Vol.
XXIV,
pp. 175-178 June 16, 1911PROCEEDINGS
OFTHE
BIOLOGICAL SOCIETY OF WASHINGTON
THE LOUISIANA PUMA.
BY
N.HOLLISTER.
[PublishedbypermissionoftheSecretaryoftheSmithsonianInstitution.]
An attempt
to identify thespecimens
ofpumas
in the collec-tionof the
United
StatesNational Museum has
resulted in the discovery that theform found
inLouisiana
isvery
differentfrom any
yet described.This new form
is represented in themuseum
collectionsby
threespecimens; two
skullsfrom Mer Rouge, Morehouse
Parish, collectedby
J. Fairiemany
years ago,and a
skinand
skull in the BiologicalSurvey
collectionfrom
Vidalia,Concordia
Parish.The Louisiana puma
ismuch more
closely relatedto thepuma
ofFlorida than
tothenorthern form
or to Felis oregonensisazteca. It isreadily separablefrom both
thefirstand
the last speciesby very marked
color differ- ences.The museum
is fortunate in possessingtwo
skullseach
of Feliscouguar from New York and
Felis corgifrom
Florida, as well asgood
seriesof thewestern
species.The
pelage ofthe Florida puma has been
carefully describedby Mr. Chas.
B.Cory,* by Mr. Outram Bangs,! and by Dr.
C.Hart Merriam,+
and Mr. Bangs has proved
that it varieslittle in colorwith
the season.From
the recordsoftheoccurrence
of thepuma
in theSouthern
States collectedand published by Dr.
F.W. True
§ itseems
certain that the distributionbetween Florida and
Louis-iana was
atone time continuous. As
so little isknown
of theanimal formerly inhabiting
that region,however, and
as theLouisiana puma
differs sogreatlyfrom
theFlorida animal
incolor it
seems
best to regard itat present as adistinct species.*Hunting andFishingin Florida,p.109, 1896.
tProc. Biol. Soc.Washington,XIII, pp.15-17, 1899.
tProc.WashingtonAcad.Sci., Ill,pp.583-585, 1901.
§ReportU.S.Nat.Mus.,1888-'89,pp. 591-608,1891.
31—Pkoc.P.iol.Soc.Wash.,Vor..XXIV,1911. (17.-,)
176
Hollister— The Louisiana Puma.
Felis
arundivaga
sp. nov.Type from 12milessouthwestofVidalia, ConcordiaParish, Louisiana.
No. 137,122, U. S. National
Museum
(BiologicalSurveyCollection), skinand
skull ofd\
adult(occipito-sphenoid sutureentirely closed); collectedJune
17, 1905, by B. V. Lilly.Generalcharacters.
— A
largepuma
of theFelis coryi type but lacking the bright ferruginous color; skullmuch
larger than in Feliscouguar, with the verybroadnasalsand highlydevelopedsaggital crest of F. coryi.Differs externally from Feliso. azteca in its darker coloration and very pronouncedcaudal stripe.
Colorof type.
—
Upperparts, including outersides oflimbs, grayishfawn color with a decided cast of ecru drab, especially on flanksand
legs;nape, withers,
and
an indefinite stripedown
backtorump
brighter, pale rufous; the entire upperparts finelymixed
with dusky, the darkercolor of the hair tips. Face from crownto nose darker, blackish, with clear black spot each side ofnose; a white streak over eye; ears blackish out- side, withedge ofgrayand lining of white hairs. Lipsand
throatalmost purewhite, shading through creamy whitetofulvouson breast. Insides of legs grayish,mixed brown and
white hairs; fur of foot pads darkbrown;
feet like legs and thinks. Tailmuch
darker than backwith a sharply defined stripe of darkbrown
along the entire length of upper surfacefromrump
totheshort black tip.Skull
and
teeth.—
Skull large, massive, with the large, broad nasalsand
high saggital crest as in F. coryi; but with larger audital bullae.Much
largerthan theskulls ofF. couguar, with welldeveloped crestand much
largernasals; bullaemuch
larger; opening of anterior nares verymuch
larger; teeth, especially the secondand
third upper premolars, larger.Measarements.
—
Skinoftype: Totallength, 2100 nun.; tail, 735(meas- urements from tanned skin). Skulls of typeand
anadult male, very slightlyyounger, fromMer
Rouge, Louisiana; the latter in parentheses:Condylobasal length, 193.5 (192); basal length, 180 (178); zygomatic breadth, 149 (
—
); palatal length, 87 (88); postpalatal length, 95.5 (97); least interorbital breadth, 43.5 (4ii); greatest length of nasals,63 (60);greatest breadth of nasals, 36.3 (35.4); length of upper premolar row, 46.1 (46); length of audital bulla, 36 (37); length of lower molar-pre- molarseries, 45.7 (46).
Remarks.
— The
rich ferruginous or intense rusty red backand
other color characters ofFelis coryi,and
the pale uniformlycolored backand
tail of Felis oregonensis azteca, are
enough
to distinguish readily the Floridaand
Texas forms from theLouisianapuma.
Pumas
are still fairlycommon
in the wilder parts of the cane brake regionofeastern Louisiana. Inthe early springof1904 whileMr.Waldo
E.ForbesofBoston, Mr. B. V.Lilly ofLouisiana,
and
Iwere huntinginthe BearLake Cane we
heardpumas
calling inthe forestsat night,and
sev- eral times succeeded in starting the animalswith the hounds, but they wereinvariably too fast forour slow bear dogsand
always quicklyleftHolliftii
—
77(6' Louisiana Pinna. 177 them in thedistance.The
following notes aretaken frommy
field note-bookof this trip:
February 23 < 1904).
Heard
panthers crying about nine o'clock last night. There were probably two ofthem
as the callswere sounded at short intervals,some
times only aboutaminuteapart,and
oneseemedalittlefartheraway.
The
animals wereevidentlymoving
alongtothe north.The
cryisalongdrawn
out,shrill trill,weirdand
startling. Itcommences
low onthescale, gradually ascends, increasinginvolume,and
then lowers at theend. ForbesandLillyput thehoundsafterthem
earlythismorning, but withoutsuccess.Heard
pantheragaininevening. February24.Made
a drive for the panther again this
morning
but did not runhim
hardenough
totree. February26.Heard
panthers to-nighton bothsides of BearLake. February27. Crossed the lake earlythismorning and made
a drive forthepanther but the dogs seemed to lose trail at lake. Lilly says that panthers takereadily towater
and
be thinksthis onecrossedto oursidebefore morning.Made
a drive on our sideand
soon started a pantherin the cane.The
animal promptlyoutdistanced the dogsafter badly slittinganearforone ofthem. February28.Heard
the panther's trilling wail across the lake to-night.We
went out in the boatand
silentlypaddled quitenear
him
inaneffortto"shine"
his eyes,butfailed to get ashot.I believe the