, ) (
: *
:
E-mail: [email protected]
رﻮﺷ ﺶﻨﺗ ﺮﺛا ي
ﺎﻫﺪﻴﺳا ﻢﺴﻴﻟﻮﺑﺎﺘﻣ ﺮﺑ ي
،ﻚﻴﺌﻠﻛﻮﻧ
ﺖﻴﻟﺎﻌﻓ ﻢﻳﺰﻧآ
ﺎﻫ ي ﻲﺘﻧآ ناﺪﻴﺴﻛا ﻞﻴﻓوﺮﻠﻛ ﺲﻧﺎﺳرﻮﻠﻓ ،
ﻨﺗ و ﻈ ﻢﻴ هﺪﻨﻨﻛ ﺎﻫ ي ﺰﻤﺳا ي اﺰﻠﻛ ﻢﻗر ﺞﻨﭘ
ﻔﻄﺼﻣ ﻲ رﺪﻴﺣ
1ي ﺮﺼﻣ ﻪﻤﻃﺎﻓ ،*
2ي ﺎﺨﻴﻛ ﺐﻨﻳز و 3
!
) " # :
$ /
% /
&&
' (*+# " # :
/ /
&&
(
هﺪﻴﻜﭼ
! "I
# "$
#
%
! &'
) (CAT
* +, "
) (APX
* -.
) (GPX
' / 0
# 1
# 2 3 /0 45* -
6 02' ! 3 73
# 8 " 1 9*
+:
%3
;< =51
>?
3@' 3 A ,'
# /B8 )
% (
A3. CD .
1 9*
C
#
Hyola308
"
Hyola401
"
Hyola60
"
Option50
RGS003
%3 !F 6 GH IJ A
# K "
LKK "
MKK NKK
&
$0
NaCl
%3 !F 6 %3. B
. D' :O 9%
# / 6 !8
#
PQ' F
# A5F3 !
I
# $
A3 R S
. I 3 = A6 .
4'& =' 4 !
6 0' GH
# 1
RGS003
B C1 A
<
F
# '&
. 6
#
1 3 $8T ' ! D
# U 45* -2
6 C1 8
. 1
Hyola401
=' 4 4%8
U .
3 S
0' NKK
&
6 A $0
P !8 F
# A6 3 73 ! .
A5F3 !
APX
U CAT
0' 8 1
# 6
# 3 F
# 5 !8
A6 GPX
. A5F3 ! 4%8 "C1 4 U CAT
1 8
Option50
1 APX RGS003
<
F
# .
4 9% V
!' ! 1
RGS003
<
6 !6 R
#
A5F3 ! & ,
I
#
%
! /W%
'&
# T C1 A .
هژاو ﺎﻫ ي ﺪﻴﻠﻛ ي :
# "$
#
% ' "!
/
# 0
#
"
"
6
# .
ﻪﻣﺪﻘﻣ
!"# $ %&'( )* +,# -,
'.
#,- /'&'0 -,
$ 123 45' 67.
'" 8. ' 123 8"
$ . %&'( 8. !;
,!< $
/'&'0 )=.'>
(Munns, 1993)
. / )(
?@ AB +'>C
3!. D"E F82
'"& '* -,
Na+
$
Cl-
G $
,HI !J'# 'K L#
,& %7.
8E,$ 8.
!.
,HI !J'# AHM /3 )&
%&'( )* +,# -,
'" 8. ?,! , /'&'0 $
)
Marschner, 1995
.(
= P'. %&'(
,'( / )( F
>E>Q $ G
8=$, R,!S, -, T!. !0 /,G !<
DE, /'&'0 )
Munns, 2000
( F U,G0
'& % !V '"&
@ , = WXE
'*=', ,(, % YE R,!S, %&'( D*M ,-B
5 % -$!.
