MAMMALS OF ANGLO-EGYPTIAN SUDAN — SETZER 511 The color in a recently taken specimen from Kamisa is rather light

above

and

^{graduall}^}shading over thesides into the

plumbeous

based whitish hairs of the belly.

The

blackish tips of the hairs is

most

intense over the shoulders

and

diminishes in

amount

over the

rump and

on theupperlegs.

The

nose

and

a ring aroundtheeye areofpm'e color near

Cinnamon

Buff.

The

skull,

which

is of

an

adult male, is

quite long with

heavy

supraorbital ridges

and heavy

z3^gomata.

The

rostrum is short

and

wide

and

the teeth are relatively massive.

The

auditory bullae are small but well inflated.

The

specimensfrom

Khartoum

are

somewhat

palerthantheanimals

from

farther east.

Leniniscomysbarbariiszebra (Heuglin) Figure6,c

Mus

zebra Heuglin, Nova Acta Acad. Caes. Leop.-Carol., Halle, vol. 31, No. 7, p. 10, 1864. (Djurand Bongo, Anglo-Egyptian Sudan.)

Specimens Examined:

Thirty-four, from: Torit, 17; Ikoto, 1;

Nimule, 2; Malek, 8

(BM);

25miles east of

Lake No,

1

(BM);

Bahr- el-Ghazal, 1

(BM);

Badigeru

Swamp,

20 miles east of Mongalla, ¹

(BM);

Fashoda, 1

(BM); Dud

Majok, 1

(BM); Wau,

1

(BM).

Measurements:

Averages

and

extremes ^of three adult males

and

three adult females

from

Torit are, respectivel}^ as follows:

Length

of head

and body

102 (94-110), 101 (98-104); lengthoftail 112 (104-120), 110 (107-116); length^ofhindfoot 24.5 (24-25), 25 (25);length of ear 14 (14), 15 (14.5-15.5); greatest length of skull 28.0 (27.1-28.5), 27.7 (27.0-28.3); condyloincisive length 24.9 (23.9-25.6), 24.6 (24.0- 25.4); length of auditory bullae 5.4 (5.0-5.6), 5.4 (5.2-5.6);

crown

length of upper toothrow 4.9 (4.9-5.0), 4.8 (4.8-4.9); least inter- orbital width 4.5 (4.5), 4.5 (4.4-4.6); width of rostrum at level of antorbital foramen 3.6 (3.5-3.8), 3.5 (3.4-3.7); length of nasals 10.2 (9.8-10.6), 10.2 (9.8-10.5); greatest width across zygomatic arches 12.9 (12.6-13.2), 13.1 (12.5-13.5).

Kemarks: There

is an apparent sexual difference in size of body, lengthoftail, lengthofhindfoot,

and some

cranialcharactersin

which

thefemales generally average slightly largerthan the males. In gen-

eral, the color is quite uniform.

Three

of the specimens

from

Torit are only slightly paler than is the rest of the series,

and

the single specimen

from Wau

^{is of the}

same

color asthe majorityof the speci-

mens from

Torit.

Thissubspeciesof striped

mouse

seemsto

have

thelargestgeographic range of

any

of the rodentsfound in the Sudan.

The

explanation for

386559—56 ⁵

512 PROCEEDINGS OF THE NATIONAL MUSEUM

^vol.loe

this apparent lack of differentiation

must

await further specimens

from

intermediate localities or until a revision is

done on

thegenus as a whole.

.Lemniscomys dunni dunnt (Thomas) Figure6,c

ArvicanihisdunniThomas, Proc. Zool. Soc.London,p. 297,Aug. 6, 1903. (Kaga

Hills, western Kordofan.)

Specimens Examined:

Eleven, allin

BM,

from:

Umm Keddada,

3;

ElFasher, 2;

Juga

Juga, 15mileseast-northeastofElFasher, 1^;76miles east of

El

Fasher, 1; 110 miles east of El Fasher, 1; Kurra, Jebel Marra, 3.

