other
known
species.They
suggest Marginifera popei(Shumard) which
isabundant
inthe Glass, Guadalupe,and Delaware Mountains
ofTexas, Thisspecieswas
identifiedby McKee from
theKaibab
lime- stoneof theGrand Canyon,
Ariz.LIOSOTELLA
Cooper,new
genusShell,based
on known
species, rangingin sizefrom
smalltomoder-
ately large; strongly concavo-convex; hinge line wider than the shell at the middle; cardinal extremities auriculate. Anterior
commissure
narrowly uniplicate; pedicle sulcus shallow to deep; brachial fold usuallylow and
obscure. Surface costate; costae spinose.Umbonal and
juvenile partsof the valvessmooth
to indistinctlycostateand
with afew
scattered smallspines. Costaeon
anteriorand body
strong, bear- ing scattered erect strong spines.A row
of strong spines appearson
the steepumbonal
slopesand
overhangs the earswhich
are smooth.Pedicle interior with adductor muscles located
on
a strongmedian
elevation justposterior to themiddle; diductor impressions largeand
flabellate.
Concave
auricles walled offfrom
visceral regionby
alow
obliqueridge. Brachial interiorwithstout,short cardinal processhav- ingatrilobedmyophore. Median
ridge low, extendingslightlyanterior to the middle; marginal ridges not strongly developed; brachial im- pressionsprominent, occupyingthemiddle;inner surfacemore
orless deeplypitted.Genotype.
—
Liosotellarugosa Cooper,new
species.Discussion.
—
This genus is recognizableby
its externalform and
ornamentation suggesting large specimens ofAvonia
ormembers
of the Dictyodostits occidentalisgroup
of Dictyoclostus.Only
a casual inspection is necessary to distinguish Liosotellafrom Avonia
because of differences in details of the ornamentation butmore
particularly in thearrangement
of the spinesand
in the interior.The
spines of Avonia, at least as that genus is identified in thePermian
rocks ofNorth
America,are scattered irregularly over thesurface particularly in the sparsely plicated anterior region.Avonia
is notprovided with prominentears separatedfrom
thevisceralchamber
by awell-defined partition. Inside the brachial valve ofAvonia
the brachial ridges di- vergefrom
the hinge line, a situation entirely different in thenew genus under
discussion.Liosotella differs
from
Dictyoclostus in at leasttwo
respects.One
of the
most
important differences is the complete lack of reticulate ornamentationon
the posterior portionsofboth valves. Furthermore, the ears of Dictyoclostus usually arevery spinose.NO. 2 SPONGES,
BRACHIOPODA,
ETC.— COOPER
37LIOSOTELLA ANGUSTATA
Cooper,new
speciesPlate iiB, figures 5-10
Shell ofabout
medium
size for the genus, subrectangular inoutline with the width slightly greater than the length. Auriculate, but with thehingeonly slightlywider than themidwidth. Deeply concavo-con-vex
; visceral area shallow. Anteriorcommissure
slightly uniplicate.Surface
marked by
irregular,uneven
costae,rounded and
confined to the flanksand
the anterior slope. Spines scatteredand
smallon anter- iorslopebut largeand
stoutina curvedrow
atthebase of theumbonal
slopes.
Pedicle valve fairly evenly
convex
in lateral profileand
with themaximum
curvature inthemedian
region. Anterior profilesomewhat
quadrate but withthe pediclevalvedepressedinthe middle.Umbonal
region swollen;
umbonal
slopes steep.Median
sulcus originating just anterior to theumbo and
extending tothe frontmargin
; sulcus shal- lowand
broad,forming
awide
shallow trough. Flanks swollenand
rounded, withnearlyverticalslopes. Earssmall,gentlyrounded
;beak small, incurved.Brachial valve deepestinthe
median
regionwith steeplateralslopes but asomewhat
less steep anteriorslope.Fold
originatingin anterior half,low, poorlydeveloped. Cardinal extremitiesflattened.Interior
unknown.
Measurements
inmm. —
Holotype,length (crestofumbo
toanteriormargin)
21.8,midwidth
24.2, hinge width (basedon
halfmeasure
of 13.7) 27.4?, thicknessat middle7.2, surfacemeasure
of pedicle valve 40.0; paratype, length (crest ofumbo
to anteriormargin)
20.6, mid- width 22.3, hinge width 24.0 plus, thickness ?, surfacemeasure
of pedicle valve 37.0.Types.
