KHOA HOC C O N G N G H
THAIXIH PHAIXI LOAI DC SOIXIG KlIXIH CO VO O MOT SO VLIIXIG Bl£l\l VIET MAIVI
Nguyen Tai Tu^ Trdn Van B^\ Sfrenko B.P T O M T A T
Nghidn cuu ve thanh phan loai 6c Song kinh co vo d vimg bien Phii Quoc (Kien Giang), Con Dao (Ba Ria - Vung Tau), Son Duong (Ha Tinh), Vinh Hy O^inh Thuan), Nha Trang (Khanh Hoa), Son Tra ff)a Ndng), Cat Ba (Hai Phong) duoc tidn hanh voi 2 dot thu mdu (dot 1 tir thang 4 - 6/2017 va dot 2 tu thang 4 - 6/2018). tai 40 vi tri. Kdt qua cho thay tong cong co 68 loai dpng vat than mdm lop Song kinh co vo dupe phat hien thuoc 11 ho trong do, sd lirtmg loai cao nhat thuoc ve viing bien Nha Trang vdi 35 loai. Chi sd tuong ddng loai dao dong tir 0 - 0,65, cao nhat thuoc vd khu vuc Vinh Hy (Ninh Thuan) va Nha Trang (Khanh Hoa). Ket qua nghien cdu cho thdy trong 3 dang ndn day, ndn day san ho chiem iru the phan bd cho lap 6c Song kinh cd v6. Ngoai ra, kdt qua cua nghidn cmi con la ngu6n dulieu de xay dung danh muc dpng vat than mdm lap Song kinh co vo cua Viet Nam.
Tii khoa; Oc Song kinh co vo, thanh phan loai, viing bien, Viet Nam.
I.eATVAMOE
Lop de Song kmh ed vo thupe nganh dpng vat than mem co kich thudc da dang [3], vdi khoang 940 loai dang sdng [9,17] va 430 lodi hda thach [8] dupe ghi nhan. Cde loai trong lop ndy cd lop vo dupe tao nen tir tam manh vd [3]. Cac manh vo nay dupe xep chdng I6n nhau. Do do, lop vo eua chiing vira cd tac dung bao ve vira cho phep chiing udn ngudi khi can di chuydn d bd mat khdng bdng phdng; chung tham chi ed the eupn trdn lai [4]. Cae loai Song kinh co vd phan bd tren todn thd gidi, ttr vung nude lanh ddn mien nhiet ddi. Chiing sdng tren edc bd mat cung nhir tren da, canh san hd ehdt, vo dpng vat than mdm ehdt.. Thiie an chu yen cua ehung la rong, cd hien, thit [1].
Cac edng trinh nghien cuu vd 6e Song kinh cd vd cdn rat ban chd d Viet Nam, phdn Idn la ciia edc tdc gid nude ngoai nhu nhiing nghien cuu eua Leloup va cs (nam 1937, 1940,1952), da lap dupe dardi luc cae lodi thudc Idp Chitons ddu tien tai Viet Nam, hao gdm 13 loai va md ta ha lodi mod [5, 6, 7). Nam 2003, Shack bien soan dupe danh luc gdm 40 lodi thupe Idp Chitons tai Viet Nam [18]. Tuy nhien, nhidu lodi trong danh luc cdn phdi xdc minh them. Cae edng trinh nghien cuu eo tinh he thdng va lau dai eiia Sfrenko va cs (nam 1988, 1992, 1998, 2012, 2014, 2016, 2017) [11,12,13,14,15,16] da phat hien ra dupe 10 lodi mdi cho khoa hpc vd hon 30 lodi mdi cho Viet Nam.
