The following paper was accepted
forpublication:DESCRIPTIONS OF NEW PSENID WASPS FROM THE UNITED STATES.
[Hymenoptera; Psenidae.]
BY
S. A.ROHWER.
If in
general
Dr.Ashmead's
classification ofthe Sphecoidea
isto
be
followed,and the genera Pscn and Pemephredon,
asunderstood by
Latreille (Free. car.gener
Insect,1796)
are tobe united
inthe same
family,the name should be Psenidae and not Petnphredonidse,
as Dr.Ashmead has
it,because Psen was described on page 122 and Pemphredon on page
128.According
tomany authors these two genera have been made the types
oftwo
distinct families,and by some have been widely
separated.Such wide separation has been due
tothe
fact thatsome authors have considered
cubitalvenation
tobe
ofthe
greatestimportance. This view
ofthe
greatimportance
ofvenation
isgradually being given up and students
arelooking
tothe body
itself forthe
principalcharacters and
treatingthe appendages
secondly.Such
a
method
ofstudy
willno doubt bring about
amore natural
classification. It is,however,
tobe expected
that inmany- perhaps
amajority
casesthe characters
ofthe appendages
will correlate so well
with the body characters
that a satisfac- toryarrangement could be
builtup on characters
easilyseen on the appendages. But
untilsuch
relationshipshave been
fullyestablished
the body should be
carefully studied.Whatever may have been the
origin ofthe Pseninae and Pemphredonina^,
therecan be but
littledoubt
that atthe present they
are closelyenough
related tobelong
tothe same
family.
An examination
ofthe type genera
ofthe two sub-
families willshow
thatthe thorax and abdomen
aremuch the same, but
thatthe head
ofPemphredon
islarger,and has the antennae
inserted close tothe
clypeus.In the wings Pem- phredon
differs inthe
loss ofthe
firsttransverse
cubitus. All these differencesdo not hold throughout the two
subfamilies,however,
for inAmmoplanus,
Diodontits,and
insome
speciesof
Passalcvcns
thehead
is smaller,and has more the shape
of speciesfound
in Pseninse.The
position ofthe antenna
1 isas inPemphredon, and
at leastone
transversecubitus
iswant-
ing.
The
difference invenation between the types
ofthe two
subfamilies
isnot
asgreat
asthe
differencebetween Stignius
and Pemphredon, both belonging
tothe Pemphredoninae.
100 PROCEEDINGS ENTOMOLOGICAL SOCIETY
For
inStigmns, and
allies,the second recurrent nervure
iswanting; and the presence
orabsence
of a recurrentnervure
is of far
greater importance than the presence
orabsence
ofone
ofthe transverse
cubiti.As
far asknown the
habits ofthe two subfamilies
aremuch the same.
The above remarks should
justifythe uniting
ofPsen and Pemphredon
inthe same
family.Psenulus (Neofoxia)
sayi,new
species.Belongs near
tibialis Cress.,but the sculpture
ofthe dorsu- lum
is not as strong, themesopleura
is sparselypunctured,
not striato-punctate,the head more
shining,and the abdomen with
abroader
redband.
Female
Length (imm.
Anterior margin ofclypeus gently rounded, notdentate ornotched; frontcloselypunctured, vertexand posterioror- bits shining and very sparsely punctured, carina from anterior ocellus complete and distinct; a depression beside each lateral ocellus; dis- tancebetweenthelateral ocelli slightly greaterthan thedistancebetween one ofthem
and the nearest eye margin; flagel robust, the first joint slightly longer than second; dorsulum and scutelwith distinct, wellsep- arated punctures;mesopleura
similarly punctured; enclosure well de.fined, with
some
oblique stria*;furrow of the posterior face deep; rest ofthemetathorax finely striated,with a tendency toreticulationsonthe sides; thirdtransverse cubitus sinuose;secondcubitalnarrowed aboutone"half above;femorarobust;
abdomen
elongate,petioleshorter than therest of the segment, flattened above: pygidium narrowed slightly apically, finely granular. Black; flagel beneath and tegulajtestaceous; fourante- rior tibiaeand tarsi, hind tibiae and tarsiat thebase whitish; allofthe second andthird abdominal segmentsbright red.Wings
clearhyaline, iridescent; venationdark brown.Type
locality:Onaga, Pottawatomie County, Kansas(Creve-
coeur).Type: No. 12356, U.
S.National Museum.
This
isthe
speciesrecorded
asMimesa pauper from Texas by Cresson
inHymenoptera Texana, but
it ismuch
differentfrom
that species, invenation and sculpture
as definedby Fox and Packard.
Psenulus (Neofoxia)
brevipetiolatus,new
species.Very
distinct inthe
short petioleand simple
clypeus.Female. Length!>
mm.
