• Tidak ada hasil yang ditemukan

( . . . . . . ) ( . / P> . LSD =% α . . N ) CCA ( . ) CCA

N/A
N/A
Protected

Academic year: 2024

Membagikan "( . . . . . . ) ( . / P> . LSD =% α . . N ) CCA ( . ) CCA"

Copied!
11
0
0

Teks penuh

(1)

!"

#

$%&

'() *

%+

&+

%+

!"#

N1

$%&' $( ) $* %+ ,

*

!"# $% & !

!"# $% &

'(

!)

* + !"# $% , -, +.!./ 0 12(

) 45 6 :

78 / 7;

/

<<

= 6

>

:

?8 /

@ /

<8 (

! " #$ $ %& ' ( !)

. * #$ $

+ , -/ *0 1 " #$ 2$ + , 3 4*/* )$5 .

"

/3

%6 7 8*0 )

9 : $ " )

;7<

9 ' ( 4*</ )*

+*

. )*

/3 " 4 -<$=

)* $ ) #*> ?* ) /**

@/ AB2 )*

/ 3 .

$=

#

)*

;7<

/3 " )

/3 " C*0 )& $ $ 9& !D* &% E 86 F*G $ )

+ ) .

H*7/ 4*IJ 4* > 4*: *K @ 0

/%/ $

A*% 'J 4*: ) L

0 ,*;

. 9

/3 +:/ 2*K *3 )*

#* 0 1 !' M$ *I * )*

) OPQR L OPQO ( / ) 3 0 .

# 4*: ) ) $ %*-2 ) !#* / T2 !:0 )*;6 8*0 /$ , U )*

M >

0 $

U #

* VW / V P>

0 H*7/

. 4Y Z 4*</ LSD

[

>

\ ) O

=%

α /3 " @/

_1* )

%/ $ ` )*

$ ` .

# 46 #* / T2 #

;7<

a 0 H*7/ > 4*: *K )

'*Gb * U 4*</

0*

. c%

* d a*

eE * fG C*0 _

N1

b* $ 0 C7< g*

M$ * C*0 ) , 3 $ 9& !D* 86 hi

[*> eG )

(CCA +, 3 8&E $ hi .

* hi

,*> eG )

(CCA Gj $ ?* -G% 3/* e $ D* 2Y J * "

I -G% * * $ + .

:

/3 "

# !) /3 ,$ )*

!) hi ,*> eG

* :

:

: E-mail:[email protected]

(2)

!"

...

! " "

#$% &'

" () "* %" + ,- ,%- .

- "! " /0!

1- 2 2 34

5! % /0!

6!

5!

- * 0 %"

7"

) Soudi, 1993 (.

# : ;< :3= >? %"

! @% #- A+*

6!$

!A

%" A !%"%*

+

! B: "%

6!$

" %"%* + )

! 1'C (

D" B:

6!$

%"%* E >

7 )

!

! E

FE .(

G D' H +

I"J ! @"4 .

&&? % 7/ %"

! C ": %

)

%E7

! ( K! %"

34 AL' G4 ME ' $

*A : D"

) 6!$

! 3!

( "% M!

" %N 4 )

Magurran, 2003 (

.

%K 6%

!

O% G4 %"

P)+

*

C

+ P) "(

"7 .

O%

C P)+

Q*7 57

! 5 #!7 #RE SE N1

...

7 .

O%

P)+

O 57

!$ G

"(

K!

!E! >3 %-

T! G4 %* % %K

"7 ) Mesdaghi, 2002; Magurran, 2003;

Vandermeer, 1981 (

. E4

!$ G

%

) UVWX (

E%D'

) Fisher et al., 1943 (

#"E! %7

H)!

H Y% Z3% H: O[

%E7

! 6!$

" ! .

: -

Motomura's geometric series

Log series

A7 P< SE <

! \' #

...

$ %N "( "%

- B:

]J %"

! !$

" ^$

" " _ 5'" ' !

