Check-list of South Korean Diplazontinae (Hymenoptera

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Check-list of South Korean Diplazontinae (Hymenoptera: Ichneumonidae) with key to South Korean genera

Jin- Kyung Choi

^1,2

, Gyu-Won Kang

²

, Ekaterina N. Balueva

²

, Jong-Chul Jeong

³

and Jong-Wook Lee

²

*

1Department of Life Sciences, the Natural History Museum, Cromwell Road, London SW7 5BD, UK

2Department of Life Sciences, Yeungnam University, Gyeongsan 38541, South Korea

3National Park Research Institute, Korea National Park Service, Wonju, South Korea

한국산 등에살이뭉툭맵시벌아과 (벌목: 맵시벌과) 의 종 목록 및 속 검색표

최진경^1,2·강규원²·발루에바²·정종철³·이종욱²*

1런던자연사박물관, ²영남대학교 생명과학과, ³국립공원관리공단 국립공원연구원

Abstract :39 species of South Korean Diplazontinae are reviewed. Until now we confused Korean name of this subfamily. In this study, we suggest new Korean name and provide list of South Korean Diplazontinae and key to genera of Diplazontinae.

Key words :National park, new Korean name, Diplazontinae

요 약 :본 연구결과는 한국산 등에살이뭉툭맵시벌아과의 목록을 정리한 것이다. 등에류에 기생하는 것으로 알려진 본 아과의 국명에 ‘등에’와 ‘등애’가 혼동되어 사용되고 있어, 본 논문을 통해 모든 국명을 ‘등에’로 재정리 하고자 한 다. 이 중 20종의 경우 11개의 국립공원에서 채집된 기록이 있어 국립공원 종 목록에 신규로 추가하고자 한다.

주요어 :국립공원, 국명 개칭, 등에살이뭉툭맵시벌아과

Introduction

The subfamily Diplazontinae is a well-studied group of ichneumon wasp, widespread distributed. At present approxi- mately 340 species within 22 genera have been described, most of the species occur in Holarctic and Neotropical regions (Gauld et al. 1997; Townes 1971; Yu et al. 2012). In Eastern Palearctic fauna, Diplazontinae studied particularly.

The Russian Far East fauna of subfamily Diplazontinae was studied by Manukyan (2007). Japanese Diplazontinae were reviewed in several studies (Uchida 1957; Nakanishi 1978, 1979, 1985, 1986) and recorded 45 species from 12 genera.

Only 29 species distributed in seven genera were reported in large territories of China (all reported species were found in Xinjiang) (Chang 2007; Ma et al. 1992; Ma et al. 1993;

Wang et al. 1992). The Korean Diplazontinae were reviewed by Uchida (1930, 1957), Kim (1955), Yang et al. (2006),

Balueva & Lee (2014, 2015, 2016a, b). The current total 39 species within 12 genera of Diplazontinae were known from South Korea.

This subfamily is one of the more easily recognized ichneumonid subfamilies. Diagnostic features include the three-toothed mandibles, the rectangular first tergite, com- paratively short propodeum, short ovipositor, and transverse head (excluding genus Episemura).

The host range of Diplazontinae seems to be very narrow, with all confirmed host records being from the hoverfly subfamily Syrphinae (Diptera, Syrphidae). Except for the two species of the derived genus Bioblapsis Förster, Diplazon- tinae furthermore only attack the aphidophagous species of this group (Fitton & Rotheray, 1982).

In this study, we suggest new Korean name. Checklist of South Korean Diplazontinae, pictures of habitus each species and key to genera of South Korean Diplazontinae are provided.

*Corresponding author E-mail: [email protected]

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Materials and Methods

All photographs were obtained using an AxioCam MRc5 camera attached to a stereo microscope (Zeiss SteREO Discovery. V20; Carl Zeiss, Göttingen, Germany), processed using Axio Vision SE64 software (Carl Zeiss), and optimized with a Delta imaging system (i-solution; IMT i-Solution Inc., Vancouver, Canada). All the materials examined are deposited in the Department of Biology, Yeungnam University in Korea.

The terminology for morphological features mostly follows that of Klopfstein (2014). The following is a list of museum and institutions used in the taxonomic revision.

ANSP: Academy of Natural Sciences of Philadelphia, 19th and the Parkway, Philadelphia, Pennsylvania, 19103, U.S.A.

AEI: American Entomological Institute, 3005 S.W. 56th Avenue, Gainesville, Florida, 32608, U.S.A. (H. Townes collection.)

CNC: Canadian National Collections, Centre for Land and Biological Resources Research, Agriculture Canada, Ottawa, Ontario, K1A 0C6, Canada.

HU: Hokkaido University, Faculty of Agriculture, Entomo- logical Institute, Sapporo, Japan.

KU: Kyushu University, Entomological Laboratory, Fukuoka, Kyushu, Japan. (A. Nakanishi collection.)

MHN: Muséum d'Histoire Naturelle, Route de Malagnou, CH-1211 Genève, Switzerland.

MNCN: Departamento Entomologia, Museo Nacional de Ciencias Naturales, J. Gutierrez Abascal 2, E-28006 Madrid, Spain.

MPM: Department of Invertebrate Zoology, Milwaukee Public Museum, 800 W Wells, Milwaukee, Wisconsin, 53233, U.S.A.

NCM: Norwich Castle Museum, Norwich, United Kingdom.

NHM: The Natural History Museum, Department of Ento- mology, Cromwell Road, London, England, SW7 5BD, United Kingdom.

NR: Naturhistoriska Riksmuseet, Sektionen för Entomologi, S-104 05 Stockholm, Sweden.

UL: Universite Laval, Department de Biologie, Faculte des Sciences, Ste Foy, Quebec, G1K 7P4, Canada.

USNM: U.S. National Museum of Natural History, Smith- sonian Institute, Washington, D.C., 20560, U.S.A.

UU: Uppsala Universitet, Zoologiska Institutionen, Entomo- logiska Avdelningen, Villavägen 9, S-75236 Uppsala, Sweden. (Thunberg collection.)

ZI: Zoologiska Institutionen, Helgonavägen 3, S-223 62 Lund, Sweden.

ZM: Zoological Museum, P. Rautatiekatu 13, FIN-00015

Helsinki, Finland.

ZMHU: Zoologisches Museum (Museum für Naturkunde), Humboldt Universität, Invalidenstrasse 43, D-101115 Berlin, Germany.

ZSM: Zoologisches Staatsammlung, Münchhausenstrasse 21, D-81247 München, Germany.

The followings are abbreviations of Korean provinces and indices used in the paper.

GG, Gyeonggi-do; GW, Gangweon-do; CB, Chungcheong- buk-do; CN, Chungcheongnam-do; GB, Gyeongsangbuk- do; GN, Gyeongsangnam-do; JB, Jellabuk-do; JN, Jellanam- do; JJ, Jeju-do. TL: Type Locality; TS: Type species; TD:

Type depository.

Results

Check-list of South Korean Diplazontinae

Subfamily Diplazontinae 등에살이뭉툭맵시벌아과 (개칭)

Genus Bioblapsis Förster, 1869 배반짝뭉툭맵시벌속 Bioblapsis Förster 1869: 162. TS: Bassus flavipes Holmgren.

Trichomastix Vollenhoven 1878: 161. TS: Trichomastix polita Vollenhoven.

1. Bioblapsis polita (Vollenhoven, 1878) 배반짝뭉툭맵시벌 Trichomastix polita Vollenhoven, 1878: 161. Type: unknown.

Bassus tibialis Bridgman, 1883: 139. Type: unknown, TL:

United Kingdom; TD: NCM.

Material examined. [Korea] GW: 1♀, Donghae-si, Samhwa- dong, Mureung valley, MT, N 37°27'52″, E 129°1'26″, 27.ix-5.x.2007, J.W. Lee.

Korean Record: Balueva & Lee, 2014, Lee et al., 2015 Distribution: Korea, Japan, Russia, Belarus, Belgium, Czech

Republic, Czechoslovakia, Finland, France, Germany, Hungary, Netherland, Poland, Sweden, Ukraine, United Kingdom.

Genus Campocraspedon Uchida, 1957 긴배뭉툭맵시벌속 Campocraspedon Uchida 1957: 237. TS: Campocraspedon

satoi Uchida.

