ART. 28
CHALCID
FLIESNORTH
OFMEXICO
BALDTJF71
72 PROCEEDINGS
OFTHE
NATIONAL.MUSEUM
vol.7&abdomen more
broadly, black; vestiture, sculpture,and
the head di- mensions, as described for female.Type
localit'if.—
Gaviota, Calif.ry;?^.—Female,
U.S.N.M.
No. 42246.Remarks. — The known
material ofthis species consistsof the type., the allotype,and
11 female paratypes; the firsttwo, with three para- typesand
the frontwing
of a female paratypeon
a slide, are depos- ited in thecollection of the United StatesNationalMuseum, and
the remainder are in the collection of the writer. All these specimenswere
rearedby
DoctorKinsey from
thegalls ofAndricuspomifor^nu
rossi (Kieffer)
on
Californialiveoak
{Querent agrifolia) atGaviota, Calif., thegallmakers
issuingon March
5, 1920.29.
DECATOMA
QUERCI-LANAE (Fitch)Plate2, Figueb26
Spalangiaquerci-lanaeFitch,Ann.Rep.
New
YorkState Agr. Soc., p. 816, 1859;
5th Rep. Nox. Ins.
New
York, p. 36, 1859.Decatoma querci-lanae Ashmead, Trans. Amer. Ent. Soc, vol. 14, pp. 196, 197, 1887.
DecatomahyalipennisWalsh,Amer. Ent.and Bot., vol.2, no. 10, p. 301,1870.
—
Provancheb, Additions k la faune hymenopterologique, p. 193, 1887.
Decatoma simplicistigma Walsh, Amer. Ent. and Bot., vol. 2, no. 10, p. 301, 1870.
Decatomufiavicollis Walker, Ent. Mag., vol. 2, p. 156, 1834. Original descrip- tion.
A
European species.—
Mate, Verb. Zool. Bot. Ges. Wieu, Band 55, pp. 532-534, 1905. Redescribed.
DecatomafiavicollisAshmead,Trans. Amer.Ent. Soc, vol. 12, p. xiii,1885.
An
American species.Decatoma fioridanaJ. B. Smith,Ent. Americana,vol. 2, p. 19, 1886.
New
namefor fiavicollisAshmead.
DecatomamicJiigamca Giba-ult,Proc U. S. Nat. Mus., vol. 58, p. 209, 1920.
Similar to foliatae
Ashmead
in size,form
of the head,and number
of setae
on
the submarginal vein, but differsfrom
it inhaving
even the coxae all yellowand
the thorax with amore
or less developed irregular dorso-median longitudinal black band, while foliatae hasmost
ofhead, theprescutum,scutel,propodeum, and most
ofabdomen
black. Specimens
having
scapulaemore
or less black are being regarded as foliatae.Fenidie.
—
^Length, 2.5 to3.5mm.,
mostly about3mm.
; fairly robust, colors yellow, brownish yellow,and
black, relative degrees of each variable;head and
thoraxhoney
yellow; scape almost concolorous with head, basal two-thirds of pedicel brown-black, its apexhoney
yellow, flagellumchestnut-brown;head
with only vertexand
occiput black; black of vertexnarrowly,orsometimesbarely, inclosingocelli, or sometimes extended to include disk of vertex but very rarely reaching eyes laterad or farther than apex of antennal scrobe orART.28
CHALCID
FLIESNORTH
OFMEXICO BALDUF 73
hind edge of head; occiput usually mostly black, this notuncom- monly
broadeningtoward
but seldom reaching the vertex exceptby
a narrow, or rarely a broad, longitudinalmedian
band, theband
sometimes extendingcaudad
varying in widthand
length, occasion- ally reaching hind edge ofpronotum; mesonotum
rarely almost immaculate, usually with amedian
longitudinal blackband
varying in widthat different pointsand
in a seriesof specimens, often broad- enedespeciallyon prescutum, sometimes enlargedtoa trianglealmost coextensive with borders ofprescutumand
extending forward under pronotum,band more
regularon
scutel,almost always full length of thesclerite,and
usuallynotbroader thanhalfthe width ofthis area, scapulaeand