Measurements: An

adult female

from Juga Juga and

an adult

male from

76 mileseast ofElFasher measure,respectively, as follows:

Length

of

head and body

116, 108; length of tail 142, 130; length of hind foot 27, 25; length ^of ear 15, 16; greatest length of skull 30.1, 30.1; condyloincisive length 26.9, 26.7; length ofauditory bullae 5.7, 6.2;

crown

length ofupper toothrow 5.0, 5.3; least interorbital width

4.5, 4.8; breadth of rostrum at level of antorbital

foramen

3.7, 3.4;

length ofnasals 11.7, 11.8.

Remarks: The

species

dunni

differs

from

L. barbarus in

markedly

lighter color, shorter anterior palatine foramina,

markedly

different shapeof the posteriorchoanae,

and

the general structureoftheptery- goidregion.

The

auditory bullae are smaller, the toothrow

somewhat

longer, anterior parts of the zygomatic arches tapering into skull insteadof flaringoutward,

and

thebraincaseis

more

extendedposterior to the posterior root of the

zygoma.

There

are three specimens in the British

Museum

without skulls

from

Kurra, Jebel Marra, that either represent a

new

speciesor are hybrids

between

L. dunni

and

L. lynesi.

From

the appearanceof the skins theyareintermediatein alldetails. I feelitbest to record these specimens

and

onlyrefer

them

toL. dunniprovisionally.

Lemniscomys dunninubalis

Thomas

and Hinton Figure6,c

LemniscomysdunninuhalisThomasandHinton,Proc. Zool. Soc. London,p.263, July6, 1923. (Talodi,

Nuba

Country, southern Kordofan.)

Specimens Examined:

Slx, all in

BM,

from: Talodi, 2;

Nuba

Mountains, 1; Delami, 1; Agur, 2.

Measurements: An

adult

male from

the

Nuba Mountains and an

adultfemalefrom Talodi measure, respectively, as follows:

Length

of head

and body

95, 90; length of tail 115, 112; length of hind foot23, 22; lengthofear13, 13;greatest lengthof skidl 28.4,?;condyloincisive

MAMMALS OF ANGLO-EGYPTIAN SUDAN — ^SETZER 513

length 25.5, ?; length of upper toothrow 4.9, 4.9; least interorbital width 4.3, 4.3;breadth of rostrum at level of antorbital

foramen

3.3, 3.3; length of nasals 10.4, 9.1.

Remarks:

L.d.nubalisis

somewhat

smallerthanthe

nominate

race

and

ispronouncedlybrighterincolor.

The

color ofdunniisa palebuff while nubalisisabrightgoldencolor.

The

sides

and

flanks,insteadof being wliite asin dunni, are

washed

with the golden ochraceous color of the dorsum.

Lemniscomyslynesi

Thomas

and Hinton

LemniscomyslynesiThomasand Hinton, Proc. Zool. Soc. London, p. 267, July6, 1923. (Central JebelMarra, Darfur.)

Specimens Examined:

Ten, allin

BM,

from: JebelMarra, 3;

Cen-

tral Jebel Marra, 6; South

Downs,

JebelMarra, 1.

Measurements: An

adult female from

South Downs,

JebelMarra, measuresasfollows:

Length

^of

head and body

101; lengthof tail 104;

lengthofhindfoot 23^;length ofear 14^; condyloincisivelengthof skull 23.9;

crown

length ofupper toothrow5.1^;least interorbitalwidth4.5;

breadth ofrostrum at level ofantorbital

foramen

3.6.

Remarks:

In the original description of this species,

Thomas and Hinton

suggested that it

was more

nearly related to L. zebra

{=L.

barbarus zebra) than to

any

^{other gi'oup in} the genus. In all cranial details, and,

most

strikingly,inthe distributionofthe dorsal lines

and

spots

and

indegree of spininess, L. lynesiis

most

closely allied to L.

striatus.

From aU

charactersstuaied it appeai-s as though this is not avalid species but merely asubspecies ofstriatus. I feelcertain that intergradation couldbe demonstrated^ifspecimenswere obtained

from

intermediatelocalities.