2. Campocraspedon satoi Uchida, 1957 긴배뭉툭맵시벌 Campocraspedon satoi Uchida 1957: 237. Type: ♀; TD:

AEI.

Material examined. This species was not found in the

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materials of Korea.

Korean Record: Uchida, 1957; Nakanishi, 1985; Balueva &

Lee, 2014.

Distribution. Korea, Japan.

3. Campocraspedon elongatus Nakanishi, 1985 참긴배뭉툭맵시벌

Campocraspedon elongatus Nakanishi, 1985: 31. Type: ♀, TL: Japan; TD: HU.

Material examined. [Korea] GB: 1♀, Gyeongsan-si, Dae- dong, Yeungnam University, MT, 1-8.2009, J.W. Lee.

Korean Record: Balueva & Lee, 2014.

Distribution. Korea, Japan. Russia.

Genus Diplazon Nees, 1819 등에살이뭉툭맵시벌속 Diplazon Nees, 1819: 292. TS: Ichneumon laetatorius Fabricius.

4. Diplazon koreanus Balueva and Lee, 2015 고려등에살이뭉툭맵시벌 (개칭)

Diplazon koreanus Balueva & Lee, 2015: 562. Type: ♀, TL:

Korea; TD: YNU.

Material examined. [Korea] JB: 1♀, Jeongeup-si, Ibam- myeon, Deungcheon-ri, Ibamm Wetland, N 35°33', E 126°50', 21.vi.2005, coll. J.G. Han; GB: 1♀, 1♂Gyeongsan-si, Dae- dong, Yeungnam Univ., 4.vi.1988, coll. E.S. Kim; 1♂, Baegun-ri, Seongju-gun, Yonggipokpo, 17.vi.2000; 1♀, Cheongdogun, Unmun-myeon, Mt. Unmun, 23.v. 2008, coll. J.W. Lee; GG: 1♂, Gwacheon-si, Makgye-dong, Mt.

Cheonggye, 4.vi.1989, coll. S.M. Ryu; 1♂, Gogang-dong, Oieong-gu, Bucheon-si, 18.v.1997, coll. J.Y. Jang; 1♂, Seoul, Gangnam-gu, Gaepo-dong, Mt. Daemo, 9.vi.1996, coll. H.J. Jung; 1♂, Seoul, Nowon-gu, Sanggyedong, Mt.

Surak, 29.v.1997, coll. S.Y. Kim; 1♀, Seoul, Dobong-gu, Dobong-dong, Mt. Bukhan, 25.v.1998, coll. M.A. Nam.

Korean Record: Balueva & Lee, 2015, Lee et al., 2016.

Distribution. Korea.

Remark. Bukhansan National Park (New record), Naejangsan National Park (New record).

Lee 등(2016)에본종의국명이 ‘한국살이뭉툭맵시벌’ 로기록되어있으나한국살이뭉툭맵시벌은 Balueva &

Lee(2014)에서 Eurytyloides koreanus의 국명으로 보고 되었기에 본 논문을 통해 Diplazon koreanus는 고려등 에살이뭉툭맵시벌로 개칭한다.

5. Diplazon laetatorius (Fabricius, 1781) 등에살이뭉툭맵시벌

Ichneumon laetatorius Fabricius, 1781: 424. Type: ♀, TL:

Germany; TD: lost.

Ichneumon dichrous Schrank, 1781: 352. Type: ♀, TL:

Austria; TD: lost.

Anomalon attractus Say, 1835: 241. Type: ♀, TL: USA; TD:

USNM.

Bassus albovarius Wollaston, 1858: 23. Type: ♀, TL: Madeira Islands; TD: NHM.

Bassus cinctipes Holmgren, 1868: 409. Type: ♀, TL: South Africa; TD: NR.

Bassus sycophanta Cresson, 1868: 112. Type: lost.

Bassus tripicticrus Walsh, 1873: 85. Type: lost.

Scolobates varipes Smith, 1878: 3. Type: ♀, TL: New Zealand;

TD: NHM.

Bassus balearicus Kriechbaumer, 1894: 246. Type: ♀, TL:

Spain; TD: MNCN.

Bassus terminalis Davis, 1895: 30. Type: ♀, TL: U.S.A; TD:

ANSP.

Bassus generosus Cameron, 1898: 31.Type: ♀, TL: New Zealand; TD: NHM.

Bassus senegalensis Ferrière, 1925: 121. Type: ♀, TL: Senegal;

TD: MHN.

Bassus ikiti Cheesman, 1936: 182. Type: ♀, TL: Vanuatu;

TD: NHM.

Bassus ikiti Cheesman, 1936: 182. Type: ♀; TD: NHM.

Material examined. [Korea] 1♀, Seoul, Dongsun-dong, 3.vii.1980, coll. J.I. Kim; 1♀, CN, Seosan-si, Haemimyeon, Daegok-ri, Hanseo Univ., 9–21.x.2006; GB: 1♀, Gyeongsan- si, Dae-dong, Yeungnam Univ., 29.V.1983, coll. H.G. Kim;

3♀, ditto, 14.iv.1986, coll. J.Y. Cha; 1♀, ditto, 10.vii.1986, coll. J.W. Lee; 7♀, ditto, 13.iv.1987, coll. E.S. Kim; JB:

1♀, Jeongeup-si, Naejang-dong, Naejangsa, Mt. Naejang- san, 11.v.2007, coll. J.C. Jeong; JN: 1♀, Suncheon-si, Songgwang, Pyeongni, Songgwangsa, 30.v.1981, coll. S.M.

Ryu; 1♀, Suncheon-si, Seungju-eup, Seonamsa, Mt. Jogye, 5–8.v.1990, coll. J.W. Lee; JJ: 1♀, Jeju-si, Jejudaehak-ro, Jeju Nat. Univ., 26.v–2.vi.2008, coll. J.W.Lee, (GenBank Acc. Nr. KP750191); GW: 1♀, Chuncheon-si, Sawong- gong, Gwangwon Prov., Alt. 81m, 2–15.v.2012, (GenBank Acc. Nr. KP750192); 1♀, Daegu-si, Dalsung-gun, Hyunpoong- myeon, Hyunpoongjoong, 26.v.2008, coll. S.G. Lee, (GenBank Acc. Nr. KP750193); JB: 1♀, Namwon-si, Sannae-myeon, Jeongteongchi, 2.ix–15.x.2001, coll. J.W.

Lee, (GenBank Acc. Nr. KP750194); 1♀, GB, Gyeongsan- si, Dae-dong, Yeungnam Univ., 27.v.2008, coll. S.G. Noh, (GenBank Acc. Nr. KP750195); 1♀, JN, Jeongeup-si, Ssangam-dong, Seoraebong (PFT), 16.vi.2004, coll. J.G.

Han, (GenBank Acc. Nr. KP750196); 1♀, Daegu-si, Dalseo-gu, Daegok-dong, Daegu Arboretum (Alt.88mm),

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4–18.iv.2012, coll. S.G. Kang, (GenBank Acc. Nr.

KP750197); 1♀, JB, Jeongeup-si, Yongsan-dong above Ternel, 15.vi.2004, (GenBank Acc. Nr. KP750198); 1♀, GN, Haman-gun, Daesan-myeon, 26.iv.1991, (GenBank Acc. Nr. KP750199); 1♀, GB, Cheongdo-gun, Gakbuk- myeon, Namsan-3ri, 6–20.vii.2008, coll. J.W. Lee, (GenBank Acc. Nr. KP750200); 1♀, CN, Seosan-si, Haemi-myeon, Daegok-ri, Hanseo Univ., 21.vi–27.vii.2011, coll. J.K.

Kim, (GenBank Acc. Nr. KP750201); 1♀, GN, Sancheong- gun, Geumseo-myeon, Wangdeungjae, 31.viii–29.x.2001, coll. J.W. Lee, (GenBank Acc. Nr. KP750202); 1♀, GB, Gyeongsan-si, Dae-dong, Yeungnam Univ., 17–24.iv.2009, coll. J.W. Lee, (GenBank Acc. Nr. KP750203); 1♀, Seoul, Dongdaemun-gu, Cheongyangri-dong, 26.ix–4.x.2005, coll.

W.I. Choi, (GenBank Acc. Nr. KP750204); 2♀, Busan-si, Yeongdo-gu, Dongsam-dong,Gosin Univ., 27.iv–7.v.2008, coll. J.W. Lee, (GenBank Acc. Nr. KP750205; Nr.