axillae immaculate,rarely in partinfuscated, sterna of thoraxblackmostlybetween coxae only; anterior portionand
groove ofpropodeum
black, often feebly so; legs yellow, coxae concolorous with sides of thorax, rarely the hind pair feeblyand
partly black- ened,femora
slightly lighter, tibiae light yellow, tarsi stramineous;
stigma brown, submarginal
band
lacking; peduncleblack, dorsal area ofabdomen
narrowly to broadly black with pointed projections extendingtoward
middle of sides,abdomen
otherwise ochreous- yellow.Head
three-fifths as long as broadfrom
above, face broadly rounded,eyes not abruptly separatedfrom
front, vertex feebly con- vex, hind edge ofhead
almost straight; ocellar area finely punctate,malar
space smooth, polished, rest ofhead
moderately reticulate- punctateand
sparselyand
inconspicuously hairy; about two-thirds as long as pediceland
a fifth longer than each ofF2
to F5, latter joints subequal, scarcely longer than broad, club not quite so long asF3
toF5
combined, flagellum rather densely hairy;pronotum
slightly narrower than
head and
mesothorax; pronotum, prescutum,and
scutel moderately umbilicately punctateand
hairy, mesopleura aciculateand
finely punctate; 12 to 14 setaeon
submarginal vein;groove of
propodeum
crossedby
a rounded carina in frontand by
feeble rugae
on
posterior two-thirds;abdomen
polished, sparsely hairyon
the sides, moderately hairyand
punctate in the region of the ovipositor sheath.Male.—The male
of this species is extremely variable in color.One
color variety is being recognized in this sex in addition to the lighterform
described herewith.Like the female in general, except
on
the average slightly darker, the occiptal black space usually larger, the scapulae, hind coxae,and abdomen more
often partlyand more
extensively black; pedicel a fourth longer than Fl, latter twice as long as wide,F2
toF4
equalin
width and
subequal in length, about five-sixths as long as Fl, club slightlylonger thanF3 and F4
combined.74 PROCEEDINGS
OFTHE NATIONAL MUSEUM
vol.79Type
locality.— New York
State(Asa
Fitch).ry;?e.—Female,
U.S.N.M.
No. 1832.No
other type material in theUnited
States NationalMuseum
collection.
Remarks. —
Redescribedfrom
the type, the types ofsynonymous
forms,and
a series ofspecimensfrom
various regionsin the eastern half of theUnited
States.Ashmead
pointed out thatD. hyalipennisWalsh
isconspecificwith querci-lanae (Fitch).He
likewiseshowed
that D. shnplidstiginaWalsh
is also this Fitch species, but onlyestablished the identity of Walsh's species with dorsalis (Fitch), adark
color variety of querci-lanae.There
are present, in addition to thedark
specimens, typical light-coloredforms
of sitiipliclstigma that agree fully with the Fitch typeand
checkwith the descriptions of the typical querci- lanae.A
comparison of the aboveforms and
the available rearing data leave no doubt about the correctness ofAshmead's
conclusions that siniplicistigmaand
querci-lanae areconspecific.Professor
Smith
placed flavicollisAshmead
insynonymy, renam-
ingthespecies -floridana in viewof the preoccupation of thename by
flavicollis Walker. D. floridana is
no
doubt just another lot of querci-lanae (Fitch). All the type specimens, except one, are males.The
female is agood
querci-lanae.,and
the males mostlycome
close to variety dorsalis (Fitch),and
agree in color variation with series since rearedfrom
gallsofAndricus flocci(Walsh) on
Quercus alba.,the hosts of thetypes of querci-lofnae.