! C ('

` #G4 %" *A a

2 ;E: 5N # E

"3! AL' G4

"* b* :4 20

) Mesdaghi,

2002; Magurran, 2003 (.

c' ' )

Xdde ( % 6!$

! < % ! \' 5 N7 F4 %"

Hc 6!$

! C %" G4

f

!"E! $: #7 .

%" O[ M! %"

O %

P)+

2 %"

%"

G4

$ @%J .

%0 ""

) Xddg ( %"

! & G4 %

< %

;c A H

P)+

@_ " +!

; $ K! %"

G ])

E%D' #! \' #A7 P< h57

!"" %N % "( "% % .

+! i!

! G4 ""

;

! G4

! \' f 20

. M "" +! ^&? H %- j

_

) #AA' M

` M #;M J%" #

kl m* $ #'- H (

@_ " %"

!

f E &l

!%"

. E )

XddX (

4 ,70 @_ $0 5< no

! "% % F- ( Yp N[ G

6%

"" %N .

H %"

6%

J%" _ ,70

! 4

!

_ <

c4 17 J%" #(% 57 $0 5<

_ <

7 $ K! %" 5&

. i!

- ,M$ (% , "" +! ^&? H

-

Broken stick

Log normal

Oliveria & Batalha

(3)

#$ %&'() * +& %, -"/

0 +12 % '3 % 45 % 67 8 58

5

! ,M$ q< 17

7

"

. 0 H ,a

%

P)+

2 )

!$ G (

!

l? 5< n )

$0 ;N #m*

( Q*7

!

A %c %" G4

:

c(J

""0 .

H %"

6%

%" : A "

R %

)

!"

F!* "7 (

E! : A "

R

) S*- 7"%

( F)! c(J %"

7 . 6a

2 "% ^= ,70 m* #$M$

6%

4 %"

U X 7 "%- .

! % %K

%- # g

,70 % '

A

"% : 6%

) UWrU UWrg (

c(J :3= G %+ @&&? M 7 /*

) Borhani et al., 2010 (

*- 6%

) UWrj (

""

> m* ,70 Z

6+a0 GE4

7 %- .

! ;

A! " %" '

@%"

= %" n

"%

6%

#: A "%-

7 .

! %K E! &l %" ;

R

@%"

U

× U

%" &l %"

R %

@%"

U

× X "$ O%E7 O%

!

"(

7 .

!" #7"% &l %"

F!* "7

&l " 7" 5'"

2 "p UX %A

!E!

l* A! %c 7 F)! %"

edd

J$

Udd A! %" 1P! DA

Ug 5J$ n @%"

Wd : DA 7 3

. %"

S*- &l ,70 @_ 0 5'

#17 ]) @c4 %"

2 "p g %A

!E!

F)! %"

7 A! ,7 Xdd

J$

gd A! %" 7 3: DA Ud

J$ n @%"s Xd

% DA 7 "

.

!E! %K

"? %" %"

"

O% 7

tE u

"( $"P 7

.

; # % 7 *7 O% E4 %"E!

6!$

3%

? u

.

&&? O% H %"

$ G SE !

#

! \' G #'

) QN!

( "( A7 P<

E!

.

? H

#

3! D!

6!$

! 0 % ;%D' v&

%?

3% 5& %" y 6!$

% ; " c!-

%?

x 7 ) 5A7 U .(

:

*A T<

6!$

! #]) 6!$

5 3! ! 3!

"" ,E! J%" %+< @%J 7 3?

"7 . ? >* 5A7

- 2 ]J

" %" *A

% "%" "% G4

) Mesdaghi, 2002; Magurran, 2003 .(

(4)

!" # ...

!

"#$%

"E ' °

"N ' °

"E ' °

"N ' °

"E ' °

"N ' °

"E ' °

"N ' °

&'(

!

#$

!

#$

!%&()

#$ ($*

!%&()

#$ ($*

)*

+","

) (m

+

/ ) (mm

+

++

0"

(%) +

,

0" % - .

/ -

01%2

+","

23 (%)

/

/ +

/

/

4 "5 41#&

56

56

41#&

% 067 8"

Bromus tomentellus- Scariola orientalis Scariola orientalis- Cousinia cylindracea

Artemisia sieberi Artemisia sieberi

&'(

9"

: : ; (

< =9 )

(cm

>?