KP750206); 2♀, JB, Jeongeupsi, Naejang-dong, Ansan, 19.v.2004, coll. M.K. Yun, (GenBank Acc. Nr. KP750207, Nr. KP750208); 2♀, Daegu-si, Hwawon-eup, 1.vi.2008, coll. J.Y. Cho, (2♀, GenBank Acc. Nr. KP750209; Nr.

KP750210); 2♀, GW, Chuncheon-si, 2–15.v.2012, (GenBank Acc. Nr. KP750211; Nr. KP750212); 2♀, CN, Seosan-si, Haemi-myeon, Daegok-ri, Hanseo Univ., 1–

20.vi.2011, coll. J.G. Kim, (GenBank Acc. Nr. KP750213;

Nr. KP750214); 1♀, CN, Seosan-si, Haemi-myeon, Daegok-ri, Hanseo Univ., 20.vii–24.viii.2011, coll. J.G.

Kim, (GenBank Acc. Nr. KP750215); 2♀, GB, Cheongdo- gun, Gakbuk-myeon, Namsan-3ri, 6–20.vii.2008, coll. J.W.

Lee, (GenBank Acc. Nr. KP750216; Nr. KP750217); 1♀, GB, Cheongdo-gun, Gakbuk-myeon, Namsan-3ri, 1.viii–

15.ix.2013, coll.J.W. Lee, (GenBank Acc. Nr. KP750218);

1♀, GB, Gyeongsan-si, Dae-dong, Yeungnam Univ., 16.vi.2006, coll. J.W. Lee, (GenBank Acc. Nr. KP750219);

1♀, JN, Jeongeup-si, Ssangam-dong, Seoraebong, 16.vi.2004, (GenBank Acc. Nr. KP750220); 1♀, CN, Buyeo-gun, Gyuam-myeon, Sumok-ri, 13–27.vi.2005, (GenBank Acc.

Nr. KP750221); 2♀, GB, Cheongdo-gun, Gakbuk-myeon, Namsan-3ri, 10–30.vi.2013, coll. J.W. Lee, (GenBank Acc.

Nr. KP750222; Nr. KP750223); 1♀, GW, Gangwon Prov.

Environment Research Park, Bukbang, N35°45'15", E127°51'1", 30.v–15.vi.2012, coll. S.J. Jang, (GenBank Acc. Nr. KP750224); 1♀, GN, Haman-gun, Gunbung- myeon, Seokgyocheon, 24.viii.2001, (GenBank Acc. Nr.

KP750225).

Korean Record: Kim, 1955; Kim, 1970; Uchida, 1957;

Townes et al., 1965; Yang et al., 2006; Choi et al., 2012;

Balueva & Lee, 2015.

Distribution: Korea, Japan, Russia, Holarctic.

6. Diplazon pallicoxa Manukyan, 1987 황다리등에살이뭉툭맵시벌

Diplazon pallicoxa Manukyan, 1987: 70. Type: ♀, TL: Russia;

TD: unknown.

Material examined. [Korea] GB: 1♀, Gyeongsan-si, Dae- dong, Yeungnam Univ., 30.viii.1988; 1♀, Seoul, Mt.

Daemo, Gaepo-dong, Gangnam-gu, 9.v.1998, coll. H.Y.

Park; GN: 1♀, Jinju-si, Gajwa-dong, Gyeongsang Univ., 22–28.iv.1989; JB: 1♀, Jeongeup-si, Naejangdong, Wonjeokam, Mt. Naejang (MT), site 8, 28.v.2006, coll.

J.W. Lee; 1♀, Jeongeup-si, Naejangdong, Wonjeokam, Mt.

Naejang (MT), site 14, 8–14.vi.2008, coll. J.W. Lee; 1♀, Iksan-si, Sinyon-dong, Wongwang Univ. (MT), 28.viii–

22.ix.2006; JN: 1♀, Naju-si, Dado-myeon, Pungsan-ri (MT -II), 30.ix–4.x.2005, coll. S.B. Yu.

Distribution: Korea, Russia, Armenia, Finland, Switzerland.

Remark. Naejangsan National Park (New record).

Figures 1-6. Habitus of Diplazontinae spp. (1) Bioblapsis polita (Vollenhoven, 1878), (2) Campocraspedon elongatus Nakanishi, 1985, (3) Diplazon koreanus Balueva and Lee, 2015, (4-♀, 5-♂) Diplazon laetatorius (Fabricius 1781), (6) Diplazon pallicoxa Manukyan, 1987.

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7. Diplazon pectoratorius (Thunberg 1822) 참홍다리뭉툭맵시벌

Ichneumon pectoratotius Thunberg, 1822: 280. Type: ♂, TL:

Sweden; TD: UU.

Ichneumon angustorius Thunberg, 1824: 366. Type: ♀, TL:

Sweden; TD: UU.

Bassus urupensis Uchida, 1935: 118. Type: ♀, TL: Russia;

TD: HU.

Bassus pulchripes Provancher, 1875a: 143. Type: ♀, TL:

Canada; TD: UL.

Homocidus akaashii Uchida, 1931: 24. Type: ♂, TL: Korea;

TD: HU.

Polysphincta pleuralis Provancher, 1875b: 312. Type: ♀, TL:

Canada; TD: UL.

Material examined. [Korea] CB: 1♂, Dannyang-gun, Gagok-myeon, Eouigok-ri, Sobaeksan National Park, Huibang-valley, 3.v.1997, coll. D.K. Chung; 1♂, Mungyeong- si, Dongro-myeon, Saengdal-ri, 25.v.2007, coll. S.J. Park;

GG: 1♀, Namnyangju-si, Jinjeop-eup, Pyeong-ri 247, Gwangneung, 20.v.1990, coll. J.A. Lee; 1♂, Yangpyong- gun, Mt. Yongmun, Yongmunsa, 8.v.1998, coll. C.Y. Cho;

GW: 1♂, Pyeongchang-gun, Jinbu-myeon, Dongsan-ri, Odaesan National Park, 26.v.1995, coll. J.M. Ryu; 1♂, Taebaek-si, Mt. Yeunhwa-san, 14.v.1997; 1♀, Donghae-si, Samhwa-dong, Mureung-valley, 10.v.2007, coll. J.W. Lee.

Korean Record: Kim, 1955; Townes et al., 1965, Choi et al., 2012; Balueva & Lee, 2015.

Distribution: Korea, Russia, Europe, Holarctic.

Remark. Odaesan National Park (New record), Sobaeksan National Park (New record).

8. Diplazon scutatorius Teunissen, 1943 등검정등에살이뭉툭맵시벌

Diplazon scutatorius Teunissen, 1943: 50–52. Type: ♀, TL:

Netherland; TD: unknown.

Diplazon pilosus Uchida, 1957: 235. Type: ♂, TL: Japan;

TD: HU.

Material examined. [Korea] 1♀, Seoul, Dobong-gu, Dobong- dong, Bukhansan National Park, 9.v.1989, coll. S.Y. Shin;

9♀, Daejeon, Donggu, Daejeon Univ., 12.iv–12.v.2007, coll. J.W. Lee; CB: 1♀, Boeun-gun, Songni-san National Park, 4.viii–26.ix.2007; GB: 1♀, Gyeongsan-si, Dae-dong, Yeungnam Univ., 3.ix.1988, coll. E.S. Kim; 2♀, Yeongyang- gun, Irwol-myeon, Mt. Irwolsan, 26.vi–15.vii.2014, coll.

H.Y. Han; GW: 4♀, Prov. Enviroment Research Park, Bukbang, 30.v–15.vi.2012, coll. S.J. Jang; JB: 1♀, Jeongeup- si, Naejang-dong, Ansarn site-10, 19.v.2004, coll. M.K.

Yun; 1♀, Sunohang-gun, Bokheing-myeon, Songgotbawi, 03.vi.2004, coll. K.B. Kim; 8♀, Wanju-gun, Dongsang- myeon, Daea Arboretum, 19–30.ix.2012, coll. J.M. Park.

Distribution. Korea, Japan, Russia, Europe.

Remark. Bukhansan National Park (New record), Naejangsan National Park (New record), Songnisan National Park (New record).