ZC )>E 1 -, /'&'0 F
#'Q
>B /[(, ) (ROS
(, !CE '"&
)
(O2
(, !C /\$&
(H2O2)
', $ '&
+($&
) (OH
E /$
. R'X(! ,
'- R'3
.!] /E,(, 4!5 -, , F'&
$!C '&
'&E, $ ,$ E 8. 1(
(
(Appel & Hirt, 2004)
.
'>E, /,(,
( . -,
>B /,(, !0 '&
,, 8( )>E ,
== /-$
"(
>&
/, -^''( 8.
) (CAT
-,(,!C R'.E, F )
(APX
-,(,!C (''0 $ )
(GPX
!( ', .
, )GB '&
"* '. %_
', /!( 'KQ!I '&
,-B
0 8. 8>. F, /'&'0 E /[(,
&'0 8 $
!< /'&'0 '*B D='KQ /,G % R
(
)
Alscher et al., 1997
.(
'.
#,- /'&'0 -,
L0 '"&
(Sairam
et al., 2002)
8XC $
(Gosset et al., 1994)
>Q ^'.
, D='KQ /,G )GB
'&
5 % -$!.
DE, U,G0 .
>Q ^'.
D='KQ /,G )GB
'&
>B /,(, )' '* % D@ /'&'0
8. +"@
)' , 8. D
,
$ =$!C '"& ('0-'E R'X(! '(
R,&.!(
,GQ'. /'&'0 +"@ /,G !. '&
. Z"&
D='KQ /,G -, ?,! , 8>E'(DNA
$
, D='KQ /,G >Q ^'. WXE 8. !, )GB
'&
($- '"&
-'(X
'.
. DE, U,G0
% ?,!
` , DaI )GB
'&
/$
,'3 /'&'0 E
Chenopodiaceae
^'.
$
(ABO-Kassem, 2007)
.
. R'K='
8b , /'2 '"
/'&'0 D@
% -, ,K
=B R'X(!
) @ '&
b'0-'E (
c";
.' F 3, R'Xb! ,
!Q B '" '&
'*B
" ,$
b ., , - R'X(!
/, 8.
#,, -, R,&.!b '&
@ ) F >'
-Q, F-'b'E
'b'EV=, $ (
R'Xb! $ 8\$!>
) E, 0 $ =$!C F8B
d'>.
(
!b ', .
)a X*. "* %_ b'0-'E R'Xb!
G"E, % D@ /'&'0 ,
(Good &
Zaplachiniski, 1994)
.
#,- /'&'0 !e(,
E!. ,G( 8"M -, '&
K>
% 8. %(,$
,!< $ 8(
E R'X(! /,G M. '*B ('0-'
B
.3 8. -& ', DE, 8>Q!0 RJ
/,G . 8.,
D='KQ ,_ '. R'X(! , )GB
'&
>B /,(,
'"&
APX
$ GPX
LK $ D f72 CAT
5'X, 8] 8( D
G"E, )a )' $ .
$
D='KQ )GB '&
>B /,(, ,!< '.
/,G 8(
D='KQ )GB '&
D3'E !0 M.DNA
B
8. D$'_ '*, 8V] $
', %_ ,G(
( . E!. %'-B , -, g& ,H=
!Sh iE
j>7
D='KQ /,G !.
)GB '&
FCAT APX
$ kC +Q$!( l'EQ $ +Q$!( !'_ FGPX
m ,G( ) ) 2( n'_ !e(, D'( +.'m k, L'm, (
$
$ /,G '. '*B 8., G )a
( G"E, '&m
'&E, +( !'_ G $ =$!C F @
$ 1(
($o )GB D='KQ
-'(X DE, .I
.
شور و داﻮﻣ
ﺎﻫ
%'-B , RJ 8.
i!5 W='m $ +>('Q
'(
pq Q' 'V2'-B 1m', ,! 8E '.
!"
# :
$% &' ()*+ !, -.%/ 0 / 123 4 ...
$'Q D-
)
!>E.