—
Holotype,U.S.N.M. No.
115465; figured paratype,U.S.N.M. No.
115466.Horizon and
locality— Rare
intheAnidanthus
zone,loc.8o6n,806-0.Discussion.
—
This is the smallest of the species occurring in theMonos
Hillsand
can be distinguishedfrom
L. rugosa, withwhich
itoccurs, by its costation being confined to the anterior portion of the flanks
and
anterior slope.LIOSOTELLA RUGOSA
Cooper,new
species Plate loC, figure 9; plate iiC, figures 11-18Shell moderately large, subrectangular to subquadrate in outline, deeply concavo-convex; auriculate withthe hinge
forming
thewidest part;lateralmarginsgentlyconvex
;anterolateralextremitiesnarrowly38 SMITHSONIAN MISCELLANEOUS COLLECTIONS
VOL. II9convex
; frontmargin
gentlycurved.Umbonal
regionsmooth
orv^^ithscattered small spines; anterior two-thirds strongly costate, costae
numbering
2 or 3 in 5mm.
at the front margin. Costae bearing scat- tered erect spines; large spines also developed in a curvedrow
of6
spines diminishing in size posteriorly near the base of theumbonal
slopeand
overhangingtheears,which
are smooth.Umbonal and body
spines slender.Pedicle valve strongly
convex
in lateral profileand
with the greatest convexityinthemedian
portion. Anteriorprofilesomewhat
rectangu- larwith amedian
depression representing the sulcus,and
with nearly vertical sides.Umbo
moderately swollen; sulcusand
costation origin- ating5 to7mm.
anteriortothebeak; sulcus narrow, shallow,and
ex- tendingtothe anteriormargin.Umbonal
slopessteep;flanksbounding
sulcusnarrowlyrounded and
withnearlyvertical sides.Ears
rectangu- lar; narrowlyrounded
in section.Beak
small, strongly incurvedand
overhanging theumbo
of the brachial valve.Brachial valve deeply concave in lateral
and
anterior profiles;um-
bonal region deeply concave,bounded
posterolaterally by flattened areas corresponding to the auricles.Median
ridge low, subangular, extending anteriorly to the frontmargin from
a point about 5 to 7mm, from
the beak. Flanks deeply concave; inner concavitybounded by
the anteriormargins
strongly directedtoward
the brachial valve.Interior, brachial valve with small delicate cardinal process having smalltrilobed
myophore and
shortshaft.Measurements
inmm. —
Holotype, length (crestofumbo
toanteriormargin)
26.0,midwidth
31.7,hinge width,35.7, thickness 8.2,surfacemeasure
of pedicle valve 52.0; paratype(U.S.N.M.
115459a) : length (crest ofumbo
to frontmargin)
28.2,midwidth
29.6, hinge width (based on halfmeasure
of 17.8) 35.6, thickness 5.8, surfacemeasure
of pedicle valve 50.0.Ty/>^.r.—Holotype,
U.S.N.M.
115458;figuredparatypes,U.S.N.M.
115459a, I.G.M.;unfigured paratype,
U.S.N.M. No.
115459b,Horizon and
locality.— Rare
inComposita
zone, loc. 806c, 8o6h, 8o6r,8o6v
; Dictyoclostus zone, loc. 806k',8o6x
; Leiorhynchoidea- Cancrinella zone, loc. 8o6i;Anidanthus
zone, loc.806m, 8o6n
; Spiri- ferellinazone, loc.8o6g.Discussion.