9. Diplazon suigensis Uchida, 1957 수원뭉툭맵시벌 Diplazon suigensis Uchida, 1957: 233. Type: ♀, TL: Korea;

TD: HU.

Material examined. [Korea] 1♀, Seoul, Geumcheon-gu, Siheung-dong, Mt. Cheonggye, 21.v.1991, coll. H.J. Ku;

2♀, Daejeon, Donggu, Daejeon Univ., 12.iv–12.v.2007, coll. J.W. Lee; GB: 1♀, Gyeongsan-si, Dae-dong, Yeungnam Univ., 9.ix.1988, coll. E.S. Kim; 1♀, Cheongdo-gun, Unmun-myeon, Geungolgyegok Valley, 12.v–8.viI. 2013, coll. J.W. Lee; GW: 1♀, Donghae-si, Mereuggyegok Valley, 2–15.v.2007, coll. J.W. Lee; JB: 1♀, Iksan-si, Sinyon-dong, Wongwang Univ., 28.viii–22.ix.2006.

Korean Record: Kim, 1970; ESK and KSAE, 1994; Choi et al., 2012; Balueva & Lee, 2015.

10. Diplazon tetragonus tetragonus (Thunberg, 1824) 검정등에살이뭉툭맵시벌

Ichneumon tetragonus Thunberg, 1822: 280. Type: ♂, TL:

Sweden; TD: UU.

Ichneumon hortorius Thunberg, 1822: 280.Type: ♂, TL:

Sweden; TD: UU.

Bassus tricinctus Gravenhorst, 1829: 351. Type: lost.

Bassus nemoralis Holmgren, 1856: 354. Type: ♀, TL: Sweden;

TD: NR.

Bassus concinnus Cresson, 1868: 111. Type: ♀, TL: U.S.A.;

TD: ANSP.

Bassus albicoxus Provancher, 1874a: 31. Type: ♂, TL:

Canada; TD: UL.

Bassus amoenus Provancher, 1874b: 55. Type: ♀, TL: Canada;

TD: UL.

Diplazon tetragonopsis Uchida, 1957: 234. Type: ♀, TL:

Japan; TD: HU.

Material examined. [Korea] CN: 1♀, Seosan-si, Haemi- myeon, Daegok-ri, Hanseo Univ., 17.iv–10.v.2006; 1♀, ditto, 3–10.vii.2008, coll. J.W. Lee; GB: 2♀, Cheongdo- gun, Unmun-myeon, Ssalbawi (U7), 12.v–9.vi.2012, coll.

J.W. Lee; 1♀, Cheongdo-gun, Unmun-myeon, Haksodae-

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pokpo (U5), 29.viii–11.x.2014, coll. J.W. Lee; GW: 2♀, Donghae-si, Samhwa-dong, Mureung-valley, 29.iv–

26.v.2007; JB: 4♀, Wanju-gun, Dongsang-myeon, Daea Arboretum, 19–30.ix.2012, coll. J.M. Park.

Korean Record: Uchida, 1957; Balueva & Lee, 2015.

11. Diplazon tibiatorius tibiatorius (Thunberg, 1824) 종아리등에살이뭉툭맵시벌

Ichneumon tibiatorius Thunberg, 1822: 280. Type: ♀, TL:

U.S.A; TD: ANSP.

Bassus albosignatus Gravenhorst, 1829: 343. Type: lost.

Material examined. [Korea] GW: 1♀, Donghae-si, Samhwa- dong, Mureung-valley, N 37°27', E 129°1', 25.iv–

20.vii.2007, coll. J.W. Lee.

Distribution: Korea, Japan, Russia, Mongolia.

12. Diplazon varicoxa (Thomson, 1890) 검정다리등에살이뭉툭맵시벌

Bassus varicoxa Thomson, 1890: 1468. Type: ♀, TL: Sweden;

TD: ZI.

Bassus japonicus Ashmead, 1906: 170. Type: ♂, TL: Japan;

TD: USNM.

Material examined. [Korea] 1♂, Daejeon-si, Donggu, Daejeon University, 13–28.iv.2006.

Distribution: Korea, China, Japan, Russia, Afghanistan, Austria, Belgium, Czechoslovakia, Finland, France, Germany, Hungary, India, Ireland, Italy, Mongolia, Norway, Poland, Spain, Switzerland, United Kingdom.

Genus Enizemum Förster 1869 검정다리배줄뭉툭맵시벌속

Enizemum Förster 1869: 142. TS: Bassus tibialis Cresson.

13. Enizemum nigrocoxatum Balueva and Lee, 2016 검정다리배줄뭉툭맵시벌

Enizemum nigrocoxatum Balueva & Lee, 2016a: 46. Type:

♀, TL: Korea; TD: YNU.

Material examined. [Korea] CB: 1♀, Dannyang-gun, Gagok-myeon, Eouigok-ri, Mt. Sobaek, Huibang-valley, 9.viii.1994, S.M. Ryu; GW: 1♀, Donghae-si, Samhwa- dong, Mureung-valley, 29.iv–26.v.2007; GB: 1♀, Gyeongsan- si, Dae-dong, Yeungnam University, MT, 29.ix.1986, J.M.

Choi.

Korean Record: Balueva & Lee, 2016a.

Remark. Sobaeksan National Park (New record).

Genus Eurytyloides Nakanishi, 1978 한국살이뭉툭맵시벌속

Eurytyloides Nakanishi 1978: 429. TS: Eurytyloides kusigematii Nakanishi.

14. Eurytyloides koreanus Balueva and Lee, 2014 한국살이뭉툭맵시벌

Eurytyloieds koreanus Balueva and Lee, 2014: 152. Type: ♀, TL: Korea; TD: YNU.

Material examined.[Korea] GW: 1♀, Yanggu, Gachilbong, 31.v.1992, coll. J.W. Lee.

Genus Homotropus Förster 1869 참어리등에살이뭉툭맵시벌속 Figures 7-12. Habitus of Diplazontinae spp. (7) Diplazon pecto-

ratorius (Thunberg 1822), (8) Diplazon scutatorius Teunissen, 1943, (9) Diplazon suigensis Uchida, 1957, (10) Diplazon tetragonus (Thunberg 1824), (11) Diplazon tibiatorius (Thunberg 1824), (12) Diplazon varicoxa (Thomson 1890).

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Homotropus Förster 1869: 157. TS: Bassus elegans Graven- horst.

15. Homotropus areolaris (Uchida, 1957) 경포참어리등에살이뭉툭맵시벌

Syrphoctonus areolaris Uchida, 1957: 237. Type: ♀, TL:

Russia; TD: HU.

Material examined. [Korea] GB: 3♀, Yeongyang-gun, Irwol-myeon, Mt. Irwolsan, 22.v–7.vi.2014, coll. H.Y.

Han; 1♀, ditto, 22.v–7.vi.2014, H.Y. Han; 1♀, Yeongju-si, Punggi-eup, Sobaeksan National Park, Jungnyeong, 12–

22.vi.2009.

16. Homotropus frontorius (Thunberg, 1824) 붉은가슴참어리등에살이뭉툭맵시벌

Ichneumon frontorius Thunberg, 1824: 297. Type: ♂, TL:

unknown; TD: UU.

Homocidus subopacus Stelfox, 1941: 115. Type: ♀, TL:

United Kingdom; TD: USNM.

Homotropus quadrangularis Dasch, 1964: 106. Type: ♀, TL:

Canada; TD: CNC.

Material examined. [Korea] GW: Donghae-si, Samhwa- dong, Mureung-valley, 10.v.2007, J.W. Lee.

17. Homotropus longiventris Thomson 1890 긴배참어리등에살이뭉툭맵시벌

Homotropus longiventris Thomson, 1890: 1476. Type: ♀, TL: Sweden; TD: ZI.

Material examined.[Korea] GW: 1♀, Donghae-si, Samhwa- dong, Mureung-valley, 26.v–29.vi.2007.

Distribution. Korea, Russia, Belgium, Finland, Germany, Ireland, Lithuania, Mongolia, Netherlands, Norway, Poland, Romania, Sweden, Switzerland, United Kingdom.

18. Homotropus megaspis Thomson 1890 꼬마참어리등에살이뭉툭맵시벌

Homotropus megalaspis Schulz, 1906: 115. Type: ♀, TL:

Germany; TD: ZSM.