Middleof the anteriormarginoftheclypens truncated, thesides rounded, without teeth; a line-likefurrowfromante- riorocellusto baseofclypeus; frontfinelygranular; vertexandposteriororbitspunctured, thevertex
more
finely so; flagel robust, the firstjoint distinctlylonger than the second; dorsulum andscutel with distinct sep- arated punctures;mesopleunt
granular, the central partmore
finely so;enclosure with irregular strut; metathorax granular with a fine rugose appearance on the sides, middle furrow distinct; femora very robust, hind tibia1 arcuate inthe middle
when
seen from above; second cubital stronglynarrowed
above: venationas inNeofoxia; petioleshorter than the remaining part of the segment, widening apically,flat above with the marginsdistinct:abdomen
narrow,elongate: pygidiumvery narrow, fullythree times as longas wide, coarsely granular. Black; flagel tes- taceous; mandibles, apex of second,and most
of the thirdsegment
rufous; teguht, turbercles,
and
four anterior tibiae and tarsi whitish:hindtarsi brownish.
Wings
hyaline, strongly iridescent; venation dark brown.Tvpc
locality:Los Angeles County,
California, (Coquillett)-Type: No. 12355^ U.
S.National Museum.
Psen (Mimesa)
similis,new
species.Belongs
togroup nigra and
in thelong antennae
is related to longicornisand
mcllipcs,but
isperhaps nearer
tonigra
ormi\ia. The slender
antennae, striato-punctate vertex,narrow distance between
the lateral ocelli,almost impunctate meso-
pleuraeand dorsulum, the
large,smooth space
inthe
enclosure,and
slightlydusky wings
will serve todistinguish thisfrom
its allies.Male. Length alittle
more
than 5mm.
Anterior marginof the cly-peusnotched; supraclypeal areafinelygranular;frontalcarinanotstrong butpresent; head finely striato-punctate, the punctures in
same
places more abundant than thestriae; distance between thelateral ocellimuch
lessthan the distancefrom one of
them
to the nearest eye margin; an- tennas a little longer than thehead
and thorax, simple, not clavate, third jointabout thesame
length as fourth; dorsulum, scutellum, and mesopleurae shining, with only a fewwidelyseparated small punctures;enclosure with a few slightly oblique striae, the diamond-shaped area largeand smooth; rest of the metathorax except the finely granular pleura1finely reticulate; legsandvenation normal,petiole trisulcate,about the
same
length as the hind femora;abdomen
broadened and not es- peciallyelongate. Black; the four anterior tarsi pale brown, silvery pubescence not dense.Wings
slightly dusky hyaline, iridescent,vena- tiondarkbrown
Type
hca/ity: Florissant,Colorado, July
17,1907
(S.A.
Rohwer).
Tvpe: No. 12357, U.
S.National Museum.
102 PROCEEDINGS ENTOMOLOGICAL SOCIETY Psen (Mimesa)
floridana,new
species.Entirely black and runs
inwith regularis
orIcncopus
inFox's
tables,but
isnot
related closely tothese
or toany North American
species described.In many ways the fauna
ofthe
coast
southeastern region
ofFlorida
is distinctfrom the
rest ofthe United
States,but has
relatedforms
inthe neighboring
islands. Dr.
Ashmead had given
this speciesthe manuscript name
floridana,and
it isused on
thataccount.
Female. Length7.5
mm.
Anterior margin of the clypeus rounded) slightly irregular medially; front striato-punctate, vertex and posterior orbitswith small separate punctures; frontal carina not reaching the clypeus; ocelli in atriangle, thedistance between the lateral ones less than the distance from oneofthem
to the nearesteye margin; flagel ro- bust, thefirstjoint distinctlylonger thanthesecond; dorsulum striato- punctate; scutelwith widelyseparated, distinct punctures, mesopleurae shining, with small separated punctures; enclosure with afew strong rugae, asmooth
diamond-shaped areaapically; metathorax above retic- ulate,belowand on thepleurae granular; petiole strongly trisulcateand longer than thehind femora; secondcubitalnarrowedmore
than a half above; legsnormal; pygidium narrowedapically, and with large sepa- rate punctures. Entirely blackexcept the whitish anterior tarsi; wings duskyhyaline,iridescent, venation black.Type
locality:Biscayne Bay,
Florida.Type: No.
12360,U.
S.National Museum.
The sculpture
of thisspecies isvery
fine,and above
descrip- tionwas made from magnification
of35 diameters.
Psen (Mimesa) erythopoda, new
species.Belongs
togroup Kohli
ofFox and may be
easily recog-nized by the rufous four
posteriorfemora, the
striated enclos-ure
atthe base
ofmetathorax, and smooth mesopleurse.
Female. Length 10
mm.