Trung tam Nhiet ddi Viet - Nga
Tai Viet Nam, eac chuong trinh nghien cuu bidn, du an, dd tdi chu ydu tap tnmg nghien cuu cac ddi tupng ed gia tri kinh te cao hay phd hidn nhu cd, dpng vat than mdm lop hai manh vd, lop chan hung, sinh vat phii du, gidp xdc,... Do dd, nhung ngbien cuu ve Idp Song kmh cd vd con han che vd nhidu khia canh cdn dupe khai thdc nhu da dang sinh hpc, ung dung trong nghien cuu bien ddi khi hau, moi trudng, ngudn thuc pham, dupe pham,...Nghien cihi thdnh phdn lodi 6c Song kinh cd vd gop phan xay dung eo sd dir heu vd nganh dpng vat than mdm noi chung va Idp oc Song kinh ndi rieng, tu dd lam co so cho nhimg nghien ciiu tidp theo.
2. VJiT UBJ VA PHUDNG PHAP NGHBV CUU Nghien cuu dupe tien hanh theo 2 dpt, dot 1 tu thang 4 - 6/2017 va dot 2 tii thdng 4 - 6/2018 tai vimg bien Phu Quoc (Kien Giang), Cdn Dao (BdRia - Vung Tau), Son Duong (Hd Tmh), Vinh Hy (Ninh Thuan), Nha Trang (Khdnh Hda), Son Trd ffJa Ndng), Cdt Bd (Hai Phdng), tdng cong cd 40 diem thu mdu. Mdu dpng vat than mdm dupe thu bdng phuong phdp lan sau thu cac gia the (vo hau, san ho ehdt, dd) tren cac ran san hd theo mat cdt Mat cdt, dia didm thu mdu dupe lua ehpn dam bao tinh ngdu nhien, tmh dai dien cho tiing kidu dia hmh vd chat day cua tiing khu \u.c. Ddi vdi cac viing trieu ed da hinh I6i lorn va nhidu dang chat day khde nhau, dat nhidu mat cdt va thu mdu d nhieu tram tiiy thupe vdo dien tich thuc te cua tiing vimg tiidu. Ddi vdi vung tiieu CO dia hmh phdng va chat day don dieu thi s6
98 N O N G NGHIEP VA PHAT TRIEN N O N G THON - KY 1 - THANG 1/2019
KHOA HOC CONG NGHE
lupng mat cat va h"am thu mdu se it hon, vi bidn dpng da dang sinh hpc vd ngudn lpi dpng vat than mdm
^VFM) hai manh vd d dang sinh canh nay hau nhu khdng Idn, sir phdn bd tuong ddi ddu. Gid the sau khi thu dupe xu ly bang nirdc bien hoac nude ngpt, ngam gia the hang Formahn 0,4% trong 0,5 - 1 h. Sau dd, tidn hdnh lua chpn mdu qua mdt hp sang vdi kich thuoc mat ludi khde nhau (1 mm, 2-3 nun vd 5-6 mm). Cudi cung, mdu c6 dmh trong dung dich Formalin 8 - 10% vd dupe van chuydn ve phong thi nghiem, tidn hdnh phan tach thanh edc nhdm shih vat ^anh ma sd vd bdo quan mdu bang cdn 80 dd.
Dinh danh ten cde giong loai eo ti-ong mdu thu bang cdch dua vdo cac tdi h&u phan loai eua Sirenko, 2012.
Phuong phap phan loai chu yeu hdng hinh thai so sanh, dua tren dae diem phan loai thdng qua edc chi tieu hmh thdi eau tao ngodi va cdu tao hen trong ciia eo the. Danh gia su tuong ddng ve thdnh phdn loai dua vdo ehi so sau:
Chi sd tuong dong loai Sorensen (Krebc, 1999).
Trong dd: A la s6 lodi frong khu vuc nghifen cuu A, B la sd loai frong khu vuc nghien cuu B vd C Id sd lodi ehung giiia 2 khu vuc so sanh.
Xu ly sd Ueu dupe thuc hien bang chucxng hinh Microsoft excel 2013.