Material examined. [Korea] JB: 1♀, Jeongeup-si, Jangseong- gun, Bukha-myeon, Sajabong (24 site), MT, 20.vii.2005,

J.W. Lee; JJ: 1♀, Jeju-si, Ara-dong, Ara valley, MT, 2–

21.vi.2008, J.W.Lee.

Distribution. Korea, Japan, Russia, Belgium, Czech Republic, Czechoslovakia, Finland, Germany, Hungary, Ireland, Lithuania, Norway, Poland, Switzerland, United Kingdom.

19. Homotropus momoii (Uchida 1957) 모모이참어리등에살이뭉툭맵시벌

Syrphoctonus momoi Uchida, 1957: 262. Type: ♀, TL: Japan;

TD: HU.

Material examined. [Korea] 1♀, Daegu-si: Dalseo-gu, Daegok-dong, Daegu Arboretum (Alt. 88 m), 3–16.v.2012, coll. S.G. Kang; 1♀, Nam-gu, Daemyeong-dong, Mt.

Apsan, 19.v.2002; 5♀, Daejeon-si, Donggu, Daejeon Univ., 1–27.ix.2006, coll. J.W. Lee; 1♀, ditto, 8.x–

30.xi.2007, J.W. Lee; 1♀, Ulsan-si, Ulju-gun, Sangbuk- myeon, Mt. Gaji, 21–26.vi.1989; CB: 1♀, Jecheon-si, Figures 13-18. Habitus of Diplazontinae spp. (13) Enizemum nigrocoxatum Balueva and Lee, 2015, (14) Eurytyloieds koreanus Balueva and Lee, 2014, (15) Homotropus areolaris (Uchida, 1957), (16) Homotropus frontorius (Thunberg 1824), (17) Homotropus longiventris Thomson 1890, (18) Homotropus megaspis Thomson 1890.

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Baegun-myeon, Mt. Sambong, 28.v.1997, D.K. Chung;

CN: 3♀, Seosan-si, Haemi-myeon, Daegok-ri, Hanseo Univ., 30.iv–9.v.2006; 2♀, ditto, 19.v–4.vi.2006, J.W. Lee;

GB: 1♀, Gyeongsan-si, Dae-dong, Yeungnam University, MT, 7.v.1987, J.W. Lee; 1♀, ditto, 30.v.1991, H.J. Kim;

GG: 1♀, Anyang-si, Manan-gu, Gwanag Arboretum, Alt.

126m, MT, 16–29.x.2008, J.O. Lim; GW: 1♀, Donghae-si, Samhwa-dong, Mureung-valley, 21–30.v.2005; 1♀, ditto, 28.v–11.vi.2006, J.W. Lee; 2♀, ditto, 29.iv–26.v.2007; 4♀, ditto, 20–25.vi.2012; JB: 1♀, Jeongeup-si, Ssangam-dong, Daegok-ri, MT, 19.v.2005, K.B. Kim.

20. Homotropus sepiatus Balueva and Lee, 2015 검은참어리등에살이뭉툭맵시벌

Homotropus sepiatus Balueva & Lee, 2016a: 52. Type: ♀, TL: Korea; TD: YNU.

Material examined. [Korea] 1♀, Daejeon-si, Donggu, Daejeon Univ., 1–17.v.2006; GW: 1♀, Donghae-si, Samhwa-dong, Mureung-valley, 13–20.v.2006; 1♀, ditto, 28.v–11.vi.2006, J.W. Lee; 1♀, ditto, 20–25.iv.2007, J.W.

Lee.

21. Homotropus spinosus (Uchida, 1957) 검은얼굴참어리등에살이뭉툭맵시벌

Syrphoctonus spinosus Uchida, 1957: 263. Type: ♀, TL:

Japan; TD: HU.

Material examined. [Korea] GB: 2♀, Gyeongsan-si, Dae- dong, Yeungnam University, MT, 6–20.v.2013, J.W. Lee;

3♀, ditto, 26.vi–15.vii.2014, H.Y. Han; GG: 1♀, Anyang- si, Manan-gu, Gwanag Arboretum, Alt. 126m, MT, 5–

19.vii.2007, J.O. Lim; GN: 1♀, Hapcheon-gun, Myosan- myeon, Banpo-ri, Gayasan National Park, 22.v.2000, S.H.

Baek; GW: 2♀, Samcheok-si, Miro-myeon, Mt. Dootasan, 21.v. 2005, J.W. Lee; 1♀, Wonju-si, Heungeop-myeon, yeonse Univ., Wonju campus, MT, 20.iv–24.v. 2007; JN:

1♀, Jangseong-gun, Bukha-myeon, Cheongnyangwon, Mt.

Naejang, MT (site 25), 26.IV–4.VI.2006; JB: 1♀, Jeongeup-si, Ibammyeon, Deungcheon-ri, Wetland, 21.VI.

2005, coll. J.G. Han.

Korean Record: Lee et al., 2016; Balueva & Lee, 2016a.

Remark. Gayasan National Park (New record), Naejangsan National Park (New record).

본 종의 국명이 영문으로 표기 되어 Lee 등(2016)에서 검은에골참어리등에살이뭉툭맵시벌로 잘못 표기 되 었기에, 본 논문을 통해 ‘검은얼굴참어리등에살이뭉 툭맵시벌’로 바로 잡는다.

22. Homotropus tenuitebialis (Uchida, 1957) 황참어리등에살이뭉툭맵시벌

Syrphoctonus tenuitibialis Uchida, 1957: 261. Type: ♂, TL:

Japan; TD: HU.

Material examined. [Korea] GW: 1♂, Donghae-si, Samhwa- dong, Mureung-valley, 21.iv–2.v.2007, J.W. Lee.

Genus Phthorima Förster, 1869 납작칼배뭉툭맵시벌속 Phthorima Förster, 1869: 162. TS: Bassus compressus Desvignes.

Phthorimus Thomson, 1890: 1474. Emendation for Phthorima.

Figures 19-24. Habitus of Diplazontinae spp. (19) Homotropus momoii (Uchida 1957), (20) Homotropus sepiatus Balueva and Lee, 2015, (21) Homotropus spinosus (Uchida 1957), (22) Homo- tropus tenuitebialis (Uchida 1957), (23) Phthorima compressa (Desvignes 1856), (24) Promethes compressus Nakanishi, 1986.

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23. Phthorima compressa (Desvignes, 1856) 납작칼배뭉툭맵시벌

Bassus compressus Desvignes, 1856: 1–120. Type: ♀, TL:

United Kingdom; TD: NHM.

Bassus ibalioidis Kriechbaumer, 1878: 211. Type: ♀, TL:

Germany; TD: ZSM.

Homotropus niger Morley, 1906: 434. Type: ♂, TL: United Kingdom; TD: NHM.

Material examined. [Korea] GW: 1♀, Ganseong, Geon- bongsa, 22.v.1992, J.W. Lee.

Korean Record: Balueva & Lee, 2014; Lee et al., 2015.

Distribution: Korea, Austria, Belgium, Canada, Czechoslo- vakia, Finland, France, Hungary, Ireland, Isle of Man, Italy, Latvia, Moldova, Poland, Romania, Russia, Spain, Sweden, Switzerland, U.S.A., United Kingdom.

Remark. 본 종의 국명은 Balueva & Lee(2014)에서 납작 칼배뭉툭맵시벌로 명명하였으나, Lee 등(2015)에서 납 작볼뭉툭맵시벌로 잘못 표기 되었음을 밝힌다.

Genus Promethes Förster 1869 납작볼록뭉툭맵시벌속 Promethes Förster 1869: 162. TS: Bassus sulcator Graven-

horst.

Promethus Thomson 1890: 1475. Emendation for Promethes.

Liopsis Förster 1869: 162. TS: Bassus sulcator Gravenhorst.

24. Promethes compressus Nakanishi, 1986 납작볼뭉툭맵시벌

Promethes compressus Nakanishi, 1986: 127. Type: ♀, TL:

Japan; TD: KU.

Material examined. [Korea] GW: 1♀, Donghae-si, Samhwa- dong, Mureung valley, MT, N 37°27'52″, E 129°1'27″, 21- 30.v.2005.

Korean Record: Balueva & Lee, 2014; Lee et al., 2015.

