřŚşȱǻŚǰȱşŝȬŗŖśȱ––Ǽǯȱ

Scolecenchelysȱ’•‹¢ǰȱŗŞŜŝȱǻ’ž›ŽȱŝǼ Specimen:ȱŗȱœ™ŽŒ’–Ž—ȱœŽ—ȱ˜ȱ‘Žȱȱ

Remarks:ȱ’œ’—ž’œ‘Žȱ›˜–ȱ‘ŽȱŽ—žœȱMuraenichthysȱ‹¢ȱ‘Žȱ™›ŽœŽ—ŒŽȱ

˜ȱŽŽ‘ȱ˜—ȱŸ˜–Ž›ǰȱ–Š¡’••ŠȱŠ—ȱŽ—Š›¢ȱǻŒ˜œ”Ž›ǰȱŗşŝŖǼǯȱŽŒŠžœŽȱ

˜—•¢ȱ Šȱ œ’—•Žȱ œ™ŽŒ’–Ž—ȱ ’œȱ ŠŸŠ’•Š‹•Žȱ Š—ȱ ‘Žȱ —ž–‹Ž›ȱ ˜ȱ ŸŽ›Ž‹›ŠŽȱ

ŒŠ——˜ȱ‹ŽȱŽŽ›–’—ŽȱŠȱ‘’œȱ’–Žǰȱ Žȱ•’–’ȱ˜ž›ȱ’Ž—’ęŒŠ’˜—ȱž™ȱ˜ȱ

‘ŽȱŽ—žœȱ•ŽŸŽ•ǯȱ

Figure 7. The worm eel Scolecenchelys from Bago River.

”¢‘›Ž—Œ‘Ž•¢œȱ£Š‹›ŠȱŠœ•ŽȱŠ—ȱŒ˜œ”Ž›ǰȱŗşşş

Specimens:ȱȱŗŚŞśŝŚDzȱǯǯȱŽ••Ž›œǰȱŘŖȱŠ›Œ‘ȱŗşŘşȱ

Remarks:ȱ ŽœŒ›’‹Žȱ Šœȱ Šȱ —Ž ȱ Ž—žœȱ Š—ȱ œ™ŽŒ’Žœȱ ‹¢ȱ Šœ•Žȱ Š—ȱ

Œ˜œ”Ž›ȱ ǻŗşşşǼǰȱ ’—Œ•ž’—ȱ Šȱ œ’—•Žȱ œ™ŽŒ’–Ž—ȱ ǻȱ ŗŚŞśŝŚǼȱ

Œ˜••ŽŒŽȱ˜ěȱ•˜’•˜ǰȱŠ—Š¢

’››‘’–ž›ŠŽ—ŠȱŒ‘’—Ž—œ’œȱŠž™ǰȱŗŞśŜȱǻ’ž›ŽȱŞǼ Specimens:ȱ ȱŗŖŖśŝ

Remarks:ȱ Ž—’ęŒŠ’˜—ȱ ™›˜Ÿ’œ’˜—Š›¢ȱ ’ŸŽ—ȱ ‘Šȱ ‘Žȱ Ž—žœȱ —ŽŽœȱ

›ŽŸ’œ’˜—ȱǻŒ˜œ”Ž›ǰȱǯȱ™Ž›œǯȱŒ˜––ǯǼǯȱ Bascanichthys sp.ȱǻ’ž›ŽȱşǼ

Specimens:ȱ ȱŗŖŖśŞDzȱȱœ’—•Žȱœ™ŽŒ’–Ž—ȱŒ˜••ŽŒŽȱ›˜–ȱ‘ŽȱœŠ—¢ȱ

ŽœžŠ›¢ȱ˜ȱ•˜ȱ’ŸŽ›ȱ™›ŽœŽ—•¢ȱŽ™˜œ’Žȱ’—ȱDzȱŒ˜••DZȱǯȱžŒ˜•ȱŠ—ȱ

™Š›¢ǯȱȱ

Figure 8. Cirrhimuraena cf chinensis.

Figure 9. Bascanicthys sp., an uncommon estuarine eel in Negros Occidental.

’››”Š•Šȱ–’œ˜•Ž—œ’œȱ ž—‘Ž›ǰȱŗŞŝŘȱǻ’ž›ŽȱŗŖǼ Specimens:ȱ ȱŗŖŖśş

Remarks:ȱ ˜•˜›Š’˜—ȱ œ’–’•Š›ȱ ˜ȱLamnostomaȱ ǻŠ˜˜”Šȱ ǭȱ ˜œ‘’—˜ǰȱ ŗşşŞDzȱ –’‘ȱ ǭȱ Œ˜œ”Ž›ǰȱ ŗşşşDzȱ ‘Š—ȱ ǭȱ œŠ’ǰȱ ŘŖŖřǼȱ ‹žȱ ŒŠ›Žž•ȱ