@

? A?

@ B?

C

@

?

>?

@ C B?

@

>?

>D

@

? DC @

>D E?

@ DC

>B

@

? DE @

>B E?

@ DE

#/

(

( (

(

&1/

&1/

&1/

( &1/

( (

/

pH / / + / +

/ +

/ +

/ + / + / + / + +

/ + / + / +

F8 G

/ / +

+

/

6 G

/ 7 7

/ 7

7

/ 7

7

/ 7

7

(5)

!"

#

$%&

'() *

%+

&+

&$

)( )

Mesdaghi, 2002; Magurran, 2003 (

!"#

$ %

&

' (

&) *+

, -

# . /01 23 1 +4 5+#-

"%

)

& &" 2

0 (

/01 23 1 89 5+#

"%

) :# ;< = >0 2 (

.

="

?0 23 ," *@ 0 & &" &

A

BC ,"# 23 D+-

"#

. A 23 + = >0 ="

BC 1

"#

. :# ;< ="

+

3

: +@+

A

"#

. % ,"# 5E 23 D+0

=" F3 D =G ,"#

/& H+FI +:

) Magurran, 2003 (.

''(

* D

"

="

,"#

J

8 K@ L) ,

@ 0 "#

/& D M 23 NF

" +- 2 ="

.

0 B++O /L3 CP Q D+R S+- T ,& Q+( ;< J

"+

.

%

UVVV

% % L@+- D+:E

W X

, K+Y(

8+& !

/&

) Magurran, 2003 (.

=" Z /L3 /:E

B'$

,

"

D++F ,"# ,

# . 5Y B'$ D

log2

HF

"#

. [-

#

KT

'$ \

- , % ="

&( ,"#

# . D++F + \ D"

#

) (S K% ! )

(N

i) (n

"#

.

)

% * ]

&

) ( S D+ ^

,"

,"#

J \ KT

B'$

,"#

, %

# . R4 *) D

,"# &( P _`

/ _ ,"# b- =" "#

=" Bc D +

# ) Magurran, 2003 (

.

D &

=" +Y%

X ,&

Z Past

"#

"

.

Krebs

(6)

&, -. /

01 20 3 *0 4 5' 607 8

...

A D+ d & \

K<

Q+(

) - 8+YT !e (

'+Q Z

F

CCA

"# ,&

) Mesdaghi, 2002 (

.

D L W ,

=&

Ufgh T ,&

( +Y% /

&

X CANOCO

i

`

"# XW .

D++ :' *)

LSD

jQ&

U

=%

α *@

m9 % F

'Q D+ A \

+ n&

F * D+ D+R n&

2T

+ 'Q

n&

) /# R) (

3

0G 2T F * D+ %

n&

)

# o" * (

m9 F

,"

# )

="3 f i K# ! p .(

="3

` q /&"

,") % Z *)

&

) (

="

>0 !:# ;<

& !=

, &" & 0

:

* $ F

=&

&

) Ufgh ]

UfgU ( *@

"

. >0 =" % *@ *) D

=" D+ ,"# Z =" DL =

,"#

F3 NG + % "#

/:

"-

/& d&

.

K#

f * L4 ,"# Z = >0

F

&

=&

Ufgh *@

"

.

rE@ ' ( &

# D1 %

D% R) F * ^

o" F * sYF

"#

. D+R

rE@ ( 5+# &

D% % #

Canonical Correspondence Analysis

Least significant different test

D@+ o" F * /& , t R) F * .

q

&"

/ ,") % *@ CCA

D+:E (

* i /

`U X ( "c iU

K% B++O "c

+3

"%

. ( /:E " 1 :

) V`

/ _ ( 0) "c ! )

gp / _ ( : X (

5+# 1 )

u / _ ( ) PH hu / _ ( *@

"

.