—
This species is characterizedby
its prominentmedian
sulcus, fairlystrong fold
on
the brachialvalve,and
prominentanterior costation.The
species differsfrom
stronglymarked members
of L.suhrugosa,
which
occurin higher strata, in thegreaterprominence
of the sulcusand
fold, a less strongly archedumbo and somewhat more
prominent costation. Itis possible that specimens referred to L. siih-NO. SPONGES,
BRACHIOPODA,
ETC.— COOPER 39
rugosaoccurring in theComposita and
Spiriferellinazones should be referred to L. rugosaand
that thesubdued
costation of thelatter, as well as the apparentlymore
archedumbo,
are features of poor pres- ervation. Unfortunately sufficient specimens are not available in the collectionsmade
to settle this point.LIOSOTELLA SUBRUGOSA
Cooper,new
speciesPlate IIA, figures 1-4; plate 12B, figures 9-17
Shell moderately large, transverselysubrectangular in outline,auri- culate,withthehinge
forming
thewidestpart;deeply concavo-convex.Lateral margins gently
rounded
; anterolateral extremitiessomewhat
narrowlyrounded
; anteriormargin
nearly straight, faintly uniplicate.Umbonal
region nearlysmooth; umbonal
slopes, anterior halfand
flanksmarked
by low, rounded, irregular costae. Spines few, erect, scatteredon
surface of venterand
in arow
of6
large spines located alongthe base of theumbonal
slopeand
overhanging the auricles.Pedicle valve strongly
convex
in lateral profile with the greatest convexity located in themedian
region. Anterior profile strongly arched, withnearlyvertical sidesand
slightly bilobatecrest.Umbonal
region moderately strongly swollen; beak small
and
overhangingbra- chialumbo.
Sulcus originating lo to 15mm.
anterior to the beak,shallow with a
narrow median furrow and
broadly sloping sides, ex- tending tothe frontmargin. Flankstumid
withsteepsides. Anterior slopeconvex, verysteep.Ears
large,narrowlyrounded
ontheextrem-ity
and
in section.Brachial valve deeplyconcavein lateral
and
anteriorprofiles;umbo forming
aconcavity within themain
visceralregion;concavity deepest in themedian
region. Fold originating near the middle,low and
not strongly defined. Sidesforming
a steep rimaround
the central concavity.Interior of pedicle valve with
umbonal
cavity thickenedby
callus;diductor impressions large, located near the middle of the valve; ad- ductor field large, elongated, elevated
on
a low,wide
ridge extending nearly to the middle. Interior of brachial valve with short, stout- shafted cardinal process having a trilobatemyophore. Median
ridge low.Adductor
scars crenulated, located posteriortothe brachial ridgeswhich
occupythemiddleregion. Marginal ridgeslowand
indistinct.Measurements
inww.—
Holotype, length (crest ofumbo
to frontmargin)
29.4,midwidth
32.5, hinge width 34.0 plus; thickness at middle 12.5, surfacemeasure
of pedicle valve 58.0.Ty/'^.y.—Holotype,
U.S.N.M. No.
115460; figured paratypes,U.S.N.M.
Nos. 115461-115463, I.G.M.40 SMITHSONIAN MISCELLANEOUS COLLECTIONS
VOL. IIQHorizon and
locality.—
Spiriferellina zone, loc, 8o6b, 8o6d, 8o6d', 8o6f, 8o6g,8o6h^
8o6s,8o6w.
Discussion.
—
This species as definedis characterizedby
its strongly archedumbo,
itssubdued
ornamentation, sulcuson
pediclevalve shal- low, faint foldon
brachial valveand
large thick cardinal process. Inallthese respects it differs
from
thewell-marked specimens occurring in theAnidanthus
zone.The
distinctionsnamed,
as already pointed out, are notclearlymarked
in all specimens buttheend members
are quite distinctive. It is possible that a large collection of better-pre- served specimens willshow
L. rugosaand
L. subrugosa to be one species. Collecting of these fossils is very difficultand
it is doubtfulifbetter-preservedmaterialistobe
found
in theMonos
Hills. Conse- quently the question cannotnow
be answered.LIOSOTELLA MAGNIRUGOSA
Cooper,new
species Plate 12A, figures i-8Shelllarge,subquadrate inoutline;strongly auriculate
and
with the hingeforming
the greatestshellwidth. Sidessloping gentlymedially;
anterolateralextremities narrowly
rounded
; anteriormargin
indented inthemedian
region. Anteriorcommissure
gentlyuniplicate. Surface paucicostate, theflanks, venter,and
anterior slopesmarked by
strong, narrowlyrounded
costae, about20
innumber, and
afew
of the larger ones bifurcating near the anterior margin. Spineslarge, scatteredon
the anterior slopeand an
obliquerow
of 3 ormore
located atthe base of theumbonal
slope.Umbonal
region withoutornament.Pedicle valve strongly
convex
in lateral profileand
with the greatest convexitylocatedinthemedian
part. Anteriorprofileslightlybilobate.Umbonal
region swollen, tapering toa small beak thatoverhangs the brachialumbo. Umbonal
slopes rounded, steep. Auricles large, well rounded.Median
sulcus originatingabout15mm.