( 'E +.,- 'V2, rst
,!M,
!0 . j>7 iE
uE '*] +'
&' )
!J
=(
S0
F
=
S1
F
=
S2
$ w
=
S3
^ 1"
%'-B , ,G( )m kC $ NaCl
E!.
>Q!0 ,!m .
G( L'm, +' ,
Hyola308
F
Hyola401
F
Hyola60
F
Option50
$
RGS003
. .
F%!>C ,G( L'm, $H. /!( , 8,M -, K.
8Z&'0 '&
+3, 8.
/,0 '&
$' ,HI @
, '_>, 1m', 0&
. -, K. -$
!0,!m ,HI ?@ /'&'0
j>7 iE F
'.
1" -, '>E, /!( 8Q'x, '. $ 8* NaCl
; /,G 8. '*B
uE !& J )
'0'E D*M
/'&'0 / (
'" +( 8>& 1 -, K. F
8.
!0 '"#, /'&'0 .
@ 8>& !& %'-B 5
,I RJ 8.
yK +'(
!0 . -, lC w -$
M /'&'0 % '"#, FD,!. /,0 uE !&
) $ DaI F/'&'0 ! /-$
G"E, (
'&m
)GB D='KQ /,! ,!"& 8. =$!C $ @ '&
FAPX
$ GPX
,& %7. $ CAT
'. '*B 82 $
!>>Q$!>zE, 'V>E -, '>E, -,,
!0 .
$ !'_
)a ( G"E, '&m $ @
'& T!. !>,M =$!C -,,
!0 . '&m
E, U$ P'E,!. $ ', -, '>E, '. @ 1=E )
, 1956 Schlegel
( -, '>E, '. =$!C $
U$
Bates et al.
(1973
-' $ GXE DQ'. -, T!.
'&
$ {,!7>E, -,,
!0 . +Q$!( /,G . ,
, '. T!.
'V>E -, '>E +Q$!( l'EQ $ SPAD
P!>E, 'V>E '.
>E !>
(Plant stress meter
Biomontor AB)
DQ!0 L';, .
I
'&'" '"#, -, K.
D,!. -, lC $
'&E, +( /,G F/'&'0
DQ'. M 1(
GXE T!.
'&
U$ '.
Chaykin (1970)
-, '>E, '. $
!>>Q$!>zE, 'V>E
-,,
!0 . D='KQ /,G
($- )GB
-'(X U$ P'E,!.I
Kunitz (1950)
GXE DQ'.
T!.
'&
-,,
!0
!0 . ,!.
a ,
T!.
'&
j>7 iE -, /'&'0 D,!. -'
8.
R ,!m !_ '.$ !_ AB 8_m
, .
/,G / ! DQ'. -, L!0 T!.
'&
,!>.E +$!C P'E,!. 8(
Kuntiz (1950)
8*
/" 8Q'x, $ /!( /" \!>'E -, K. F.
/ E E ,, 8( ,!>.E !Q'. @ -,
pH=
- )GB D='KQ /,G . ($
-'(X 5 I
{ } $ Dd,!m !>' -,,
!0 .
! "#$
%
&
'(
!Q'. 8*
Ice–Cold Extraction
: +' @ ,
R'Q )E'>C !Q'. @
^ '.
= pH
$
@
EDTA 0.1 mM
);
~cc
. .
!.
, @
1" $ -, R'Q )E'>C
KH2PO4
$
K2HPO4
'>E, .
@ ,>., @ 8* D*M
^
, -, L,( !& -, 1"
'&
lzE 8*
E E '*B -,
); 8. $ n7 )& '. FD,!.
E E 'E
. @ ,pH
!0 )a .
'( )*+
EDTA
, : ); @
E E DaI '. $
/ 8>3'E ^ .
,!.
8*
!Q'.
Ice–Cold Extraction
F } !Q'. -, !>=$!
,!"& 8. R'Q )E'>C
!>=$!
D,!.EDTA
); 8. $
~cc
'E .