" v B9- )

# K i ( ,") /&"

( U

p

s$

n&

) /& /&

( + n&

) w4 /&

(

$

"3 T 8

"+

. D+R

K#

,"#

*@

R) F * % "

) /&

D+- w4 (

*+ D1

"#

'Q

&

n

o" *#

) /&

D+- /&

( 5+% + % ,"# 2T 8 J

/:

'Q *:

,"#

% ( ,

*) * x 'Q J 50 .

D+R

K#

,"#

*@

"c !" K< & % "

5+# 0) # /y 1 : "

0G 1 : " % .

(7)

!"

#

$%&

'() *

%+

&+

&

!

"

#$%

&' *

1

) N

"+

"+,

(

! N1

! " #

/

/

/

/

/ /

#!"

$%&

/

/

! - . ./0 1

233 &'

1

) N

"+

"+, (

$%

& ' () *(

+

&

!

&

F P

'(" )"

/

/

*

***

/

*

+

/

/

,-

/

***

) ]]

* ]]

( R) a

) ]]

* ]]

( /#

b

)

]]

* ]]

(

#

* c

) ]]

* ]]

( o"

c

D++

Ug Up

h

_

4 56 7.8

&'

1 9:* ;.< : N

) LSD =a b >?@' A B c C

3 DEF 3

% 3 (

S&

A r#

K+

N1

"c

/y 1 : " *+ 0) *@ 1 :

"

$ % %

A t

"

.

="

F * L4 ,"# Z 2

D@+

E

"# = >0 ="

)

*+

@+ & %P

` %

"#

( {y&

=&

UfgU ! Ufg`

Ufgh

=" J |+ %

F * 5& ,"#

"@ , r+E@ o"

% =G , K+0

=&

,"#

/&

) ="3

` .(

K#

f i *@

R) F * % "

D@+

"#

. K#

f ,"

# 2 %

: * +

" R) F @+ = >0 =

/&

) ( B<

,"#

5+#

%

"#

.(

(8)

&:

-. /

01 20 3 *0 4 5' 607 8

...

! H I.@

JFB 2*

K:B 9:* : )

( 233 B L.:

9 7

C

< 3 F

"+,

"+

,(- .

/0

12 34 ! 50

P P

P P

/

/ /

/

/

/ /

* /

/

/ /

/

/

/

/ /

/

/ /

/ /

*

/

/

* /

/

/ / /

/

*

/

/ /

/

/ /

/ /

*

/

/ /

* /

/ /

/ /

*

/ /

/

6) .

/0 12 34

! 50

P

P P

P

/

/ /

* / /

*

/

/ /

/

/

* /

/ /

/

/

* /

* /

/ /

/ /

/

* /

* /

/

/ /

/ /

/

/

* /

/

/

* /

/ /

/

/ /

/

/

* /

/ /

*

/

/ /

!( / ( .

/0 12 34

! 50

P

P P

P

/

/ /

/ /

/

* / /

/

/ /

/ /

/

/ /

/

*

/ /

*

/ /

*

/

* /

* /

/

/

* /

/ /

/

/ /

/

/ /

/ /

/ /

/

/

/ /

/ /

*

/

/ /

7.!

. /0

12 34 ! 50

P

P P

P

/

/ /

/ /

*

/

/ /

/

*

/ /

*

/ /

/

* / /

/

/

* /

*

/ /

/

/ /

/

/ /

/ /

/

/ /

* /

/ /

/ /

/

/ /

/

/ /

/ /

/

* / /

P

/ ! .

(9)

!"

#

$%&

'() *

%+

&+

&;

0.00 0.50 1.00 1.50 2.00 2.50 3.00

0 20 40 60

ﺎﻫ ﻪﻧﻮﮔ ﺐﻴﻛﺮﺗ

ﺒﺴﻧ رﻮﻓو باﺪﻧﻮﺧﻪﭽﺧآ

ﺖﺷدرو نﺎﻳدﺎﺷ ﻪﻧدﺮﮔ

(log2 P8 B)"+, F 3 C 9 M NO 3 "

- ./0 C@ 8E )