Remark. 본 종의 국명은 Balueva & Lee(2014)에서 납작 볼뭉툭맵시벌로 명명되었으나, Lee 등(2015)에서 납작 칼배뭉툭맵시벌로 잘못 표기 되었음을 밝힌다

Genus Sussaba Cameron, 1909 호리뭉툭맵시벌속 Sussaba Cameron 1909: 728. TS: Sussaba bicarinata Cameron.

25. Sussaba cognata albicoxa Thomson, 1890 흰밑마디호리뭉툭맵시벌

Promethus albicoxa Thomson 1890: 1479. Type: male; TD:

ZI.

Material examined.[Korea] GN: 1♀, Yeongju-si, Punggi-

eup, Sobaeksan National Park, Jungnyeong, site 99, 12. vi- 23. vii. 2012, S.H. Oh; GB: 1♀, Cheongdo-gun, Gakbuk- myeon, Namsan-3ri, MT, N35°41', E128°35', 29. vi-6. vii.

2008.

Distribution: Korea, Japan, Russia, Austria, Belgium, Czechoslovakia, Denmark, Finland, France, Germany, Hungary, India, Ireland, Italy, Lithuania, Netherlands, Norway, Poland, Romania, Sweden, United Kingdom.

26. Sussaba sugiharai (Uchida, 1957) 스기하라호리뭉툭맵시벌

Promethes sugiharai Uchida, 1957: 245. Type: ♀, TL: Russia;

TD: HU.

Material examined.[Korea] GW: 1♀, Wonju-si, Baegunsan, 6-19.vi.2011, coll. H.Y. Han; 1♀, Wonju-si, Heungeop- myeon, Yeonse Univ., Wonju campus, 29.vi-15.vii.2011;

1♀, Chuncheon-si, Sanong-dong, Gwangwon Prov. Arboretum, Alt. 81m, N 35°38'08″, E 129°1'27″, 2-15.v.2012, G.Y.

Lee; 1♀, Gwangwon Prov. Environment Research Park, Bukbang, N 35°45'15.6″, E 127°51'1.7″, 30.v-15.vi.2012, S.J. Jang.

Distribution: Korea, Japan, Russia, Taiwan.

Genus Syrphoctonus Förster, 1869 어리등에살이뭉툭맵시벌속 (개칭)

Syrphoctonus Förster 1869: 162. TS: Bassus exsultans Graven- horst.

27. Syrphoctonus laevis Brues, 1908 레비스어리등에살이뭉툭맵시벌

Syrphoctonus laevis Brues, 1908: 48–56. Type: ♀, TL:

U.S.A.; TD: MPM.

Material examined. [Korea] GB: 1♂, Gyeongsan-si, Dae- dong, Yeungnam Univ., 8.v.1989, J.Y. Cha; GN: 1♂, Jinju- si, Gajwa-dong, Gyeongsang Univ., 22–28.iv.1989.

Distribution: Korea, Canada, U.S.A.

28. Syrphoctonus tarsatorius (Panzer, 1809) 어리등에살이뭉툭맵시벌 (개칭)

Bassus tarsatorius Panzer, 1809: 8–10. Type: lost.

Bassus exsultans Gravenhorst, 1829: 328. Type: lost.

Bassus insignis Gravenhorst, 1829: 349. Type: lost.

Bassus flavus Desvignes, 1862: 219. Type: ♂, TL: United

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Kingdom; TD: NHM.

Bassus indicus Cameron, 1909: 728. Type: ♂, TL: India; TD:

NHM.

Homocidus tarsatorius Morley, 1911: 97. Type: ♀, TL: lost.

Homotropus flavitrochanterus Uchida, 1956: 99. Type: ♀, TL: Japan; TD: HU.

Material examined. [Korea] 1♀, Daejeon-si, Donggu, Daejeon Univ., 13–28.iv.2006, J.W. Lee; 3♀, ditto, 1–

17.v.2006, J.W. Lee; 1♀, ditto, 1–27.ix.2006, J.W. Lee; 1♀, ditto, 12.iv–12.v.2007, J.W. Lee; GB: 1♀, Cheongdo-gun, Gakbung-myeon, Namsan-3ri, MT, 13–21.vi.2007, J.W.

Lee; 1♀, ditto, 26.iv–5.v.2009, J.W. Lee; 1♀, Gyeongsan- si, Dae-dong, Yeungnam Univ., 14.v.1988, J.Y. Cha; 1♂, ditto, 14.iv.1989; 1♀, ditto, 12–27.v.2007, J.W. Lee; 1♀, ditto, 17.x–18.xii.2008, J.W. Lee; 1♀, ditto, 6–20.v.2013, J.W. Lee; 1♀, ditto, Mungyeong-si, Maseong-myeon, Sangnae-ri, Mt. Baekhwasan, 16.viI.2013, J.K. Choi; 1♀, Cheongdo-gun, Unmun-myeon, Ssalbawi (U7), MT, 21.ix–

11.x.2012, J.W. Lee; GN: 1♀, Jinju-si, Gajwa-dong, Gyeongsang Univ., 8–14.iv.1989; 1♀, ditto, 2–9.vV.1989;

1♀, ditto, 6–12.v.1989; 1♀, ditto, 27.v–2.vi.1989, E.S.

Kim; JN: 1♀, Jeongeup-si, Ibammyeon, Deungcheon-ri, Naejangsan National Park, Ibamm Wetland, 21.vi.2005, coll. J.G. Han; GW: 1♂, Donghae-si, Samhwa-dong, Mureung valley, MT, 20–25.vi.2012, J.W. Lee; 1♀, Taebaek-si, Geumganggol, 19.v.1991, J.G. Kim.

Korean Record: Uchida, 1930; Kim, 1955; Yang et al., 2006; Choi et al., 2012.

Distribution: Korea, China, Japan, Russia, Holarctic.

Genus Syrphophilus Dasch 1964 꼬마등에살이뭉툭맵시벌속

Syrphophilus Dasch 1964: 59. TS: Bassus bizonarius Graven- horst.

29. Syrphophilus bizonarius (Gravenhorst 1829) 꼬마등에살이뭉툭맵시벌

Bassus bizonarius Gravenhorst, 1829: 1097. Type: lost.

Bassus cingulatus Holmgren, 1856: 396. Type: ♂, TL: Sweden;

TD: NR.

Bassus frontalis Brischke, 1878: 104. Type: ♂, TD: lost.

Bassus saginatus Provancher, 1879: 227. Type: ♂, TL:

Canada; TD: UL.

Zootrephes inconstans Davis, 1895: 23. Type: ♀, TL: U.S.A.;

TD: ANSP.

Homocidus iwatensis Uchida, 1930: 257. Type: ♀, TL: Japan;

TD: HU.

Homocidus satoi Uchida, 1930: 257. Type: ♂, TL: Japan;

TD: HU.

Material examined. [Korea] 1♀, Daegu-si Dalseo-gu, Daegok-dong, Daegu Arboretum (Alt. 88 m), 18.iv–

3.v.2012, coll. S.G. Kang; 1♀, ditto, 3–16.v.2012, coll. S.G.

Kang; 1♀, Ulsan-si, Sangbuk-myeon, Icheon-ri, 23.vi.1987, E.S. Kim; GB: 1♀, Cheongdo-gun, Gakbung-myeon, Namsan-3ri, MT, 13–21.vi.2007, J.W. Lee; 1♀, ditto, 12–

18.vii.2007, J.W. Lee; 1♀, Gyeongsan-si, Dae-dong, Yeungnam University, 22.iv.1985; 1♀, ditto, 22.iv.1987;

3♀, 23.iv.1987, E.S. Kim; 1♀, ditto, 5.v.1987, J.Y. Cha;

1♀, ditto, 8.v.1987; 1♀, ditto, 29.v.1987, J.W. Lee; 4♀, 28.iv.1988, J.Y. Cha; 2♀, ditto, 3.v.1988, J.W. Lee; 1♀, ditto, 16.vi.1988, E.J. Park; 1♀, ditto, 29.iv.1991, H.J.

Kang; 1♀, ditto, 22.v.1991, C.A. Kim; 1♀, ditto, 20.vi.992, M.N. Kim; 1♀, ditto, 11–24.v.2007, J.W. Lee; 1♀, ditto, 15–25.vi.2007, J.W. Lee; 1♀, ditto, 25.iv–6.v. 2013, J.W.