Similarly, micJdganica Girault, represented in the
United
States NationalMuseum
collectionby
the typeand
a paratype, both fe- males, are true querci-lanae (Fitch), agreeingin all essentialrespects with the typical light colorform
ofthis species.No
other species of this genus has perhaps been collectedand
reared socommonly
as this one.- Following are the records athand
:
The
female type alone remains of a series rearedby
Doctor Fitch inNew York
Statefrom
galls ofAndricus
flocci(Walsh)
{Cynips quercus-lana Fitch);Walsh'sseries {simplicistignia)came from
galls of Cynips pezomachoides OstenSacken and
Biorhiza forticornis (Walsh), bothon
Q. alha^and A.
petiolicola (Bassett)on swamp
white oak {Q. bicolor), all presumably
from
northern Illinois;Ash-
mead's floridana (flavicollis) specimens were"bred from
an unde- scribedCynips
gall," Jacksonville, Fla.;and
the Girault types of mdchiganicaarefrom
a"whiteoak woolly twiggall " (Ace. Cat.737, Agr. Coll.Mich.,June
27-28and
July9, 1887) in Michigan; anotherAshmead
seriesfrom
Jacksonville, Fla., hason
several pins the label"TV. (Neurotenos)
ma
jcdis,on
Q.alh a^\'aiew
old s-pecimens are labeled"-4. flocci,''^
and
DoctorKinsey
reared a large series, bothsexes,from
a variety ofthis gall as follows: Jacksonrille, Tex., Q. stellaia;Big
ABT.28
CHALCID
FLIESNORTH
OFMEXICO — BALDUF 75
StoneGap,
Va., Q. alba-, a fewfrom
galls of A. aciculatus Beuten- mueller on Q. stellata^ Austin, Tex.;and
one, same,from Yoakum,
Tex.; the gall
makers
of theseKinsey
lots issued between October 25and December
4, 1919. Several specimens datedJune
29, 1924,from
galls of A. 'petiolicola var. on Q. Mcolor atWinona
Lake, Ind.,were
collectedby
F.Payne and
sentme
through Doctor Kin- sey; I havereared a femalefrom
the stem gall of DisholcaspisTnam-mM (Walsh)
on Q. rrmcrocarpa, Urbana, 111.,May
23, 1924,and
3 females issuedMarch
21, 1929,from
galls of D. globulus (Fitch) collectedon and
under Q. michauxii at Catlin, 111., onFebruary
24, 1929; a female (463*^) rearedby
J.W.
Lloyd, Cincinnati, Ohio,on June
24, 1895,from
gall of Neurotenus saltatorius'^.\ several fe-males (3126°^)
from
N. hatafus'^. {Cynips q-hatahis'^.), on Q. alba,June
30, 1883, Virginia,by Theodore
Pergande; 2 (2610«) issuedMay
4, 1882,from
cynipid galls received at Washington, D.C, on March
23,and
collectedfrom oak by H. K.
Morrison at Fort Grant, Ariz.; another (2744") withsame
data as precedingissued in transit toWashington, D.
C.,and was
receivedon June
12, 1882; a female (189="°)came from
Cadet, Mo.,where
itwas
presumably rearedon February
10, 1883; 3 females (158'') on multiple pointmount
arefrom
J.W.
Letterman, Allenton, Mo., A. jloecl (Walsh),Q. alba,
February
14; 2from
oak,Washington,
D.C, May
12, 1895;1BiscayBay,Fla.,2
from Texas
(Belfrage) ;2from New
Brunswick, N. J.; 1, probablyfrom
a gall, is dated October 27, 1876; another (17^)from oak
gall, St. Louis, Mo., C. V. Riley; A. T.McClay
reared a female
from
an unidentified gall at Hillview, 111., April6, 1928;L.
H. Weld
rearedand
sentme
a small seriesof each of the followingfrom
cynipid galls, thenumbers
in parentheses being Weld's recordnumbers;
Evanston, 111. (210), Callirhytis scitula (Bassett), Q. coccinea;same
locality (209), C. tumiflca (Osten Sacken), Q. borealis;same
place (21), C. seminator (Harris), on Q. alba; Moline, 111. (168), C. flampes (Gillette), Q. macrocarpa;Winnetka,
111. (35), Acraspis erinacei Beutenmueller, Q. dlha; Iron- ton,Mo.
(660),Andrlcus
paAtoni (Bassett), Q. stellata;and
East Falls Church, Va. (1124), A. murtfeldtaeAshmead,
Q. stellata.I find in the National
Museum
collection one female(Hopk. U.