However,

untilsuch timeasspecimens

show-

ing intergradation areobtainedI prefertoletthe

name

stand as a fuU species.

The

skull, as well as the skin, is

markedly

different

from

dunni.

The

color isthe darkest of

any

ofthe kindsin theSudan.

The

skull differsfromthatofdunniinthe

markedly

differentshapeofthe bullae, in thelengthofthe anterior palatine foramina,

and

intheverydifferent structm'eof thepterygoidregion.

Lemniscomysmacculus macculus(Thomas and Wroughton)

ArvicanihismacculusThomasand Wroughton, Trans.Zool. Soc. London, vol. 19>

p. 515, March 1910. (Mokia, southeastern Ruwenzori, Uganda.)

Specimens Examined:

Three, from: Torit, ₁;Nimule,2.

Measurements: The measurements

ofanadult

male from

Nimule, are asfollows:

Length

of

head and body

96; length ^oftail 121^;length ofhindfoot 25; length of ear 16; greatestlength of skull 26.2; condy-

514 PROCEEDINGS OF THE NATIONAL MUSEUM

^vol.loe

loincisive length 24.2; length of auditory bullae 5.2;

crown

length of upper toothrow 4.8; least interorbital width 4.5; breadth of rostrum at level of antorbitalforamen 3.3; lengthofnasals 10.2.

Remarks:

In L. macculus the proportion of the least interorbital width to the width of the rostrum ranges from 71.2 to 73.3 percent (79.7 to 81.6 percent in striatus)

and

the proportion of the length of the nasals to the total length of the skull ranges

from

37.8 to 38.9 percent (36.0 to 36.7 percentinstriatus). It

may

be noted thatin the

firstproportion the size dijfference betweenL. macculus

and

L.striatus is apparent, butinthesecondproportion, thatofnasalsto totallength of skull, theratioisreversed,

which shows

that thenasals,even

though

measuring shorterin macculus,

occupy more

^ofthe totallengthofthe skull thanin striatus.

These two

proportions appear to hold

good on

other specimens of these

two

species in both the U. S. National

Museum and

the British

Museum

collections so that they perhaps will be valid for critically distinguishingthe

two

species

where

theyoccurtogether.

In general, macculus is a smaller animal than is striatus. In the former the hind foot usually measures less than 26

mm.,

while in the latter thehindfootisusually larger.

Lemniscomysstriatus massaicus(Pagenstecher)

Mus

{Lemniscomys) barbarus L. var. Massaicus Pagenstecher, Jahrb. Hamburg-

ischen Wiss. Anst., vol.2,p. 45,1885. (Lake Naivasha.)

Specimens Examined:

Thirty, from: Gilo,

Imatong

Mountains, 11 (4,

MCZ)

^; Torit, 2;Nimule, 5 (3,

MCZ)

; Katire,9 (AICZ);Mag\ve, 2 (AICZ); TalangaForest, Imatong Mountains, 1

(MCZ).

Measurements: An

adult

male and an

adult female

from GUo, Imatong

Mountains,respectively

measure

as follows:

Length

of

head and body

123, 107; length of tail 130, 130; length ^of hind foot 28, ?;

lengthofear 11, 12;greatestlengthof skull 29.4, 29.0; condyloincisive length 26.5, 26.2; length of auditory bullae 5.9, 5.0;

crown

length of upper toothrow4.9, 5.1^;leastinterorbital breadth,4.8, 5.0; breadth of rostrum at level of antorbital foramen 3.5, 3.8:length ofnasals 10.6, 10.8.

Remarks:

This species can easil}^ be confused with L. macculus.

There

are, however, several pronounced cranial differences between the

two

species. In the present series, the proportion of the least interorbital width to the width of the rostrum is greater;

and

the proportion of the length of th(; nasals to the total length of the skull is less in the larger species striatus.

MAMMALS

OF

ANGLO-EGYPTIAN SUDAN — ^SETZER 515