) (CCA

= (

ﺐﻴﺷ ph

ﻚﻫآ ﺪﺻرد ﻲﻟآ داﻮﻣ ﺪﺻرد

هﺰﻳﺮﮕﻨﺳ و ﮓﻨﺳ ﺪﺻرد

ﻲﮔﺪﻧرﺎﺑ

ﺎﻣد

عﻮﻨﺗ ) ﻞﻴﻫ ﺺﺧﺎﺷ N1

(

-1/0 1/0

-

1/0

1/0

Ac.spAc.gr

Ag.int Agr.tri Aj.cha

An.fr

Ar.sz

Ar.si

As.ad As.cam

As.cal As.cep

As.fr

As.mac

As.mol

As.pod As.sil

As.sus As.ver

Bo.cy Br.to

Bu.en Ca.st

Cen.vir Cir.br

Col.sp Cou.ba

Cou.cy Cou.la

Da.gl

Ech.rob

Eph.in

Eup.ch Eup.di

Erem.per

Eren.bu

Eren.bi Eth.sp

Eup.bun Eur.ce

Fe.ov

Fe.ov

Ga.ga Ga.ce

Ga.au

Ger.sp Ger.tub

Gyp.vir

Her.ang

Ix.ta Ju.bu

La.ac No.ma On.me

Orn.te

Pe.ha Ph.an

Ph.ol

pi.au Poa.bo

poa.si

Poli.sp

poly.pa

Psa.fr Pte.ca

Ran.ar

Sal.sy Sc.au

sc.pe Scar.or

Sco.ph

Sco.ra

Se.in Sil.er

Sta.ac

Sta.in St.ba Ta.pi

Tar.ros

Tri.an

Tha.iso

The.ko

Tra.ca Tul.st

باﺪﻧﻮﺧ

نﺎﻳدﺎﺷ ﻪﻧدﺮﮔ' ﺖﺷدرو

ﻪﭽﺧآ

(10)

! "#

...

K+0" D

!@+

D % /& 1 A R) F * K# AI i

K+ A r# &

N1

R) F *

# D /& rE@

X 4 & "

}< % " t

eE# * + %

# + * +

K+0" eE#

~ S o) d# e o

'Q

A AW % "#

/# " = .

K#

f % +

% ( /& o" F * S#

1

) 50 (

,"

"#

. D 1 /+0

# NG K+0 F * )

Artemisia

sieberi (

X"< D 5& : d# !/&

/& ,"# *) +4 }< "# K; 4 .

*Lc Fc ,@ X( * +Y%

%

@% B'+'(

+c *Lc *& F+$ 2

8+ "T @ .

- Borhani, M., Arzani, H., Feizi, M., Saidfar, M., Khodagholi, M.& Ghaedi, A., 2010. Final report range assessment program in different areas of Isfahan province, Isfahan Research Center for Agriculture and Natural Resources.

- Ebrahimi Kebria, KH., Shokri, M.& Tamartash, R., 2002. Effect of topography and grazing factors on vegetation percent and diversity changes for river basin Harraz. MSc thesis, Mazandran University. 54 PP.

- Fisher, R.A., Corbet, A.S.& Williams, C.B., 1943. The relation between the number of species and the number of individuals in a random sample of an animal population. Animal Ecology 12 : 42-58.

- Krebs, C.J., 1999. Ecological methodology, 2ed Edition. Happer & Row, New York, 620p.

- Magurran, A. E., 2003. Measuring biological diversity. Blackwell Publishing, 256p.

- Mesdaghi, M., 2002. Vegetation description and analysis. Mashhad University Publications, 287P.

- Mirdavoodi, H. R.& Zahedi Poor, H., 2005. Determination of suitable species diversity model for Meyghan playa plant association and effect of some ecological factors on diversity change. Pajouhesh & Sazandegi 68: 56-65.

- Motomura, I., 1932. On the statistical treatment of communities (in Japanese). Zool. Tokyo 44: 379-383.

- Oliveira, F.F.& Batalha, M.A., 2005. Lognormal abundance distribution of woody species in a cerrado fragment. Revista Brasil. Bot. 28(1): 39-45.

- Soudi, P., 1993.Living treasure saving earth's threatened biodiversity. Dept. of the Environment, 53P.