3. K ^ QUA NGHEN Ciiu
3.1. Thanh phan lodi dOng v$t than mdm lap Song kmh cd vd
Kdt qua nghien cuu qua edc dpt dieu fra tai vung hidn Phii Qudc (Kien Giang), Con Dao (Bd Ria - Viing Tdu), Son Duong (Hd Tmh), Vinh Hy (Ninh Thuan), Nha Trang (Khanh Hda), Son Trd (Da Ndng), Cdt Bd (Hai Phdng) da xdc dinh dupe tiidnh phan lodi dc Song kinh tai day gom 68 loai thudc 22 giong, 11 ho. Danh luc thanh phdn lodi duoc thd hifen dhang 1.
Bang 1. Thdnh phdn loai dOng vdt than mdm Ido Sons kmh cd v6 tai mdt s6 viine bidn Viet Nam TT
I 1 2 3 11 4 III 5 IV 6 7 8 9 10 V 11 12 13 14 15
Ten lodi Hp Leptoehitonidae
Leptochiton vietnamensis Sfrenko, 1998 Leptochiton muelierSir&nko etSchwabe, 2011 Parachiton poh'tnsS<dto, 1996
Hp NiersfraszelUdae
Nierstraszella lineata Niersfrasz, 1905 Ho Ahyssoehitonidae
Ferredaella takdWu et Okutani, 1984 Ho Callochitonidae
Callochiton longispinosus Leloup, 1952 Callochiton multidentatus Pilsbry, 1892 Callochiton subsulcatus Kaas et Van Belle, 1985 Callochiton dawydo^Sir^vko, 2012 Callochiton sonchaensis Sfrenko sp. n.
Hp Ischonoehitonidae Ischnochiton albinusThiele, 1911 Ischnochiton boninensis Bergenhayen, 1933 Ischnochiton /"ourRDupuis, 1917 Ischnochiton comptus Gould, 1859 Lepidozona bisculptaVa^nj, 1892
Dia diem 1 2
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3
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+ 5
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6
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KHOA HOC CONG NGHE 16
17 18 19 20 VI 21 VII 22 VIII
23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 DC 45 X 46 47 48 49 50 51 52
Lepidozona chnstiaensiNan Belle, 1982 Lepidozona coreanica Reeve, 1847 Lepidozona vietnamensis Sfrack, 1991 Lepidozona acostata Sirenko, 2016 Stenoplax alata^o^evhy, 1841 Hp CaUistoplacidae
Caldstochiton graniterHuM, 1923 Ho Schizoehitonidae
Schizochiton incisus Sowerby, 1841 Ho Chitonidae
TegulaplaxhululensisEA. Smith, 1903 Rhyssoplax bullock Sirenko, 2012 Rhyssoplax burmana Pilsbry, 1893 Rhyssoplax komaiana Is. et Iw. Taki, 1929 Rhyssoplaxci. maldivensisE.N Smith, 1903 Rhyssoplax pulcheninia Sowerby, 1842 Rhyssoplax speciosa Niersfrasz, 1905 Rhyssoplax venusta Hull, 1923
Acanthopleura loochoana Broderip & Sowerby, 1829 Acanthopleura spinosa Bruguiere, 1792 Acanthopleura tenuispinosa Leloup, 1939 LiolophurajaponicaLischke, 1873 Lucilina camosa Kaas, 1979 Lucilinaci. dilectaTnvAe, 1911 Lucilinact. tilbrookiWiXnc,l%^8 Lucilina lamellosa Quoy&Gaimand, 1835 lAicilina sowerbyi\\\ex'^\xa^z, 1905 LuciUnas^.l
Lucilina sp.^
Lucilina tenuicostata Sirenko, 2016 Onithochiton siraci/'Sfrenko, 2012
Onithochiton helenaevietnamensis Sirenko, 2016 Ho Mopaliidae
Plaxiphora bucklandnlcksi S\rcr\k.o, 2012 Ho Acanthochitonidae
Acanthochiton aachates Gould, 1859 Acanthochiton abednalh Pilshry, 1894 Acanthochiton abifonnis Niersfrasz, 1905 Acanthochiton abritayevi Sirenko, 2012 Acanthochiton asaltoi Sirenko, 2012 Acanthochiton asavmkini Sirenko, 2012 Acanthochiton aostieaphila Sirenko et Saito, 2017
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-1- + +
-H
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00 NONG NGHIEP VA PHAT TRIEN NONG THON - KY 1 - THANG 1/2019
KHOA HOC CdNG N G H £
53 54 55 56 57 58 59 60 61 62 63 64 65 XI 66 67 68