Clypeus rather stronglyconvex, the ante- riormargin broadly notched medially; mandiblesbroad andvery robust:deep furrowsnearly encircling thelateralocelli: head shining with afew rather small punctures, the punctures closer in the postocellar area, dorsulum with larger punctures, posteriorly thesepunctures
become
con.fluentand irregularstriae take the place of the punctures; scutellum with well-separated punctures; mesoplurae and metapleurae shining,al- most impunctate: the enclosure with strong, longitudinalstrut; posterior faceof the metathorax rugoso-renticulate; petiole longer than the hind femora; hind tibiaewith shortspines, its inner spurslightly flattenedand broadened medially;
prygidum
long, well defined, granular, a seriesof punctures aroundthe margin. Black, with the usual silvery pile; basalhalf ofantennas, including scape, tegulae, anterior tibia;-, four posterior femora andtibiae(themiddlefemoraare blackishbasally), red; tarsi pal-
lid; wingshyaline,iridescent,venation pale brown.
Type
locality:Great
Falls, Virginia,August 22
(F.Knab).
Type: No. 12362, U.
S.National Museum.
Psen (Mimesa) punctata
v. Carolina,new
variety.Female. Length VI
mm. Very
\\\aepttnctata Fox,butis larger,the basal jointsof the flagel, the tibiae and tarsi, and petiole are red, the pubescenceof the faceis slightly golden, and the facialquadrangle is slightly narrower.T-vpelocality:
Raleigh, North
Carolina,September
12, 1905.Type: No. 12363, U.
S.National Museum.
Psen (Mimesa)
coquilletti,new
species.Related
toproximo.
Cress.,but the
distancebetween
the lateral ocelli is greaterthan the
distancebetween one
ofthem and the
nearesteye margin, not equal
to it;enclosure has
amedian furrow; head
isnot
"closelypunctured"; and other
characters.The
speciesdescribed by Cameron, colorado-
cnsisand
interstitialis,have the clypeus
different,and the male
differsfrom Cameron's
description inthe metathorax not being
reticulate,the
tarsiarenot
tectaceous, etc.Female. Length a little overit
mm. The
sharply defined produced middleportion of the clypeus with low roundedteethinthe middle, front closelypunctured, vertexandocciput shining, withwidelyseparate punc- tures; nofrontal carina,although a broad depressed line extendsback-ward
from the anteriorocellus; adepressed areaatthe sideof each lat- eral ocellus; flagel stout, thefirstjoint distinctly longer than the fourth;dorsulum
and
scutel dull, with widely separate punctures; mesopleura- granular; enclosure not sharply defined, with slightly oblique stria:, amedium
furrow extendingdown
the posterior face; metathorax with rather strongstriaedorsally ontheplurae,the striae are weak; second re- current nervure interstitial with the second transverse cubitus; second cubitalnarrowed abovealittlelessthanhalf; petiole stout, alittleshorter than the hind femora, nearly flat above; pygidium rather broad, with distinctpunctures;abdomen
elongate. Black; flagel and teguhi- testa- cous; tarsi andpartofthe tibiae brown;most
of abdominal segments two and three dull red.Wings
clear hyaline, iridescent: venation dark- brown. Silverypilerathermore abundant
than usual.Male.
What may
bethemale has the clypeus gently rounded and simple, the dorsulummore
shining, and the flagel darker.T\pe
locality:San Diego County,
California, female;males Los Angeles,
California (Coquillet);and two
males,Southern
California, inthe Fox
collection.Type: No. 12861, U.
S.National Museum.
104 PROC. ENT.
SOC.WASH.,
VOL,. XII, 1010.Ceratophorus
gennelli,new
species.Related
to tcna.\(Fox), but may
easilybe distinguished by the smooth,
itnpunctateenclosure
ofthe metathorax, more
sparsely
sculptured head and dorsulum,
shorter tibial spurs,and other
characters.Female. Length about(5
mm.
Anterior margin of the clypeusshal- lowly tridentate; headwith small separate punctures, notmuch
thicker onthefront; thirdantennaljoint slightly longer than the fourth; spacebetween
thelateral ocellimuch
lessthan the distancefrom oneofthem
to theinnereyemargin; dorsulum
and
scutellum sculptured similarto the head; mesoplurnesomewhat more
coarsely sculptured: enclosureof themetathorax smooth, shining, impunctate, except at the base,which hasshort, longitudinalstriae: the rest of the metathorax with small, irregular punctures,median
furrow deep; petiole rather stout, a little longer than halfof the hindfemora, above with rather large, irregular punctures; legsandvenation normal;abdomen
smooth,shining; pygidial area with separate punctures. Entirely black; venation black,wings dusky; hair gray.Type
locality:Pasadena,
California,May
31,1909
(F.Grinnell, jr.).
Type: No. 12865, U.
S.National Museum.
Passaloecus melanognathus, new
species.Easily known from other described Nearetic
speciesby the black mandibles.
Female. Length