Ž¡Š–’—Š’˜—ȱ˜ȱ‘Žȱœ™ŽŒ’–Ž—œȱ›ŽŸŽŠ•Žȱ‘Šȱ‘Ž¢ȱ‹Ž•˜—ȱ˜ȱ’››”Š•Šȱ’—ȱ

‘ŠŸ’—ȱ‘’‘Ž›ȱŸŽ›Ž‹›Š•ȱŒ˜ž—ȱŗŜŖȬŗŜŘǯȱ’Ĵ•Žȱ’œȱ”—˜ —ȱ˜—ȱ‘Žȱ‹’˜•˜¢ȱ

˜ȱ‘ŽȱŗŘȱœ™ŽŒ’Žœȱž—Ž›ȱ‘ŽȱŽ—žœǯȱȱœ’—•Žȱœ™ŽŒ’–Ž—ȱ’—ȱȱ•Š‹Ž•Žȱ

ŠŽŒž•Šȱ –’œ˜•Ž—œ’œȱ ǻ’Ž—’ꮍȱ ‹¢ȱ ǯǯȱ ˜—£Š•ŽœǼȱ Œ˜••ŽŒŽȱ ‹¢ȱ ȃǯǯȱ Ȅȱ’—ȱ‘Žȱ—˜›‘Ž›—ȱŽ—ȱ˜ȱŠž‹˜‘Š—ȱœ•Š—ǰȱ™’Š—ȱŠœœǰȱŠ—Š¢ȱ

ŠȱŗŖȱŠ‘˜–ȱ˜—ȱžžœȱřǰȱŗşŚŞǯȱȱ Ophichthus grandoculisȱǻŠ—˜›Ǽ

Specimens:ȱ ȱŗŖŖŜŖDzȱ•˜ȱ’ŸŽ›ȱœžŠ›¢DzȱǯȱŠŒŠ•’˜ŠȱŠ—ȱ™Š›¢ǯ

Figure 10. The worm eel (Yirrkala misolensis) from Bago River.

Ž–Š›”œDZȱ‘Žȱ˜—•¢ȱ”—˜ —ȱœ™ŽŒ’–Ž—ȱ’œȱ‹Ž’—ȱŸŽ›’ꮍȱŠȱ‘Žȱǯ Pisodonophis boroȱǻŠ–’•˜—ǰȱŗŞŘŘǼ

Specimens:ȱȱœ’—•Žȱœ™ŽŒ’–Ž—ȱŒ˜••ŽŒŽȱ›˜–ȱŠ˜ȱ’ŸŽ›ȱ’—ȱŘŖŖşǯȱ

FAMILY MURAENIDAE (Morays)

Strophidon satheteȱŠ–’•˜—ǰȱŗŞŘŘȱǻ’ž›ŽȱŗŗǼ Specimens:ȱ ȱŗŖŖŜŗ

Remarks:ȱžŸŽ—’•ŽȱœŠ–™•Žœȱ Ž›Žȱ˜ž—ȱ’—ȱ‘Žȱ–ž¢ȱŽœžŠ›¢ȱ ‘’•Žȱ

‘Žȱœ’—•ŽȱŠž•ȱœ™ŽŒ’–Ž—ȱ ŠœȱŒŠ™ž›Žȱ›˜–ȱ‘Žȱ–˜ž‘ȱ˜ȱ‘ŽȱŠ˜ȱ

’ŸŽ›ǯȱ Ž››Žȱ ǻŗşŘřǼȱ ›Ž™˜›Žȱ ‘’œȱ œ™ŽŒ’Žœȱ Šœȱ Evenchelys macrurus ǻ•ŽŽ”Ž›Ǽȱ›˜–ȱ•˜’•˜ǯȱ

ŒŒ˜›’—ȱ˜ȱŠȱ‹›’Žȱ‘’œ˜›’ŒŠ•ȱ‹ŠŒ”›˜ž—ȱ˜ȱ‘Žȱœ™ŽŒ’Žœȱ™›˜Ÿ’Žȱ

˜‘•”ŽȱǻŗşşśǼǰȱ•ŽŽ”Ž›ȱǻŗŞŜřǼȱŽ››˜—Ž˜žœ•¢ȱŒ˜—œ’Ž›Žȱ‘Šȱsatheteȱ Šœȱ

œ’–’•Š›ȱ ˜ȱbrummeriǰȱ ŽŸŽ—ȱ ‘˜ž‘ȱ Š–’•˜—ȱ Š—ȱ Œ•Ž••Š—ȱ ŠŸŽȱ

’Š—˜œ’ŒȱŒ‘Š›ŠŒŽ›œȱǻ‘Žȱ™˜œ’’˜—ȱ˜ȱ‘ŽȱŠ—žœǰȱ‘Žȱ—ž–‹Ž›ȱ˜ȱ˜›œŠ•Ȭȱ

Š—ȱŠ—Š•Ȭę—ȱ›Š¢œǼǯ

Figure 11. The moray eel Strophidon sathete from Bago River estuary.