D*M -,,
!0 )GB '&
F
/ '. $ D,!. T!. GXE DQ'. -, L!0
~ E E . !Q'
Ice – cold extraction
!E /$'&
pq'(
F'E 8.
RJ $B V"&
. HI'( -, V"& n7
Q'J R 8. $ X#
$ '. 8_m }
\!>'E .
^'. -'Q lzE
'# /,# 8.
$!C ,!.
"GB D='KQ %;E
'>E,
!0 . R'"# , 8"&
&
' '
~ 8M
>'E ,!0
DQ!0 L';, .
D*M D'*
-,,
!0 )GB D='KQ
-^''(
) U$ -, (CAT Beers & Sizer (1952)
)GB F
R'.EB -,(,!C
) (APX
U$ -,
Nakano & Asada
(1981)
-,(,!C (''0 $ )
(GPX
U$ -,
Urbanek
et al.
(1991)
'>E, .
'*
D , '&
'>E, '. %'-B , -, +J'
L! -, ,GQ, V' 8'_ $ 8G; SAS
P'E,!. '&
8, ] /-B ,
uE ,
% DQ!HC L';, .
,!.
8'!. -, $,M $ '&," )E '&
EXCEL
$
!0 '>E,WORD
.
ﺚﺤﺑ و ﺞﻳﺎﺘﻧ
&
$
G; k'>
'B 8 , '&
$M r
/'2 &
R$' K , !S, $ j>7 iE FL'm, .
D='KQ !. $ , +.'_>
)GB '&
>B /,(, -^''(
) (CAT
-,(,!C R'.E, F )
(APX
$ (''0
-,(,!C )
(GPX
>, M$
. /,G >Q ^'. '.
' s
^ )GB $ D='KQ /,G !.
$ APX
82 $ ,& %7. $ !& CAT
5 8.
K , )GB D='KQ -, ', F$GQ, $ , GPX
%7.
5 8.
K , 8>E'(
. ,!. %,GQ, ,
$APX
uE CAT
s
^ 8. DX
!.,!. 82 W! 8. &'
/ t $
/
J $
,& %7.
'K W! 8.
/
$
/
. J
) $M
.(
)GB $ '* D='KQ /,G >Q ^'.
$APX CAT
82 $ ,& %7. $ 8Z&'0
'&
,G(
/'2
&
!Sh )GB 8E . -, )GB $ ,
>B /,(, !.
8. +"@ /,G
/'&'0 ,
'.
(Appel
& Hirt, 2004)
. 2'-B
Alvesda Costa et al.
(2005)
% !S, !( Lq#,
D='KQ /,G !.
)GB $ $APX
L'm, 82 $ ,& %7. $ GPX
)GB D='KQ -, $ $GQ, L0E j>7 8>E'( CAT
. +.'_> R,!S, 8. n.! k'>
, /'2 L'm, $
5 % '"#,
)m F
Optlon50
uE
S3
) w
^ (NaCl
,, !>2.
)GB D='KQ /,G ,& %7. )&APX
)& $
82 %7.
'.
$ !& )GB , D='KQ /,G $
'K &' '" 8. DX %7.
w / t~
DE, J
) + '&
$ .(
-, . , )GB D='KQ /,G
>Q ^'. '. ,!"& 82 $ ,& %7. $ !& GPX
uE
' w
^ '"
,G( L'm,
8>E'(
. '" %&'( ,
!0 &'2 L'm,
2&'( $ ',
)m )GB ,
Hyola401
$ !&
m, !'E -, !>"( W,! 8. 82 $ ,& %7.
. L' )
+ '&
w $
~ .(
)GB + 8( 8ZB P'E,!. FCAT
'&
$ } &'2 5 F % '"#,
!.
'" G )GB , D='KQ /,G
!& $ ,G( L'm,
/,G !^'. ', F $GQ, 82 $ ,& %7. $ uE )GB ,
w
^ !& NaCl
)m 8. n.! %7. $
RGS003
. . %,GQ, /,G ,
,!. &' '" 8. DX
'K 82
/ $ J
,!.