S.14636") reared
from
a gall, determinedby William
Middleton as a Oallirhytissp., on Q. alba, at EastFalls Church, Va.,April 27, 1920.It has the color
and form
of the typical querci-lanm, but departs strangely in possessing a faintband
on the wing. This is distinctly narrower than the length of the marginal veinand more
than twice as long as broad.Such
instances of the occasional occurrence of aband on wings
of a species typically lacking it are rare in material accumulated to date.A
single female(Hopk.
U. S. 15639<^), rearedby
L.H. Weld.
Feb- ruary 3, 1922, at Oracle, Ariz., supposedlyfrom
a cynipid gall, on76 PROCEEDINGS
OFTHE NATIONAL MUSEUM
vol.79Q. arizonica^ has the legs in part lightly clouded, but is otherwise a typical5"wem-?an«6 (Fitch).
30. DECATOMA QUERCI-LANAE var. DORSALIS (Fitch)
Spalangia qucrci-lanae var. dorsalis Fitch, Ann. Rep.
New
York Agr. Soc, pp.81G-817, 1859; 5th Rep. Nox. Ins.
New
York, p. 36, 1859.Decatotna querci-lana-dorsalis (Fitch), Ann. Ent. Soc.Amer., vol.7, p. 8, 1914.
Decatoma querci-lanae Ashmeiad, Trans. Amer. Ent. Soc, vol. 14, pp. 196, 197, 1887.
Differs
from
thelighterform by
itssmaller sizeand
inhaving thehead
always continuously blackfrom
vertex to occiput,and
the rest of thebody
generally blacker.No
such dark forms areknown
inthe female sex.Male.
—
Length, 1.9to 2.5mm.,
mostly intermediate size, relatively slender, colors blackand
yellow, black predominating; in extreme black specimens,head
black except oral areaand narrow
circum- ocular ring, otherwise degree of yellow variable, the ocular ring sometimes dilating to include cheeksand
templesand
yellow of oral area extending toocciput behindand up
the face to antennal fossae or higher, but vertex almost always entirely blackfrom
antennal scrobe to ocular band, black area of vertex always broadly contin- uouswith thatofocciput, latter also entirely black; antennae colored ason
lightermales,and
the females;
pronotum
ofdarkest specimens withan
almost square black area full length ofmedian
third,and
rest of sclerite
lemon
yellow; butmedian
black space sometimes not solongaspronotum and
notsobroadasmedian
thirdofthissegment,and
sometimes hind edge ofpronotum
narrowly yellow or with asomewhat
rounded yellow area;on
one specimen the lateral thirds black, these separatedfrom
mesal black area onlyby
a fine oblique line ofyellow on each side; mesothorax usually mostly black, some- times entirely so with only tegulae yellow brown, mesopleura occa- sionally with yellow patches,more
oftenscapulaeand
axillae inpartbrown
yellow, rarely a yellowlineornarrow band
ineach parapsidalfurrow and
continuingcaudad
over lateral thirds of scutel; meta- thoraxand propodeum
almost always entirely black, occasionally a yellowish patchon
metapleura; legs mostly yellow, coxae almost alwaysmore
orlessblack,hind pairmostlyso, disks ofmiddlefemora and
tibiae sometimes lightly infuscated, those of hind legs darkerstill, rarely entirely black, peduncle black,
abdomen
likewise, fre- quently shadingtobrown
blackon
ventral half.Dimensions
ofhead and
funicular segment,and
sculpturingand
vestiture as in lighter
form
males.Type
locality.— New York
State.Type.—
Mule,U.S.N.M.
No. 1833 (Dr.Asa
Fitch).No
other typematerial hasbeen found.ART.28
CHALCID
FLIESNORTH
OFMEXICO BALDUF 77
Remarks.—
Keclescribedfrom
the typeand
agood
series of reared specimensfrom
various localities in the eastern half of the United Statesand
Texas,D. querci-lanae var. dorsalis (Fitch) represents a unique color
phenomenon
in the genus. Sufficient intergrading variations be- tween the lightermale and
variety dorsalis occur toshow
that they are conspecific.The
specimens that have the black of the vertexand
occiput interrupted,namely
the lighter form, are notmore
nu-merous
than variety dorsalis^and among
the latter the majority ap- proach the extreme black state rather than the lighter male form.Fitch (loc. cit.) states:
"Specimens
frequently occur, so very differ- ent in their colors that theymight
almost bedeemed
a distinctspecies."