Lee; 1♀, ditto, 6–20.v.2013, J.W. Lee; GG: 1♀, Gwacheon- si, Makgye-dong, Mt. Cheonggye, 4.viI.1989, E.S. Kim;

Figures 25-30. Habitus of Diplazontinae spp. (25) Sussaba cognata albicoxa Thomson, 1890, (26) Sussaba sugiharai (Uchida, 1957), (27) Syrphoctonus laevis Brues, 1908, (28-♀, 29-♂) Syrphoctonus tarsatorius (Panzer 1809), (30) Syrphophilus bizonarius (Gravenhorst 1829).

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GN: 1♀, Geoje-si, Dongbu-myeon, Sannyang-ri, 28.vi.1990, S.M. Ryu; 2♀, Changnyeong-gun, Yueo-myeon, Daedae- ri, Uponeup, 27.iv.1985; 3♀, Yeongju-si, Sunheung- myeon, Sobaeksan National Park, Jukgye, 24.iv.2008, J.W.

Lee; GW: 1♀, Taebaek-si, Taebaeksan National Park, Geumganggol, 13.v.1991, J.K. Kim; 1♀, 14.v.1991, J.K.

Kim; 1♀, 17.v.1991, J.G. Kim; JB: 1♀, Namwon- si,Sannae-myeon, Mt. Jiri, Baemsagol, 11.vi–8.vii.2001, J.W. Lee; 2♀, Jeongeup-si, Ibammyeon, Deungcheon-ri, Wetland, 21.vi.2005, coll. J.G. Han.

Distribution: Korea, China, Japan, Russia, Holarctic.

Remark. Naejangsan National Park (New record), Jirisan National Park (New record), Sobaeksan National Park (New record), Taebaeksan National Park (New record).

30. Syrphophilus tricinctorius (Thunberg, 1822) 검정꼬마등에살이뭉툭맵시벌

Ichneumin tricinctorius Thunberg, 1822: 278. Type: ♂, TL:

U.S.A.; TD: USNM.

Bassus cinctus Gravenhorst, 1829: 327. Tyep: lost.

Bassus lateralis Gravenhorst, 1829: 342. Type: lost.

Bassus albicinctus Desvignes, 1862: 218. Type: ♂, TL:

United Kingdom; TD: NHM.

Bassus scapulatus Provancher, 1883: 11. Type: ♀, TL:

Canada; TD: UL.

Homocidus takaozanus Uchida, 1930: 259. Type: ♂, TL:

Japan; TD: HU.

Syrphophilus niveus Dasch, 1964: 70. Type: ♀, TL: U.S.A.;

TD: AEI.

Material examined. [Korea] CB: 4♂, Dannyang-gun, Gagok-myeon, Eouigok-ri, Mt. Sobaek, Huibang-valley, 12.v.1994, S.M. Ryu; 4♂, ditto, 3.v.1997, coll. D.K.

Chung; 1♂, ditto, 13.v.1997; 1♂, ditto, 22.v.1997, J.W.

Lee; 1♂, ditto, 23.v.1997, H.G. Park; 1♂, ditto, 24.v.1997, C.Y. Bae; GB: 2♂, Bonghwagun, Mt. Mirim, 4.v.1997, J.C. Jeong; 2♂, ditto, 4.v.1997, S.M. Ryu; 1♀, Yeongju-si, Punggi-eup, Sobaeksan National Park, Jungnyeong, 22.iv–

6.v.2009; GG: 1♂, Paju-si, Jeokseong-myeon, Mt. Gamak, 3.v.1998, S.M. Ryu; GW: 1♂, Pyeongchang-gun, Jinbu- myeon, Dongsan-ri75-6, Odaesan, 18.ix.1971, C.H. Kim;

3♀, Sokcho-si, Mt. Seolaksan, Misiryeong, Sinseongbong, 1.iv–24.vi.2010, J.C. Jeong; 1♂, Taebaek-si, Sodo-gol, Mt.

Taebaek, 23.vi.1989, J.Y. Cha; 3♂, ditto, 14.v.1992, J.W.

Cha; 5♂, ditto, 6.v.1999, J.W. Lee.

Remark. Odaesan National Park (New record), Seolaksan

National Park (New record), Sobaeksan National Park (New record), Taebaeksan National Park (New record).

Genus Woldstedtius Carlson, 1979 두점뭉툭맵시벌속 Woldstedtius Carlson, 1979: 332. TS: Bassus biguttatus

Gravenhorst.

31. Woldstedtius biguttatus (Gravenhorst, 1829, Bassus) 두점뭉툭맵시벌

Bassus rufipes Gravenhorst, 1829: 337. Type: lost.

Bassus confusus Woldstedt, 1874: 63. Type: female; TD: ZM.

Material examined. [Korea] 1♀, Daegu-si, Dalseo-gu, Daegok-dong, Daegu Arboretum, Alt. 88m, 20.vi–4.vii.2012, coll. S.G. Kang; GB: 1♀, Gyeongsan-si, Dae-dong, Yeungnam University, 21.iv–19.v.2004, coll. J.W. Lee; 1♀, ditto, 11–24.v.2007, coll. J.W. Lee; 1♀, ditto, 18.vi–

10.vii.2008; 3♀, Mt. Gayasan National Park, LT, 1–

2.vii.2000; GN: 1♀, Yeongju-si, Punggi-eup, Sobaeksan National Park, Jungnnyeong, site 99, 22.vi.2005, coll. K.B.

Kim; 7♀, ditto, 12.vi–23.vii.2008, coll. K.B. Kim and S.H.

Oh; 3♀, ditto, 12.vi–23.viii.2012, coll. S.H. Oh; 1♀, Sancheong-gun, Geumseo-myeon, Wangdeungjae, 29.vi–

31.vii.2001, J.W. Lee; 1♀, GG: Gapyeong-gun, Buk- myeon, Mt. Myeongji, 29.vi.1999, coll. J.C. Jeong; GW:

2♀, Donghae-si, Samhwa-dong, Mureung valley, MT, 21–

30.v.2005; 1♀, ditto, 16.x–25.xi.2005, coll. J.W. Lee; 1♀, ditto, 2–13.v.2006; same locality, 1♀, 22.vii–3.viiiI.2006, coll. J.W. Lee; 1♀, ditto, 2–15.v.2007, coll. J.W. Lee; 1♀, GW, Wonju-si, Baegun-myeon, Baegunsan, MT, 26.vi–

15.vii.2007; 2♀, ditto, 6–19.vi.2011, coll. H.Y. Han; JB:

2♀, Namwon-si, Sannae-myeon, Bamsadol, 11.vii–5.ix.2001;

2♀, Jeongeup-si, Ssangam-dong, Dapgok-ri, MT, site-3, 18.vi.2004, coll. J.W. Lee; 1♀, Jeongeup-si, Naejang-dong, Ansam, site-10, MT, 19.VI.2004; 1♀, Jeongeup-si, Naejang-dong, Geumseongyegok, Yonggul, Mt. Naejang, MT, site-14, 13–29.v.2007, J.W. Lee; 1♀, ditto, 8–

14.vi.2008, coll. J.W. Lee; 4♀, Jeongeup-si, Naejang-dong, Geumseong, MT, site-15, 20.vi.2005, coll. D.K. Chung.

Korean Record: Balueva & Lee, 2016b.

Distribution: Korea, Russia, Austria, Azerbaijan, Belgium, Bulgaria, Czech Republic, Czechoslovakia, Denmark, Finland, France, Germany, Hungary, Iran, Ireland, Latvia, Lithuania, Netherlands, Norway, Poland, Romania, Spain, Sweden, Switzerland, United Kingdom

Remark. Gayasan National Park (New record), Naejangsan National Park (New record), Jirisan National Park (New record), Sobaeksan National Park (New record).

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32. Woldstedtius corvinus Balueva & Lee, 2016 검정두점뭉툭맵시벌

Woldstedtius corvinus Balueva & Lee, 2016b: 390. Type: ♀, TL: Korea; TD: YNU.