AcanthochitonalanaeSirenko etSaito, 2017 Acanthochiton anigra Sfrenko et Saito, 2017 Acanthochiton acondaoi Svrenko et Saito, 2017 Notoplax conica Is. et Iw. Taki, 1929 Notoplaxci. richardiKaas, 1990 Notoplax rosea Leloup, 1940 Notoplax nhatrangi Sirenko et Saito 2017 Notoplax odysseyi Sirenko et Saito, 2017 Leptoplax coarctata Sov/erhy, 1841 Leptoplax doederleim'Thieie, 1909 Leptoplax spi
Leptoplax tongkingi Sirenko et Saito, 2017 Craspedochiton laqueatus Sowerby, 1842 Hp Cryptoplaeidae
Cryptoplaxd. burrowi E A Smith, 1884 Cryptoplax dawydo:^ljsk>up, 1937 Cryptoplax oculataQuoy et Gaimard, 1835 Tong
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6 +
19
-1-
-H + 20
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27
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21 + +
-1-
19
* Chii diich: 1 - Phii Qudc (Kien Giang); 2 - Cdn Dao (Ba Ria - Vung Tau); 3 - Son Duong (Ha Unh); 4 - Vmh Hy (Nuih Thuan);5-Nha Trang (KhanhHda);6-San Tra (BaNdng); 7- CatBa (HaiPhdng).
dupe cdng bo tai Vidt Nam cho thdy, day la dir Ueu mdi cho cac viing hien duoe dieu fra.
Trong cde dia diem khao sdt, vimg bien Nha frang cd s6 lupng thdnh phdn lodi phong phu nhdt (35 loai), tidp theo la Vinh Hy (27 lodi). Son Tra (21 lodi). Son Duong (20 loai), Cdn Dao (19 lodi), Cat Bd (19 lodi), thap nhat tiiupc ve vimg bidn Phu Qudc (6 loai). Didu ndy chung td, su khde biet ve vi ti-i dia ly, khi hau giira hai mien Bae - Nam anh hudng khdng rd ret ddn sd lupng lodi ctta Idp Song kmh ed vd.
3.2. Chi sd tuong ddng lodi Bang 2. Chi sd tuong ddng lodi giua cac vung nghidn ctru Da didu fra dupe 68 lodi ddng vat than mdm lop
Song kmh cd vo tai cac vimg bien: Phu Qudc, Con Dao, Son Duong, Vinh Hy, Nha Trang, Son Trd, Cdt Bd. Sd lupng thdnh phdn lodi frong danh luc da thdnh lap d fren nhidu hon so lupng thdnh phan lodi ma Sfrenko vd cdng su da dieu fra duoc tai cde viing bidn Viet Nam tii nam 2012 den nam 2016 (61 loai). Cd 7 lodi trong danh luc ndy chua dupe edng hd frong tdi Ueu cua Sirenko Id cac lodi cd sd thu tu frong bang (6), (22), (25), (29), (32), (47), (63). Ddi chidu vdi cde tdi heu da cong bd ve tbdnh phdn lodi de Song kinh
Vimg nghifen ciiu
Phii Qudc Con Dao Son Duong
VihhHy Nha Trang
Son Tra CatBa
Phii Quoc 1 0,32 0,15 0,18 0,15 0,07 0
Con Dao 0,32 1 0,46 0,43 0,52 0,15 0,26
Son Duong
0,15 0,46 1 0,47 0,47 0,24 0,26
Vinh Hy 0,18 0,43 0,47 1 0,65 0,29 0,17
Nha Trang 0,15 0,52 0,47 0,65 1 0,39 0,30
Son Tra 0,07 0,15 0,24 0,29 0,39 1 0,35
CatBa 0 0,26 0,26 0,17 0,30 0,35 1 Chi so tuong dong loai cao nhat thuoc ve khu
vuc Vinh Hy (Ninh Thuan) va Nha Trang (Khanh Hoa) la 0,65. Do day la hai khu vuc co vi tri dia ly va di4u kien khi hau tuong dong. Miic do tuong dong
day la hai khu vuc nghien cuu each xa nhau nhat trong cac khu vuc nghien cuu, hai khu vuc nay chiu anh huong cua thoi tiet khi hau va vi tri dia ly khac
101
KHOA HOC CdNG NGH biet. Vimg hien Cat Bd thudc vinh Bdc bp, mang khi
hau dae trung ciia mien Bdc Viet Nam vd vimg bidn Phu Quoc tbude vinh Thai Lan, mang khi hau midn Nam Viet Nam. Cd hai khu vuc nay ed so lupng lodi didu fra dupe thap nhdt frong cde vimg nghien cuu (Cat Bd 19 loai, Phu Qudc 6 loai). Ket qua nay Id co sd dd nghien cuu anh hudng ciia didu ki6n mdi
Bang 3. Phan hd edc lodi BVTM lop Song kinh cd vd frdn cac dang ndn day
trudng den su phan bo cua dpng vat than mem lop Song kinh cd vd.