anteriortothe beak, extending to the front margin, broadand
shallow, occupying about one-third the width at the front margin. Flanks narrowly rounded, with vertical sides tothelateralmargins. Anteriorslope very steep.Brachial valve
most
deeplyconcave inthemedian
regionand
with theumbo
strongly concave. Anteriormargins
strongly deflected bra- chiallyand
with steep inner margins. Fold low, broad, not strongly developed; ears strongly concave.Details of theinterior
few
; cardinal process with smallmyophore
;
brachial ridges strong, widely spaced.
Measurements
inmm. —
Holotype,length (crestofumbo
toanteriormargin)
29.8,midwidth
29.6, hinge width41.0 (restored), thicknessNO. 2 SPONGES,
BRACHIOPODA,
ETC.— COOPER
4Iatmiddle9.6,surface
measure
of pediclevalve,beak
toanteriormargin
51.0,depth23.0;paratype (I.G.M.) :length (crestof
umbo
toanteriormargin)
25.7,midwidth
25.5, hinge width 24.0 plus, thickness 8.5, surfacemeasure
of pedicle valve 46.5, depth 15.6.Types.
—
Holotype,U.S.N.M. No.
115464;figuredparatype, I.G.M.Horizon and
locality.—
Leiorhynchoidea-Cancrinellazone, loc. 8o6i.Discussion.
—
This species is notuncommon
in the Cancrinella zone but specimens are difficultto prepare. Consequently afew
specimens onlyareavailablefor study.As
revealedby
thesespecimensthe species is characterizedby
its large size, strong ears, swollenumbo,
broad, shallow sulcus and, consequently, broadand low
foldon
the brachial valveand
finallyby
the strongcostae. In comparisontoL. rugosathe foldand
sulcus of L.magnirugosa
are lesspronounced and much
broader, the costae are
much
stronger, theumbo
issomewhat more
swollen
and
the earsmore
extended.MUIRWOODIA
species Plate gA, figures 1-7Shell large for the genus, strongly concavo-convex, subquadrate to subrectangularinoutline;auriculate,withthehinge
forming
thewidest part. Anteriorcommissure
narrowly uniplicate. Lateral margins con- cave anterior to the auricles, narrowlyrounded
at the anterolateral extremitiesand
emarginateatanteriormargin. Surfacemulticostellate, costellaenarrow, rounded,separatedby
striaesomewhat narrower
than the costellae. Costellaenumbering
about 10 in 5mm. on
thebody
of the shelland
along the front margin. Spines not preserved.Pedicle valveveryunevenly
convex
in lateral profilewiththevisceral area gently convex. Anterior profile bilobate.Umbonal and
visceral regions moderately swollen;median
sulcus originating 3 to 5mm.
anterior to the beak, narrow, deep, widening
and
deepening to the anteriormargin. Flankson
visceral areamoderately swollenand
with short concave slopes to the cardinal extremities. Geniculation occur- ring20 mm.
anteriortothebeak;geniculatedportionnarrowlyrounded
withtumid
areasbounding
themedian
sulcus. Traillong, deeplysul- cateand
with well-roundedflanksbordering thesulcus.Umbonal
sur- faceand
trail surface approximately parallel. Lateral slopes steep.Beak
small, protruding slightly posterior to the posterior margin.Brachial valvepoorlyexposed infour available specimens butindi- catingashort
and
shallowvisceralareaand
asharply geniculated por- tion closely fitted to the trail of the pedicle valve.A
sharp butlow
foldis indicated by thedeep sulcus.42 SMITHSONIAN MISCELLANEOUS COLLECTIONS
VOL. II9Measurements
inmm. —
Figured specimen, length (crest ofumbo
to anterior
margin)
27.6,midwidth
41.0, hinge width 47.0 plus (au- ricles not preserved), thickness ?, surfacemeasure
of pedicle valve 62.0.Figured
specimens.— U.S.N.M. No.