Gymnothorax polyuranodonȱǻ•ŽŽ”Ž›ǰȱŗŞśřǼȱǻ’ž›ŽȱŗŘǼ

Specimen:ȱŗȱœ™ŽŒ’–Ž—ȱǻŽ–™˜›Š›’•¢ȱœ˜›Žȱ’—ȱŠȱ˜—œ˜•ŠŒ’˜—ȱ˜••ŽŽȬ

ŠŒ˜•˜ǼDzȱŒ˜••DZȱǯȱ’—Šž˜ȱŠ—ȱ™Š›¢ǯȱ

Remarks:ȱŽ››ŽȱǻŗşŘřǼȱ’—Œ•žŽȱ‘’œȱœ™ŽŒ’Žœȱ’—ȱ‘’œȱ•’œȱ˜ȱ›Š›ŽȱŽŽ•œȱ’—ȱ

‘Žȱ‘’•’™™’—Žœǰȱ ‘’Œ‘ȱ ŽȱŒ˜—Œž›ǯ Gymnothorax tileȱǻŠ–’•˜—ǰȱŗŞŘŘǼ

Specimens:ȱŽ—ȱŒŠž‘ȱ‹¢ȱ꜑Ž›œȱ’—ȱ›’ŸŽ›ȱ–˜ž‘œǯȱŽŒŽ—ȱœŠ–™•Žœȱ

Š›Žȱ™›ŽœŽ—•¢ȱ‹Ž’—ȱŒŠŠ•˜žŽȱŠȱ‘Žȱǯȱ Gymnothorax pictusȱǻ‘•ǰȱŗŝŞşǼ

Specimens:ȱŗŘȱœ™ŽŒ’–Ž—œȱ’—ȱȱŒ˜••ŽŒŽȱ‹¢ȱ’••˜œ˜ȱŽȱŠ•ǯȱ’—ȱŽ‹›žŠ›¢ȱ ŗşŝśȱŠȱ˜Šœȱœ•Š—ǰȱ—’—’Ȭ¢ǰȱ—’šžŽǯȱ

Remarks:ȱ˜ž—ȱ‹•˜Œ‘ŽœȱŸ’œ’‹•Žȱ’—ȱœ™ŽŒ’–Ž—œǯȱ

Figure 12. The moray eel Gymnothorax polyuranodon.

ACKNOWLEDGMENTS

Žȱ ’œ‘ȱ˜ȱŠŒ”—˜ •ŽŽȱ‘Žȱ˜––’œœ’˜—ȱ˜—ȱ’‘Ž›ȱžŒŠ’˜—ȱǻǼȱ˜›ȱž—’—ȱ

‘Žȱ›ŽœŽŠ›Œ‘ȱ™›˜“ŽŒȱȃ¢›˜•˜¢ȱŠ—ȱŒ˜•˜¢ȱ˜ȱ‘ŽȱŠ˜ȱ’ŸŽ›ǯȄȱ›œǯȱ—Ž•ȱǯȱ•ŒŠ•Šȱ ǻ’›ŽŒ˜›ȱ˜ȱȬȱȱŠ—ȱǼȱŠ—ȱ›Ž—Œ’˜ȱǯȱŠŒ‘’ŒŠȱŠ›Žȱ‘Š—”Žȱ

˜›ȱ‘ŽȱŽ—Œ˜ž›ŠŽ–Ž—ȱŠ—ȱž’Š—ŒŽȱ‘›˜ž‘˜žȱ‘ŽȱŒ˜—žŒȱ˜ȱ‘Žȱœž¢ǯȱ‘Žȱ‘Ž•™ȱ

˜—ȱ‘ŽȱŠ¡˜—˜–’Œȱ’Ž—’ęŒŠ’˜—ȱ™›˜Ÿ’Žȱ‹¢ȱ›ǯȱŠŸ’ȱ ǯȱ–’‘ȱǻǼȱŠ—ȱ›ǯȱǯȱ ǯȱŒ˜œ”Ž›ȱǻŠ•’˜›—’ŠȱŒŠŽ–¢ȱ˜ȱŒ’Ž—ŒŽœǼȱŠ›ŽȱŠ•œ˜ȱŠ™™›ŽŒ’ŠŽǯȱǯ ǯȱ–’‘ȱŠ•œ˜ȱ

™›˜Ÿ’ŽȱŠ’’˜—Š•ȱ›Ž•ŽŸŠ—ȱ›ŽŽ›Ž—ŒŽȱ–ŠŽ›’Š•œȱžœŽȱ’—ȱ‘’œȱœž¢ǯȱ›ǯȱǯȱ’ŸŽ›Šȱ˜ȱ

‘Žȱ‘’•’™™’—ŽȱŠ’˜—Š•ȱžœŽž–ȱ’œȱŠ•œ˜ȱ‘Š—”Žȱ˜›ȱŠ••˜ ’—ȱ‘ŽȱœŽ—’˜›ȱŠž‘˜›ȱ˜ȱ

Ž¡Š–’—ŽȱŠ’’˜—Š•ȱœŠ–™•Žœǯ

REFERENCES

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