3.3. Phan \)6 thanh phdn lodi tiieo dang ndn day Nghidn cuu phan bd thanh phan loai theo dang ndn day tai cac khu vuc nghien cuu thu dupe ket qua dbang 3.
TT 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32
Ten loai Leptochiton vietnamensis Sfrenko, 1998 LeptochitonmuellerSirenko etSchwabe, 2011 Parachiton pohtiis Saito, 1996
Nierstraszella lineata Niersfrasz, 1905 Ferretaella takdWu et Okutani, 1984 Callochiton longispinosusLehup, 1952 Callochiton multidentatus Pilsbry, 1892 Callochiton subsulcatus Kaas et Van Belle, 1985 Callochiton da wydo^'Sirenko, 2012 CaUochiton sonchaensis Sfrenkosp. n.
Ischnochiton albinusT\Ae\e, 1911 Ischnochiton boninensis Bergenhayen, 1933 Ischnochiton Aowrpz Dupuis, 1917 Ischnochiton comptus Gould, 1859 Lepidozona bisculpta Pilsbry, 1892 lepidozona christiaensi^an BeUe, 1982 lepidozona coreanica Reeve, 1847 Lepidozona vietnamensis Sfrack, 1991 lepidozona acostata Sfrenko, 2016 Stenoplax alataSovierhy, 1841 Caldstochiton granderHuU, 1923 Schizochiton incisus Sowerby, 1841 TegulaplaxhululensisEA. Smith, 1903 Rhyssoplax bullock Sirenko, 2012 Rhyssoplax burmanaFHshry, 1893 Rhyssoplax komaiana Is. et Iw. Taki, 1929 Rhyssoplaxci. maldivensisEA. Srmth, 1903 Rhyssoplax pulcherrima Sowerby, 1842 Rhyssoplax speciosa Niersfrasz, 1905 Rhyssoplax venusta HuU, 1923
Acanthopleura loochoana Broderip & Sowerby, 1829 Acanthopleura sp/oosa Bruguiere, 1792
Cae dang ndn ddv Sanhd
chet + -t-
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-f
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Vo ddng vat than mdm chet
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-1-
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N'dn da
-h
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102 NONG NGHIEP VA PHAT TRIEN NONG THON - KY 1 - THANG 1/2019
KHOA HQC CONG NGHl
33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68
Acanthopleura tenuispinosa Leloup, 1939 Liolophurajaponica Lischke, 1873 Lucilina camosa Kaas, 1979 Lucilinact dilectaThiele, 1911 Lucilinaci. tilbrookiMi\ne,\^b8 Lucilina lamellosa Quoy&Gaimand, 1835 Lucilina sowerbyi^ierstrasz, 1905 Lucilina sp.l
Lucilina sp.2
Lucilina tenuicostata Sirenko, 2016 Onithochiton stiackiSirenko, 2012