115568, I.G.M.Horizon and
locality.—
Dictyoclostus zone, loc. 806k, 806-I. Lei- orhynchoidea-Cancrinellazone, loc. 8o6i.Discussion.
— The
specimens figuredand
described herein probably represent anew
species of this interesting genus but are too poor to describe asnew.
This genus has hitherto not been reportedon
this continent but itis representedby
severalspecies.Productus
genicula- tus Girtyfrom
thePark
City formationand
P. multistriatusMeek
ofthe Phosphoria formation are representatives. Dictyoclostus
deminu-
tivus
Cloud from
thePermian
of Coahuila, according to descriptionand
figures,belongs tothegenusMuirwoodia.
The
specimensfrom Monos
Hills aremost
similar toMuirwoodia
midtistriata(Meek)
butdiffer inhaving amuch
lessinflatedumbonal and
visceral disc region, a stronger, deeper,and more
angular sulcuson
the pediclevalveand
amuch
strongerdegreeof geniculation.WAAGENOCONCHA MONTPELIERENSIS
(Girty) Plate 13, figures 1-14ProductusmontpelierensisGirty,U.S. Geol. Surv.Bull. 436,p.30,pi. 2, figs.5,6, 1910; U.S. Geol. Surv. Prof. Pap. 152,p. 80, pi.28, figs. 12,13, 1927.
Pustula montpelierensis (Girty) Branson, Univ. Missouri StudiesQuart.,vol.5,
No.2, p. ^2, 1930.
Shell large, variable, slightly wider than long in the young, with length
and
width subequal in theyoung
adultand
with the lengthforming
the greatest dimension in old specimens.Hinge narrower
than the greatest widthwhich
is located at or anterior to the middle.Cardinal extremities rounded. Nearly planoconvex in section. Sur- face
marked by
fine,elongated pustuleswhich,inlife,bore veryslender spines; spine bases arranged in quincunx, thearrangement
generally cleareron
the brachial thanon
the pedicle valve.Growth
undulationscommon on
the surfaces of old specimens.Pedicle valve strongly
convex
in lateral profileand
with themaxi-
mum
convexitylocated in theumbonal
region. Anterior profile form- ingtwo somewhat
narrowlyrounded
lobes separated by a shallow sulcus.Umbonal
region swollenand
protrudingconsiderably posterior totheposteriormargin.Beak
small, stronglyincurvedand
overhang- ing the hingeand
asmall portion of the brachialumbo.
Sulcus origi-NO. 2 SPONGES,
BRACHIOPODA,
ETC.COOPER 43
natingjustanteriortotheumbo and
about 7 to lomm.
anteriortothe beak. Sulcus strong inyoung
shells,with abroadlyV-shaped
section, anteriorlybecoming somewhat
less deep in adults but presentfrom umbo
to frontmargin
at all ages. Flanksbounding
sulcus with steep tomoderately steep slopesinto the sulcus,with narrowlyrounded and
inflatedcrests
and
steeptonearlyvertical slopestothelateralmargins.Brachial valve nearly flat in the posterior two-thirds but strongly geniculated in a brachial direction in the old adults.
Umbonal
region concaveforabout5mm.,
afterwhich
a low,broad fold originates that extendstothe anteriormargin.Low,
oblique foldsbound
theconcaveumbonal
regionand
extend anterolaterallyabout 10mm.,
afterwhich
they disappear. Cardinal extremities gently sulcate between the car- dinalmargin and
the oblique folds referred to above. Lateraland
anterior margins of adults geniculated brachially,about 20 to 25mm.
anterior to the beak, the geniculation extending for a considerable distance inold adults.
Interior of pedicle valve with broadly flabellate diductors as usual inthe Productacea. Brachial valvewith large cardinal process having a longshaft
and
trilobatemyophore
stronglyrecurvedintotheumbonal chamber
of the pediclevalve.Median
ridge slender, extending forat leasttwo-thirds the length of thevalve. Place of geniculationmarked by
a curvedrow
of strongbut short spines.Measurements
inmm. — Hypotypes: (U.S.N.M. No.