!.,!. ,& %7.
/ . J .
+
L'm, 8(
Hyola401
$
Hyola60
)GB %,GQ, uE ' '*CAT
^
^'. '. F.
uE >Q
%7. /B ,_ -,
8>E'( L'm, , ,&
. !a P'E,!.
Mckersie &
Leshem (1994)
)GB FL$G(,!C CAT
(> $ DEqC$!( F)EqC>E
$ , M$
+X WXE
H2O2
8.
H2O
$
O2
. Z"&
)GB /$!>=, & /,# 8. R'.$!E, -, APX
,>., /'0 +E
'>E, R'.E,
( $
"* %_
,- D"E
H2O2
, E .
R'__@ P'E,!.
Shen et al.
(1997)
F WXE
+X
O2-
8.
H2O2
c' !, , F E /$
-'Em B!Q D'* $ ='( 83!] D='KQ
/'&'0
. ^'. ,H=
)GB D='KQ >Q '&
>B /,(, '"&
CAT
, YE R,!S, -,
+2
H2O2
-'Em B!Q !.
DEqC$!(
!0M ( .
8. n.! k'> P'E,!.
)GB '&
>B /,(,
/, 5 ,G( j>7 L'm, 8( DQ!0 8;>
% '. / 8M,
D='KQ /,G !. '*
$
)GB $APX
)GB D='KQ -, $ $GQ, CAT GPX
8>E'( '*B 82 $ ,& %7. $ !&
. ,!.'.
8. +"@
)GB $ '* ,G(
>B /,(,
$APX
!S CAT
. %'-B , k'> 8.'2
Tohidi-
Moghadam et al.
(2009)
5 8( !( U,G0
23 % -$!.
)GB D='KQ /,G -, G
GPX
,_ !. $ 8>E'( ,G(
$GQ,SOD
.
Appel &
Hirt (2004)
&'0 80 8. 8>. 8( !( Lq#,
` $
)GB F%
'&
>B /,(, J'3
'KQ +"@ D*M
!"
# :
$% &' ()*+ !, -.%/ 0 / 123 4 ...
6
. ,!.
'e
Heidari & Mesri (2008)
U,G0
@ L0 j>7 L'm, 8( !(
% D
/,G !.
)GB 8E !& D='KQ FCAT
$ APX
$GQ, GPX
.
0 0.02 0.04 0.06 0.08 0.1 0.12 0.14
0 100 200 300
(NaCl ر ) ر APX (µ mol H2O2 g-1 FW)
Hyola308 Hyola401 Hyola60 Optlon500 RGS003
اه
1@/ APX -9+; $% &' / -< =>, 7/ ?0 4 8 7#
9*
0 0.02 0.04 0.06 0.08 0.1
0 100 200 300
(NaCl ر ) ر
APX (µ mol H2O2 g-1 FW)
Hyola308 Hyola401 Hyola60 Optlon50 RGS003
ر
9*A APX -9+; $% &' / -< =>, 7/ ?0 4 8 7#
0 0.3 0.6 0.9 1.2 1.5 1.8
0 100 200 300
(NaCl ر ) ر GPX (µ mol H2O2 g-1 FW))
Hyola308 Hyola401 Hyola60 Optlin50 RGS003 اه
9* 1@/ GPX -9+; $% &' / -< =>, 7/ ?0 4 8 7#
B
0 0.5 1 1.5 2
0 100 200 300
(NaCl ر ) ر GPX (µ mol H2O2 g-1 FW)
Hyola308 Hyola401 Hyola60 Optlon50 RGS003
ر
7#
8 -9+; $% &' / -< =>, 7/ ?0 4 A GPX
9*
0 0.01 0.02 0.03 0.04 0.05 0.06
0 100 200 300
(NaCl ر ) ر CAT (µmol H202 g -1 FW)
Hyola308 Hyola401 Hyola60 Optin50 RGS003
اه
9* 1@/ CAT -9+; $% &' / -< =>, 7/ ?0 4 86 7#
0 0.02 0.04 0.06 0.08 0.1
0 100 200 300
(NaCl ر ) ر CAT (µ mol H2O2 g-1 FW)
Hyola308 Hyola401 Hyola60 Optin50 RGS003
` ر
9*A CAT -9+; $% &' / -< =>, 7/ ?0 4 8B 7#
$M
&'2
!Sh K ,
$ c"; /,G !.