Onithochiton helenae vietnamensis Sirenko, 2016 Plaxiphora bucklandnicksi Sirenko, 2012 Acanthochiton aachates Gould, 1859 Acanthochiton abednaldVUshry, 1894 Acanthochiton abdomiis Niersfrasz, 1905 Acanthochiton abritayevi Sirenko, 2012 Acanthochiton asaitoiSirenko, 2012 Acanthochiton asavmkini Sirenko, 2012 Acanthochiton aostreaphila Sfrenko et Saito, 2017 Acanthochiton a/anae Sfrenko et Saito, 2017 Acanthochiton anigra Sirenko et Saito, 2017 Acanthochiton acondaoi Sirenko et Saito, 2017 Notoplax conica Is. et Iw. Taki, 1929 Notoplaxci. richardiKaas, 1990 Notoplax rosea heloup, 1940 Notoplax nhatrangi Sirenko et Saito, 2017 Notoplax odysseyi Sirenko et Saito, 2017 Leptoplax coarctata Sowerby, 1841 Leptoplax doederleiniThiele, 1909 Leptoplax spi
Leptoplax tongkingi Sirenko et Saito, 2017 Craspedochiton laqueatus Sowerby, 1842 Cryptoplaxd. buirowiEA. Smith, 1884 Cryptoplax dawydo^Leloup, 1937 Cryptoplax oculata Quoy et Gaimard, 1835 Tong
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20
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26 Kdt qua d bang 3 cho thay, frong tdng sd 68 lodi
thi CO 45 loai phan bd fren ndn day san hd ehdt, chidm 49,5%, so loai phdn bd fren ndn ddy da Id 26 lodi, chidm 27,5%, so lodi phan bd fren vo dpng vat than mem la 20 loai, chidm 23%. Trong do co 4 loai
phan bd fren ca 3 dang ndn day, cd 11 loai phan bd fr6n hai dang nen day san hd va vo dpng vat than mem, ed 6 loai phan bd fren hai dang ndn day san ho va ndn day da, ed 2 loai phan bd fren hai dang ndn day vo dpng vat than mdm va day da.
103
KHOA HOC CONG NC Nhu vay, ndn day san hd chiem uu thd phan hd cho lop 6c Song kinh ed vd. Tuy nhien, do dae didm dia hinh, cau tnic ran d eac khu vuc khde nhau cd nhiing dae trung rieng se tac ddng len su phan bd cua chiing. Cae dia diem nghien cuu chu ydu fren cde ran san ho chet, nen cdn cd cae nghien cuu tidp theo va sau hon dd khdng dinh muc dd uu the eua ndn day san ho doi vdi su phan bd cua Idp de Song kinh cd vd.
4. KET LUAN
Da didu fra dupe 68 loai ddng vat than mem Idp Song kmh cd vo tai cde vimg bidn Phu Qudc (Kien Giang), Con Dao (Bd Ria - Vimg Tau), Son Duong (Ha Tmh), Vinh Hy (Ninh Thuan), Nha Trang (Khanh Hoa), Son Trd (J)a Ndng), Cat Ba (Hdi Phdng).
Sd lupng lodi da didu fra dupe cao nhat thupe vd vung bien Nha Trang vdi 35 lodi.
Clu sd tuong ddng lodi cao nhat thuoc vd khu vuc Vinh Hy (Nmh Thuan) vd Nha Trang (Khdnh Hda) la 0,65.
Nen day san hd chidm uu thd phan bd cho Idp dc Song kmh ed vo.
TAI L P THAM KHAO
1. Bamawell, E. B. (1960). The carnivorous habit among the Polyplacophora.
2. Britayev T. A , Pavlov D. S. (Eds). Bentiiic fauna of the Bay of Nhatrang, South VietnamN.2, Moscow, KMK.346 p.
3. Chisholm, Hugh, ed. (1911).
"Chiton". Encyclopdia Britannica. 6 (11th ed.).
Cambridge University Press, pp. 247-251.