The
sex of his specimens thus described is not given, but his decision to regard dorsalis as a variety proved sound in view of subsequentrearings. First,no
mostly black femaleshave been taken to date that can be placed in variety dorsalis;and
second, I find several instances inwhich
only the dorsalis variety of malesand
the typical light-colored females were rearedfrom
thesame
lot of galls.A
further point of interest is the occurrence of similar mostly blackmales in foliataeAshmead,
foliatae arizonica^new
variety,and
quinqueseptae,new
species, in addition to a lighter-colored form, as in querci-lanae (Fitch).On
the other hand, D. wiltzae,new
species, ismostly blackin both sexes,and
its male has the colorsand
pattern of variety dorsalis (Fitch).So
similar are the males of these spe- cies that to date nomeans
of distinguishingthem
has been discov- ered. Thissimilarity extends also tosize, dimensions, sculpture,and
vestiture, but the differences between the females, supplemented
by some
distinctions inhostrelations, constitutegood
reasonsforregard- ing these species as distinct.The
only presenthope of determining the black males of these several species is to rearthem
with the femalesfrom known
oakand
cynipid hosts.Variety dorsalishas beenobtained
from
mostof the sourcesnamed
underthe typical querci-lanae.Only
the localitiesneed to benamed
here,the dates, oaks,
and
cynipidsbeing as al'ready detailed:Winona
Lake, Ind.,Urbana and
Catlin, 111., Jacksonvilleand Yoakum,
Tex.,Michigan
Agricultural College,Texas
(Belfrage),and
Jacksonville, Fla. \N. majalis (Bassett)]. Additional records are:One from
"oak woolgall,"And7^icusflocci (Walsh),labeled "Illinois,"
emerged November
16, 1908; one (158^°')was
collectedby Mr.
Koebele, Feb- ruary 1, 1883, on Q. macrocarpa., Washington, D. C.; one taken in April, Riley County, Kans.,by
Doctor Marlatt; onefrom
gall of Callirhytis tumifica (Osten Sacken), at Braintree (near Boston, Mass., July 6, 1918, determinedby
Doctor Kinsey; one (Quaintance No. 24477) collectedby H. K.
Plank,November
17, 1924,Wagners
Ferry, La.;
two from
J. G. Barlow, Cadet, Mo., collectedMay
3,78 PROCEEDINGS
OFTHE
NATIONAL,MUSEUM
vol.79 1885, "on buds of whiteoak
";and
data are athand
for aMichigan
specimen (Agr. Coll. Mich., Ace. Cat. 592c) rearedMay
20, 1888,from
the gall ofNeurotems
noxiosus (Bassett) on "swamp
oak."31,
DECATOMA
QUINQUESEPTAE, new speciesNear
D. foliataeAshmead
in having the yellow on the prescutum variableand
the scutel almost always entirely black,and
in thenum-
ber of setae
on
the submarginal vein; D. quinquese'ptae issomewhat
largerand
less robust,and
differsmost
conspicuously in having most of the sides of theabdomen
ochreous-yellow, whereas these are en- tirelydark
in foliatae.Female.