)a ( G"E, '&m @
,G( 82 G $ ,& %7. GXE DQ'. =$!C $ ,
. /,G >Q ^'. '.
' w
^ !.
82 $ ,& %7. $ !& =$!C DaI ', $GQ,
,& %7. R,&.!( /,G -,
,_ !. $ 8>E'(
$!Q, 82 '* /B )
$M .(
!"
# :
$% &' ()*+ !, -.%/ 0 / 123 4 ...
C
D!E 8 -9+; ()*+ !, -.%/ 0 F+ 93 0+;
-G23 !.*
!22*
7'* F+ ,' 9, 9*
! "#$
! "#$
%&'
APX
APX
GPX
GPX
CAT
CAT
F0
Fv
Fv/Fm
DNaseI
(') "* %'
!"ns
/
!!#**
/
$**
/
!$**
/
%"**
/
!
&&**
/ **
/ '"*
/
#ns
/
"**
/
&**
/ '**
/
"#!'
!ns
/ '($&
!ns
/
!**
/
!&
!**
/
!"#' (**
/
!'
) ' '&**
/
#**
/
&
!(**
/
&
&!**
/ (%**
/
&(**
/
%**
/
&(*
/ **
/
!#**
/
!'**
/
"*
/
&##
ns
/ ''!(
&ns
/
&*
/ ' '*
/ '""&
"**
/ '
)
*
!&
!(ns
/
"**
/ (**
/
#**
/
#**
/
*
/
!**
/
!&ns
/
&ns
/
"**
/
!**
/
%**
/
"!$"
$/ns
"!'&
!ns
/
&**
/
"
**
/
&#'%"
'**
/
&
+, '#
/ (#
/
"
/
&
/
# /
!"
/
&
/
!%
/ ' / /
&
/
# /
%%' ( /
&%#'#
% / ( /
"
/
!!&&#
$#
/
' %CV
/$
/$
&
/
# '(
/
#
&
/ ( /
!"
' /
!"
!&
/
!#
!' /
!$
$ /
&
% /
!#
( /
!!
$/
!!
# / '
# /
!'
# /
% ' /
!#
ns
*
**
.
D!E 8 HI+ . ? )*+ !, -.%/ 0 / J/ ' K L4
-9+; ( 0+;
-G23 !.*
!22*
7'* F+ ,' 9, 9*
- . 0, 123
40 123
567 8 89 :;<
40 89
:;<
567 8
APX
40
APX
567 8
GPX
40
GPX
567 8 CAT
40 CAT
567 8
DNaseI
)
>?
( )
( )
( (µmol H2O2 min-1 mg-1 prot)
F0 Fv Fm/Fv
AB
SPAD Unit/ml g/mgFwt
C
Hyola308 '"a
/ (a
/
!
!b
/
!
!a
/
!d
/ '&b
/
!b
/
%%b
/
&ab
/
! 'b
/
!#d
/
$/bc
&""
#a
/
!#
(!a
/
"&b
/$
$/a
!(#' '#a
/
"
Hyola401 '!a
/
"&a
/
!
"b
/
!
#a
/
&&a
/
!
!&c
/
&'c
/
&(ab
/
!
!$ab
/
!
&c
/ 'a
/
"c
/
&$
#a
/
!&(
(&a
/ '&b
/$
b
/
!%&
#c
/
Hyola60 'a
/
#a
/
!
!!a
/
!
&!b
/
&&d
/
!"c
/
&!c
/
&'ab
/
!
