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Vol.

XXIV,

pp. 175-178 June 16, 1911

PROCEEDINGS

OFTHE

BIOLOGICAL SOCIETY OF WASHINGTON

THE LOUISIANA PUMA.

BY

N.

HOLLISTER.

[PublishedbypermissionoftheSecretaryoftheSmithsonianInstitution.]

An attempt

to identify the

specimens

of

pumas

in the collec-

tionof the

United

States

National Museum has

resulted in the discovery that the

form found

in

Louisiana

is

very

different

from any

yet described.

This new form

is represented in the

museum

collections

by

three

specimens; two

skulls

from Mer Rouge, Morehouse

Parish, collected

by

J. Fairie

many

years ago,

and a

skin

and

skull in the Biological

Survey

collection

from

Vidalia,

Concordia

Parish.

The Louisiana puma

is

much more

closely relatedto the

puma

of

Florida than

tothe

northern form

or to Felis oregonensisazteca. It isreadily separable

from both

thefirst

and

the last species

by very marked

color differ- ences.

The museum

is fortunate in possessing

two

skulls

each

of Felis

couguar from New York and

Felis corgi

from

Florida, as well as

good

seriesof the

western

species.

The

pelage of

the Florida puma has been

carefully described

by Mr. Chas.

B.

Cory,* by Mr. Outram Bangs,! and by Dr.

C.

Hart Merriam,+

and Mr. Bangs has proved

that it varieslittle in color

with

the season.

From

the recordsofthe

occurrence

of the

puma

in the

Southern

States collected

and published by Dr.

F.

W. True

§ it

seems

certain that the distribution

between Florida and

Louis-

iana was

at

one time continuous. As

so little is

known

of the

animal formerly inhabiting

that region,

however, and

as the

Louisiana puma

differs sogreatly

from

the

Florida animal

in

color it

seems

best to regard itat present as adistinct species.

*Hunting andFishingin Florida,p.109, 1896.

tProc. Biol. Soc.Washington,XIII, pp.15-17, 1899.

tProc.WashingtonAcad.Sci., Ill,pp.583-585, 1901.

§ReportU.S.Nat.Mus.,1888-'89,pp. 591-608,1891.

31—Pkoc.P.iol.Soc.Wash.,Vor..XXIV,1911. (17.-,)

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176

Hollister

— The Louisiana Puma.

Felis

arundivaga

sp. nov.

Type from 12milessouthwestofVidalia, ConcordiaParish, Louisiana.

No. 137,122, U. S. National

Museum

(BiologicalSurveyCollection), skin

and

skull of

d\

adult(occipito-sphenoid sutureentirely closed); collected

June

17, 1905, by B. V. Lilly.

Generalcharacters.

— A

large

puma

of theFelis coryi type but lacking the bright ferruginous color; skull

much

larger than in Feliscouguar, with the verybroadnasalsand highlydevelopedsaggital crest of F. coryi.

Differs externally from Feliso. azteca in its darker coloration and very pronouncedcaudal stripe.

Colorof type.

Upperparts, including outersides oflimbs, grayishfawn color with a decided cast of ecru drab, especially on flanks

and

legs;

nape, withers,

and

an indefinite stripe

down

backto

rump

brighter, pale rufous; the entire upperparts finely

mixed

with dusky, the darkercolor of the hair tips. Face from crownto nose darker, blackish, with clear black spot each side ofnose; a white streak over eye; ears blackish out- side, withedge ofgrayand lining of white hairs. Lips

and

throatalmost purewhite, shading through creamy whitetofulvouson breast. Insides of legs grayish,

mixed brown and

white hairs; fur of foot pads dark

brown;

feet like legs and thinks. Tail

much

darker than backwith a sharply defined stripe of dark

brown

along the entire length of upper surfacefrom

rump

totheshort black tip.

Skull

and

teeth.

Skull large, massive, with the large, broad nasals

and

high saggital crest as in F. coryi; but with larger audital bullae.

Much

largerthan theskulls ofF. couguar, with welldeveloped crest

and much

largernasals; bullae

much

larger; opening of anterior nares very

much

larger; teeth, especially the second

and

third upper premolars, larger.

Measarements.

Skinoftype: Totallength, 2100 nun.; tail, 735(meas- urements from tanned skin). Skulls of type

and

anadult male, very slightlyyounger, from

Mer

Rouge, Louisiana; the latter in parentheses:

Condylobasal length, 193.5 (192); basal length, 180 (178); zygomatic breadth, 149 (

); palatal length, 87 (88); postpalatal length, 95.5 (97); least interorbital breadth, 43.5 (4ii); greatest length of nasals,63 (60);

greatest breadth of nasals, 36.3 (35.4); length of upper premolar row, 46.1 (46); length of audital bulla, 36 (37); length of lower molar-pre- molarseries, 45.7 (46).

Remarks.

— The

rich ferruginous or intense rusty red back

and

other color characters ofFelis coryi,

and

the pale uniformlycolored back

and

tail of Felis oregonensis azteca, are

enough

to distinguish readily the Florida

and

Texas forms from theLouisiana

puma.

Pumas

are still fairly

common

in the wilder parts of the cane brake regionofeastern Louisiana. Inthe early springof1904 whileMr.

Waldo

E.

ForbesofBoston, Mr. B. V.Lilly ofLouisiana,

and

Iwere huntinginthe Bear

Lake Cane we

heard

pumas

calling inthe forestsat night,

and

sev- eral times succeeded in starting the animalswith the hounds, but they wereinvariably too fast forour slow bear dogs

and

always quicklyleft

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Holliftii

77(6' Louisiana Pinna. 177 them in thedistance.

The

following notes aretaken from

my

field note-

bookof this trip:

February 23 < 1904).

Heard

panthers crying about nine o'clock last night. There were probably two of

them

as the callswere sounded at short intervals,

some

times only aboutaminuteapart,

and

oneseemeda

littlefartheraway.

The

animals wereevidently

moving

alongtothe north.

The

cryisalong

drawn

out,shrill trill,weird

and

startling. It

commences

low onthescale, gradually ascends, increasinginvolume,

and

then lowers at theend. ForbesandLillyput thehoundsafter

them

earlythismorning, but withoutsuccess.

Heard

pantheragaininevening. February24.

Made

a drive for the panther again this

morning

but did not run

him

hard

enough

totree. February26.

Heard

panthers to-nighton bothsides of BearLake. February27. Crossed the lake earlythis

morning and made

a drive forthepanther but the dogs seemed to lose trail at lake. Lilly says that panthers takereadily towater

and

be thinksthis onecrossedto oursidebefore morning.

Made

a drive on our side

and

soon started a pantherin the cane.

The

animal promptlyoutdistanced the dogsafter badly slittinganearforone ofthem. February28.

Heard

the panther's trilling wail across the lake to-night.

We

went out in the boat

and

silentlypaddled quitenear

him

inaneffortto

"shine"

his eyes,butfailed to get ashot.

I believe the

pumas

call

more

in the early spring thanatanyother season,

and we

were probablyvery fortunate in the time of ourvisit to the cane,as the experience

was

one of very great interest. Mr. Lilly's efforts toobtain agood specimenforthe BiologicalSurveyresulted inthe captureofthe type the followingyear.

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