— Length
2.4 to 2.7mm.,
colors yellowand
black; head black, onlynarrow
ocularband
with sometimes an oblique projectionfrom
iton
vertextoward
scrobe,and
oral area, labium,and
space belowantennal fossaeand
betweenmalar
groovesmore
orless,yellow- ishbrown;
scapelemon
yellow, its radicle yellowish brown, pedicel darkbrown
outwardly, only small part of apex lighter, flagellum uniformly light chestnut-brown;pronotum
mostlylemon
yellow, black of neck extendingcaudad and forming
subquadrangular patch on middle third ofpronotum and
occasionally reaching hind edge, usually leavingnarrow
border ofyellowon
hind edge ofpronotum
;
mesonotum
mostly black, parapsidal grooves rather broadly,and
posterior part of side of, ormore
oftenmore
than posterior half of scapulae, yellowish orange, tegulaelight brown,upper
edge of pleura brown, scutel, axillae,metanotum, propodeum, and
venter of thorax, black; legsmostly immaculatelemon
yellow, onlybase of coxae some- times narrowly black,and
tarsi stramineous-yellow; stigmadark
brown, faint rudiment ofsubmarginalband
present; peduncle black,upper
two-thirds of first abdominal segment black to brown, lighter beneath,upper
third of second segmentbrown
black, lower edges of dark areas irregular,dorsum
of posterior half ofabdomen
lightly infuscated,and most
orall ofitslateralsurfacesand
venter ochreous- yellow.Head from
above three-fifths as long as itsmaximum
width, face rather well-rounded transversely between eyes,more
prominent dorso-ventrally, eyesand
face not abruptly set apart at their junc- tion, vertex moderately convex, hind edge ofhead
feebly convex, hencehead
not approaching rectangulate-ovatefrom
above; lower portion of genaeand most
ofmalar
space polished, shiny,upper
part of cheeksbecoming
strigose-reticulate, ocellar area finely punc-tate, also bearing small mnbilicate punctures, rest of head
more
coarsely reticulate-punctate,and
moderately hairy, antennaemore
denselyhairy; pedicel scarcelytwice longer thanitsmaximum
thick-ness, conical, about a fourth longer than Fl,
Fl
nearly a fifth longerART.28
CHALCID
FLIESNORTH
OFMEXICO — BALDUF 79
than each ofF2
to F5, latter jointsuniform
in dimensionsand
barely longer than wide, club as thick as funicular joints, tapering to a blunt tip,and
slightly longer thanF4 and F5
combined; pro-notum and mesonotum
rather coarsely umbilicate, anterior-lateral sclerite of mesopleura densely papillose, posterior area aciculate- punctate, polished;propodeum
coarsely reticulate, or rugose, groove limited in frontby
arounded
carina,and
with a polished depres- sion in anterior portion; 12 or 13 setae on submarginal vein; abdo-men
polished, smooth, with only small parts granulose, area of ovi- positor sheath denselyand
rather coarsely punctate, hairy.Male.
— Length
2.2 to2.5mm.,
fairly slender, blacker than female;head
black,exceptoral areanarrowly brown,scape sometimesduslcy;
sides of
pronotum
yellow, the yellow area emarginate in frontand
below, rest of thorax, thepropodeum and
peduncle, black, legslemon
yellow except basal half to two-thirds of coxae black; abdo-men
brown-black above, shading tobrown on
the sides; funicular joints notmuch
longer than broad. In proportions of the head, sculpture,and
vestiture of the submarginal vein, thetwo
sexes are alike.Type.—
Female,U.S.N.M.
No. 42244. Alpine, Calif. (A. C.Kinsey).
The
allotype is with the type in the collection of the United States NationalMuseum.
Remarks. —
Describedfrom
the type, allotype,and
12 maleand
female paratypes rearedby
Dr. A. C.Kinsey
in the type localityfrom
the galls of Plagiotrochiis quinqueseptum var.on
Quercvs engehnanni.The
gallmaker
issuedon February
24, 1920.All the males of this species at
hand
are predominantly black, resembling closely thedark
variety dorsalis of D. querci-lanae (Fitch).The
females are darker than those of querci-lanae^ butdo
not reach the degree of blackness present in the male of either ofthesetwo
species.32. DECATOMA FOLIATAE Ashmead
Decatoma foliatae Ashmead, Can. Ent., vol. 13, no. C, p. 136, June, 1881.
Decatoma batatoides Ashmead, Can. Ent., vol. 13,no. 6, p. 136, June, 1881.
Eudecatoma batatoides Ashmead, Trans. Amer. Ent. See, vol. 21, p. 319, 1894.
(D. batatoides Ashmead designated type of new genus Eudecatoma Ashmead.)
Differs