!%ab
/
!
&!cd
/ 'b
/
!a
/ ' 'a
/
! (!a
/
&a
/ ( 'b
/
!&"#
(!c
/ '
Option50 '"a
/
&#b
/
!
%&c
/
#a
/
%"b
/
a
/
$$a
/
!b
/
!
!!b
/
!
!#d
/
!d
/
$/c
&$
"a
/
!"!(
(&a
/
"!c
/
"
(b
/
!'$'
#%c
/ '
RGS003
&"a
/
"a
/
!
%c
/
&#b
/
$$c
/
!$b
/
"#a
/ a
/
! '&a
/
!
$(a
/
&&c
/ ab
/
&##
&a
/
!""
#%a
/
%!b
/$
&a
/
&#"
&%a
/$
) )
<
D<
(NaCl
8)
"a
/
%&d
/ 'c
/
&#c
/ (&a
/
#c
/
!$d
/
!a
/
!
&a
/
!
!%c
/
!'c
/ a
/
&(&
$/a
!' (&a
/ '"a
/
#
"b
/
!$%
"%a
/
"
! '"b
/
"!c
/
!
!!b
/
!
&(c
/ (a
/
&!b
/
&(c
/
!%ab
/
!
!(a
/
! '"b
/
&"b
/
!ab
/
&$$
$/a
!#$
(!ab
/
$$ab
/$
"b
/
!(%
#$b
/
&
&'bc
/
"%b
/
!
!"b
/
! b
/
#(a
/
&#b
/
"b
/
&!ab
/
!
&b
/
!
!a
/ '$a
/
"ab
/
&$%
"a
/
!"' (!ab
/
$$ab
/$
$/a
!#"!
('a
/
"
'
!$c
/
#%a
/
!
&%a
/
!
"&a
/ '&b
/
!a
/
#&a
/
!$b
/
!
"c
/
! 'a
/
&(b
/ b
/
&%
$/a
!#$
(b
/
!b
/$
!a
/
!#'%
#%a
/
"
! "#$
#%&
' (#
.
82 W(! $ , %,GQ, /'2
&
'>E,
,!. '*B -, G"E, )
'.
. -, j>7 __@
8"M
Martin et al.
(1993)
$
Heidari & Mesri
(2008)
$ L0 =$!C /,G %,GQ, -,
Sultana et
al.
(1999)
% D@ k!.
!X3
&
.
Cavelierl (1983)
/'&'0 =$!C %,GQ, !( Lq#,
% D@
# cm,$
8. '0 g!5 -, %(,$
DE, 82 ?@ AB +'>C %&'(
. /'- ,
G"E, +'>C /!( )( '. =$!C
E '&
82
,!. L-^ ?,!
$ AB AHM )&,!Q , ,HI !J'#
( . '&m DaI !0 f72 %'-B ,
uE 82 =$!C $ @
S3
8. DX 2,GQ, -, &'
'K /
~t ', F. ,3!. J
,& %7.
,!. %,GQ, ,
'K =$!C }/
}}
. J )
$M .(
V>X"& %'-B ,
K ,
>Xe $ '&m .
=$!C $ @
. 82 %7.
8 B DE .
', )GB
'&
>B /,(, V>X"& 8( !0 f72
=$!C .
'.
GPX
)GB $ '. '* $ $CAT
%7. APX
82 K , DE, DXe $ )
$M w .(
k'> P'E,!.
l',$ 8G;
, '&
$M
&'2 8(
$ )m +.'_> !S,
'&m W(! $ !&
82 $ 8m'E %7. $ =$!C $ @ K
,
DE, .
!0 ,!m /'-
iE !K L'm, ,
j>7
%7. $ =$!C DaI !^'. F
,&
uE $ 82 $
w
^ n.! F
8.
)m
Hyola401
2,GQ, -, W! 8. 8( .