115515), length (crest ofumbo
to frontmargin)
26.9,midwidth
30.5, hinge width ?, thickness at middle 12.3;(U.S.N.M. No.
115512), length(umbo
tofrontmargin)
33.5,midwidth
35.0,hinge width25.0?,thick- ness at middle 9.0(somewhat
crushed) ;(U.S.N.M. No.
115511), length (crestofumbo
to frontmargin)
42.6,midwidth
(basedon
halfmeasure
of 23.0) 46.0, hinge width 33.5, thickness at middle 14.2;
(U.S.N.M. No.
1155 10), length (crestofumbo
tofrontmargin)
53.0,midwidth
42.0 hinge width 28.0?, thickness atmiddle 22.3.Hypotypes.—
U.S.ISIM, Nos. 115510-115513, 115515-Horizon and
locality.— Anidanthus
zone loc. 8o6n, 806-0; Dictyo- clostus zone, loc. 806k, 806k', 806-I, 8o6p, 8o6z; Leiorhynchoidea- Cancrinella zone, loc. 8o6i;Composita
zone, loc.8o6h
; Spiriferellina zone,loc. 8o6b, 8o6d, 8o6d',8o6f, 8o6g.Discussion.
— Waagenoconcha
montpelierensis, as interpreted herein, ismore
broadly viewed than hitherto. Girty's type specimen is that of a small shell butmay
be ayoung
specimen.Young
individuals of theMexican
specimensconform
almost precisely to Girty's descrip- tionand
figures.These
specimens, however, are clearly theyoung
of44 SMITHSONIAN MISCELLANEOUS COLLECTIONS
VOL. II9 largerones. Ingeneral the geniculationatthefront of the valves takes place about20 mm.
anterior to the beak of the brachial valve but afew
specimens geniculate at a slightly earlier stage. Thismay
be apremature
assumption of adult characters or these specimensmay
represent another species.
Enough
specimens to prove this point are not available for study.A
fairlycomplete series of stages is present to indicate thatyoung
having the characters ofW.
montpelierensisgrow
toa large size,asize largerthan hitherto suspected.LEIORHYNCHOIDEA CLOUDI
Cooper,new
species Plate 14C, figures16-31Pugnax weeksinobilis Girty, U. S. Geol. Surv. Bull. 436, p. 32, pi. 3, fig. 7 (not5,6), 1910. R. E. King,Univ. TexasBull. 3042,p.105, pi.33, figs.9-11, 1930.
Shell large, suboval in outline, with length
and maximum
width nearly equal. Posteriormargins
gently convex, anterolateral extremi-ties
somewhat
narrowlyrounded
; anteriormargin
broadly convex.Anterior
commissure
broadlyuniplicate. Surfacecostate, costaebroadand
subangular,2costaeoccupyingthesulcus, 3marking
thefold,and 4
to 5presenton
theflanks.Pedicle valve with gentle curvature in lateral profile, the greatest curvature locatedinthe
umbonal
region; anteriorprofile very broadlyand
gentlyconvex.Umbonal
region swollen,smooth
; costationorigi- nating6
to8 mm.
anterior to the beak. Sulcus originating at about the middle, shallowand
broad. Flanks with long anteromedian ex- tremities that are moderately elevated. Flanks gentlyconvex
posteri- orly butsomewhat
flattened anteriorly.Beak
strongly incurvedand
closely pressed onto the brachial
umbo; foramen
minute, permeso- thyrid.Beak
ridges elongate but false areas very narrow. Deltidial plates conjunct butshowing
line of junction, flat to concave.Brachialvalvein lateral profilegently
convex
withthe greatest con- vexity in theumbonal
region; anterior profile very broadlyconvex
butmore convex and
deeper than the pedicle valve.Umbonal
region gently swollen,smooth.Fold
originatingataboutthemiddle, flattenedand
not risingabove the flanks except in the anterior region. Flanks gentlyconvex and
with long anterolateral extensions tomeet
the re- entrantformed by
the anteromedian elevation of the flank of the pedicle valve.Beak
incurved under,and
hidden by, the beak of the pediclevalve.Pedicle interior with short, shallow delthyrial cavity; dental plates reducedto