4. Connors, M. J., Ehrhch, H., Hog, M., Godeffroy, C , Araya, S., Kallai, I., Gazit, D., Boyce, M., Ortiz, C. Three-Dimensional Structure of the Shell Plate Assembly of the Chiton ToniceUa Marmorea and Its Biomeeharueal Consequences. Journal of Structural Biology vn (2):
314-328.
5. Leloup E., 1937. Diagnoses de sbc nouvelles espeees d' Amphineures polyplacophores de la region IndoPaeifque. Bulletin du Musee royal d'Histofre nattu-elle de Belgique, 13(38): 1-3.
6- Leloup E., 1940. Les Chitons du geme Cryptoplax Blamville, 1818. Bulletin du Musee royal d'Histofre naturelle de Belgique, 16(33): 1-32.
7. Leloup K. 1952 Polyplacophores de I'Ocean Indien et des cotes de I'Indochme francaise. Institiit Royale des Sciences Naturelles de Belgique, Memotes, (2)7:1-69.
8. Puchalski, S., Eemisse, D. J., Johnson, C. C.
(2008). The effect of sampUng bias on the fossil record of chitons (Mollusea, Polyplacophora). ./Ime/Tcan Malacological Bulletin. 25: 87-95.
9. Schwabe, E (2005). A catalogue of recent and fossil chitons (Mollusea; polyplacophora) addenda. Novapex. 6:89-105.
10. Serb, J. M., Eemisse, D. J. (2008). Charting Evolution's Trajectory: Using Molluscan Eye Diversity to Understand Parallel and Convergent Evolution". Evolution Education and Outieach 1 (4):
439-447.
11. Sirenko B. L, 2012. Chitons (Mollusea, Polyplacophora) of Nhatrang Bay, South Vietnam.
Benthic fauna of the Bay of Nhatrang, South Vietiiam. Britayev T. A., Pavlov D. S. (Eds). V.2, Moscow, KMK. P. 56-122.
12. Sfrenko B. I., 2014. New tinds of minute chiton of genus Leptochiton (Mollusea, Polyplacophora) in Vietnamese waters. Ruthenica 2014,24(2).P. 65-73.
13. Sirenko B. I., 2016a. Chapter 10. A supplement to the Recent Chitonidae (Mollusea- Polyplacophora) of Vietnam. Biodiversity of the western pari of the China Sea. A.V. Adrianovet K. A Lutaenko (eds). Vladivostok. Dahiauka. P. 481-502.
14. Sirenko B. I., 2016b. Seven-year taxonomica!
investigation of chitons in Vietnam. Seven-year taxonomieal investigation of chitons in Vietnam.
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POLYPIACOPHORA MOLLUSCS SPECIES IN SOME VIETNAMESE MARINE AREAS Nguyen Tai Tu, Tran Van Dat, Sirenko B.I Summary
The study of Polyplacophora species composition in Phu Quoc (Kien Giang provmce), Con Dao (Ba Ria - Vung Tau province). Son Duong (Ha Tinh province), Vinh Hy (Ninh Thuan province), Nha Trang (Khanh Hoa province). Son Tra (Da Nang city), Cat Ba (Hai Phong city) marine areas was conducted with two sampling periods (period 1:4- 6/2017 and penod 2 : 4 - 6/2018), at 40 sites. The results showed that total of 68 species belonging to 11 families were recorded among them, Nha Trang marine area had the most diverse species composition with 35 species. The similarity index of Polyplacophora species composition between studied areas was around 0 - 0.65, Vinh Hy (Nmh Thuan) and Nha Trang (Khanh Hoa) manne areas had the most sirailanty index. The results showed that in three types of bottom substrates, coral substrates had the most distribution of Polyplacophora. In addihon, the results of this research would be used as database for development of checklist Polyplacophora of Viet Nam.
Keywords: Polyplacophora. species composition, marine areas. Viet Nam.
NguW phan biSn: TS. ViJ Vi^t H4 NgSy nhM bai: 28/9/2018 Ngay thftng qua phan bi?n: 29/10/2018 Ngiy duyM dang: 5/11/2018