'K
/ } $ / t . ,3!. &' '" 8. DX J
) + '&
$ t .(
'&m
%7. F @
uE >Q ^'. '. 82
$GQ, /B /,G !.
.
uE !^'.
) w
^ ( )m F
Hyola401
,, !>2.
2,GQ, -, 8( . ,_
'K
/
~}
. ,3!. &' '" 8. DX J
) + .(
,& %7. ',
,_ )m "& $
uE ' '* R,&.!(
^ %,GQ,
uE >Q ^'. '. $ , /'2
' w
^
8>E'( /B /,G -, .
m, !V 8( '"& L'
+
%,GQ, '. ,!"&
-,
'&m /,G
8>E'( @ .
Munns (1993)
!( Lq#,
\
'&
+"@>
D!E 8 M -9+; ()*+ !, -.%/ 0 H/ I0.N O 0+;
-G23 !.*
!22*
7'* F+ ,' 9,
* 9
!
&
'
"
$
# (
%
!
!!
!&
!'
!
)
! ( 0, .
!
)
&
( 40 123
"**
/ E
!
) ' ( 8 567 123
"!**
/ E
%'**
/
!
) ( 40 :;< 89
&%*
/ E '**
/
&*
/
!
)
"
( 8 567 :;< 89
!$ns
/
&!ns
/ E
&#*
/ E
&!ns
/
!
)
$ ( 40 APX
!#ns
/ E '!*
/ '(**
/ '$**
/
!%ns
/ E
!
)
# ( 8 567 APX
$**
/ E
!**
/ '(**
/ '"**
/
!ns
/ E
$!**
/
!
) ( ( 40 GPX
!"ns
/
&"*
/ E '!*
/ E ''**
/ E
!'ns
/
&&*
/ E
&$*
/ E
!
)
% ( 8 567GPX
"'**
/
"**
/ E
""**
/ E '%*
/ E
!&ns
/ ''**
/ '!*
/ E
!'ns
/
!
)
! ( 40 CAT '*
/ E '$**
/
&"*
/
"ns
/ E
%ns
/ E
$ns
/ '#**
/ '"ns
/ E '&ns
/
!
)
!!
( 8 567CAT
&#*
/ E '&*
/ '&*
/ (ns
/
!ns
/
%ns
/ E
&ns
/ E
%ns
/ E
!!ns
/ E
#ns
/
!
)
!&
( Fm/Fv
!%ns
/
&!ns
/
!$ns
/ E
!!ns
/ E
!!ns
/
!#ns
/ E
&"*
/ E
!&ns
/ E '%ns
/ E '%ns
/ E
$ns
/
!
)
!' ( 'ns
/
!%ns
/ E
!ns
/ E
!"ns
/ E '#**
/ E '$**
/
&'*
/
!$ns
/
&!ns
/
&(ns
/ E
$ns
/
#ns
/ E
!
)
! ( DNaseI
#ns
/ E
&"*
/
!"ns
/ (ns
/ E
"ns
/ E
$!ns
/ '"**
/ (ns
/ E
!&ns
/
$'**
/ '#ns
/
!"ns
/ E
$&ns
/ E
!
)
!"
(
&**
/ E
$!**
/
!$ns
/
!!ns
/ E
&$*
/
"'**
/
&'*
/ E
!'ns
/ E
#!**
/
&"*
/
!'ns
/ E
!#ns
/
&(*
/ E
!#ns
/ E
ns
*
**
.
0 0.2 0.4 0.6 0.8 1 1.2 1.4 1.6
0 100 200 300
(NaCl ر ) ر و µmol g FW
Hyola308 Hyola401 Hyola60 Optlon50 RGS003
اه
8 C 7#
1@/ H%P 9 / -< =>, 7/ ?0 4 9*
0 0.5 1 1.5 2 2.5
0 100 200 300
(NaCl ر ) ر و µmol g FW
Hyola308 Hyola401 Hyola60 Optin50 RGS003
ر
9*A H%P 9 / -< =>, 7/ ?0 4 8 7#