- Vandermeer, J. H., 1981. Elementary mathematical ecology. Wiley Publishers, 294p.

(11)

397 - 387 . pp . 2010 , 3 . No , 63 . Vol , Iranian Journal of Natural Resources ,

Journal of Range and Watershed Management

397

Distributions of Species Diversity Abundance Models and Relationship between Ecological Factors with Hill (

N1

) Species

Diversity Index in 4 Range Sites of Isfahan Province

E. Ghehsareh Ardestani*1, M. Bassiri2, M. Tarkesh3 and M. Borhani4

1MSc.Graduate, Isfahan University of Technology, Isfahan, I.R. Iran

2 Associate Prof., College of Natural Resources, Isfahan University of Technology, Isfahan, I.R. Iran

3Assistant Prof., College of Natural Resources, Isfahan University of Technology, Isfahan, I.R. Iran

4Scientific member, Research Center for Agriculture and Natural Resources of Isfahan, Isfahan, I.R. Iran

(Received: 9 January 2010, Accepted: 20 September 2010)

Abstract

Our planet is threatened by nuclear wars, climate change and degradation of biodiversity. The first two dangers are relatively focused on, but biodiversity degradation is not properly considered yet.

Species diversity is mostly composed of biodiversity and is considered an index of changes in ecosystems. Many of researchers think of higher species diversity as an index of bio-system’s stability.

This research aimed to study parametric models of species diversity on range vegetations of study sites and relationship of ecological factors such as climate, soil and topography with diversity index. Four range sites were selected on different geographic areas in Isfahan province and randomly- systematic sampling was conducted to collect density data on plant species by quadrates along transects of each site during six years (2002-2007). Broken stick, log normal, logarithmic series and geometric series models were fitted for each site, and log normal showed to be the most proper model of distribution with P>0.05. LSD’s test indicated a significant different at α=1% level between species diversity of steppe1 and semi-steppe2 sites. Log normal model seemed to be the most proper model in all sites.

Generally, when this model is fitted successfully communities are considered stable. On the basis of some related studies with this research, Hill (N1) showed to be most proper index for species diversity in study sites, therefore effects of ecological factors such as soil, climate and topography on this index were studied by CCA method. Results of CCA revealed a high positive correlation between species diversity and percent organic matters of soil as well as precipitation levels and negative correlation with temperature.

Keywords: Species diversity, Species abundance model, Canonical correspondence analysis

*Corresponding author: Tel: +98 913 2111977 , Fax: +98 311 3912841 , E-mail: [email protected] 1- Artemisia sieberi community

2-Rangeland's communities of Zagross mountains

Referensi

Dokumen terkait

Vol , Iranian Journal of Natural Resources , Natural Environmental Journal of 223 Impacts of Shahid Rajaii Dam on Genetic Variation and Differentiation Siah Mahi, Capoeta

Vol , Iranian Journal of Natural Resources , Natural Environmental Journal of 277 An investigation on accumulation of heavy metals in roots and leaves of Avicennia marina the

Vol , Iranian Journal of Natural Resources , Natural Environmental Journal of 73 Habitat Suitability Modeling for Wild Goat Capra aegagrus in Kolah Ghazi National Park, Esfahan

Vol , Iranian Journal of Natural Resources , Natural Environmental Journal of 91 Study of the Effect of Benthic Community Structure of Taleghan River on the Determination of

Vol , Iranian Journal of Natural Resources , Natural Environmental Journal of 119 Study of Trend of Chemical and Microbial Changes of Rainbow Trout Oncorhynchus Mykiss to

Vol , ral Resources Iranian Journal of Natu , Natural Environmental Journal of Effects of Stunning by Ice and Clove Oil, and Asphyxia on Stress Responses and Quality

Vol , Iranian Journal of Natural Resources , Natural Environmental Journal of 187 Inhibitory Effect of Bacillus Licheniformis on G1 and G2 Aflatoxins in Culture Media and Kilka

Vol , Iranian Journal of Natural Resources , Natural Environmental Journal of Effects of Dietary Lipid Level on Growth, Feed Utilization and Survival of Juvenile of Beluga