Material examined. [Korea] JB: Jeoneup-si, Ibam-myeon, Deungcheon-ri, Wetland, MT, 21.v.2005, coll. J.G. Han;

1♀, Jeongeup-si, Ssangam-dong, Dapgok-ri, 19.v.2004;

2♀, ditto, 19.v.2005, coll. K.B. Kim; 1♀, ditto, 20.vi.2005;

1♀, Jeongeup-si, Naejang-dong, Geumseongyegok, Yonggul, Mt. Naejang, MT, site-14, 13–29.v.2007, coll. J.W. Lee;

1♀, same locality, 29.v–8.vi.2007, coll. J.W. Lee; 1♀, Jeoneup-si, Ibam-myeon, Deungcheon-ri, Wetland, MT, 19.v.2005, coll. J.G. Han; 3♀, ditto, 21.vi.2005, coll. J.G.

Han; GB: 1♀, Cheongdo-gun, Unmun-myeon, Munsu- seonwon (U2), MT, 12.v.2012, coll. J.W. Lee; 1♀, Yeong- yang-gun, Irwol-myeong, Mt. Irwolsan, 22.v–7.vi.2014, coll. H.Y. Han; GW: 2♀, Wonju-si, Heungeom-myeon, Maejiri 234, Yeonse Univ., Wonju campus, MT, 4–24.V.

2007, coll. J.W. Lee; 1♀, Hoengseong-gun, Gapcheon- myeon, Hadae-ri 594, 19.IV.2008, coll. J.W. Lee.

Distribution: Korea.

33. Woldstedtius citropectoralis (Schmiedeknecht 1926, Homocidus) 황두점뭉툭맵시벌

Homocidus citropectoralis Schmiedeknecht, 1926: 3412.

Type: ♂, TL: Germany; TD: ZMHU.

Bassus abdominator Bridgman, 1886: 336. Type: ♀, TL:

United Kingdom; TD: NCM.

Material examined. [Korea] CB: 1♀, Danyang-gun, Cheondong-ri, Mt. Sobaek, Bukbusa, MT, 14.x–24.xi.2005, coll. J.W. Lee; GW: 1♀, Wonju-si, Heungeom-myeon, Maejiri 234, Yeonse Univ., Wonju campus, MT, 21.v–

27.vi.2014, coll. H.Y. Han.

Distribution: Korea, Belgium, Canada, Czechoslovakia, Finland, Germany, Ireland, Lithuania, Netherlands, Poland, Spain, Sweden, Switzerland, U.S.A., United Kingdom.

34. Woldstedtius flavolineatus (Gravenhorst 1829) 노랑줄뭉툭맵시벌

Bassus flavolineatus Gravenhorst, 1829: 1097. Tyep: lost.

Bassus interruptus Holmgren, 1858: 359. Type: ♂, TL:

Sweden; TD: NR.

Bassus bimaculatus Holmgren, 1858: 360. Type: ♂, TL:

Sweden; TD: NR.

Bassus agilis Cresson, 1868: 111. Type: ♂, TL: U.S.A.; TD:

ANSP.

Bassus frontalis Cresson, 1868: 111. Type: ♀, TL: U.S.A.;

ANSP.

Mesoleius junctus Provancher, 1883: 10. Type: ♀, TL: Canada;

TD: UL.

Material examined.[Korea] GN: 1♀, Miryang-si, Sannae- myeon, Samyang-ri, Mt. Gajisan, 25.v.2001, coll. J.W. Lee.

Korean Record: Townes, Momoi & Townes 1965, Manukyan, 2007.

35. Woldstedtius holarcticus (Diller 1969, Syrphoctonus) 딜러두점뭉툭맵시벌

Syrphoctonus holarcticus Diller, 1969: 548. Type: ♀, TL:

U.S.A.; TD: ZSM.

Figures 31-36. Habitus of Diplazontinae spp. (31-♀, 32-♂) Syrphophilus tricinctorius (Thunberg 1822), (33) Woldstedtius biguttatus (Gravenhorst 1829, Bassus), (34) Woldstedtius corvinus Balueva & Lee, 2016, (35) Woldstedtius citropectoralis (Schmiedeknecht 1926, Homocidus), (36) Woldstedtius flavolineatus (Gravenhorst 1829).

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Material examined. [Korea] CN: 1♀, Seosan-si, haemimyeon, Daegok-ri 880, Hanseo Univ., MT, 20.ix–10.x.2009, coll. J.W. Lee; GW: 1♀, Wonju-si, Heungeom-myeon, Maejiri 234, Yeonse Univ., Wonju campus, MT, 21.v–

27.vi.2014, coll. H.Y. Han.

Distribution: Korea, Russia, Belgium, Finland, Germany, India, Poland, Switzerland, U.S.A.

36. Woldstedtius karafutensis (Uchida, 1957) 우치다두점뭉툭맵시벌

Homocidus karafutensis Uchida, 1957: 252. Type: female;

TD: HU.

Material examined. [Korea] 1♀, Daegu-si, Dalseo-gu, Daegok-dong, Daegu Arboretum, Alt.88mm, 30.viii–

5.ix.2012, coll. S.G. Kang; CB: 1♀, Jecheon-si, Naejang- dong, Malheorigol, MT, site-4, 21–24.iv.2007, coll. K.B.

Kim; 1♀, Jecheon-si, Deoksan-myeon, Worak-ri, Mt.

Worak National Park, 17–31.viii.2006; GB: 1♀, Yongin-si, Suji-gu, Mt. Gwanggyo (Alt. 231m), MT 8, 26.vii–

7.viii.2008, coll. J.O. Lim; 2♀, Yeongju-si, Punggy-eup, Sobaeksan National Park, Jungnyeong, 12–22.vi.2009;

GG: 2♀, Goyang-si, Deogyang-gu, Goyang-dong, MT, site-5, 11–23.vii.2007, coll. J.O. Lim; 1♀, Yangpyeong, Yongmun, Yeonsu, Mt. Youngmun (Alt. 324m), 16–30.vii.

2009, coll. J.O. Lim; 1♀, Gwangju-si, Docheok-myeon, Chugok-ri, Mt. Taehwa (Alt. 153m), 28.vi–14.vii.2008, coll. J.O. Lim; 1♀, Gwangju-si, Docheok-myeon, Chugok- ri, Mt. Taehwa (Alt. 219m), 8.v–7.vi.2007, coll. J.O. Lim;

GN: 1♀, Danyang-gun, Danyang-eup, Cheondong, 12.vi–

22.vii.2008, coll. J.W. Lee; GW: 1♀, Chuncheon-si, Dong- myeon, Jinae-ri, 29.viii–5.ixX.2005, coll. S.J. Jang; 2♀, Hongcheon-gun, Hongcheon-eup, Jangjeonpyeong-1ri, Geomdungae village, MT, 6–26.vii.2007, coll. J.W. Lee;

1♀, Hongcheonh-gun, Bukbang-myeon, Gwangwon Prov.

Environment Research Park, 1–17.vi.2013, coll. S.J. Jang;

1♀, Wonju-si, Heungeom-myeon, Maejiri, Yeonse Univ., Wonju campus, MT, 4–24.V.2007; 1♀, ditto, 23.vi–

8.vii.2009, coll. H.Y. Han; 1♀, ditto, 2.ix–20.x.2009, coll.

H.Y. Han; JB: 1♂, Jeongeup-si, Naejang-dong, Malheorigol, MT, site-4, 21–24.iv.2007, coll. K.B. Kim.

Distribution. Korea, Japan, Russia.

Remark. Naejangsan National Park (New record), Sobaeksan National Park (New record), Woraksan National Park (New record).

37. Woldstedtius nigrosignatus Balueva & Lee, 2016 검정무늬두점뭉툭맵시벌

Woldstedtius nigrosignatus Balueva & Lee, 2016b: 394.

Type: ♀, TL: Korea; TD: YNU.

Material examined. [Korea] GB: 1♀, Gyeongsan-si, Dae- dong, Yeungnam University, 6–20.v.2013, coll. J.W. Lee;

JB: 1♀, Wanju-gun, Dongsang-myeon, Daea Arboretum (Alt. 174m), 1–16.x.2012, coll. J.M. Park; 1♀, Jeongeup- si, Ssangam-dong, Dapgok-ri, MT, 19.v.2005, coll. K.B.

Kim; 1♀, Jeongeup-si, Naejang-dong, Mt. Naejang, Naejangsa (BL), 11.v.2007, coll. J.C. Jeong; 1♀, Jeongeup- si, Jangseong-gun, Bukha-myeon, 21.vi.2005, coll. K.B.

Kim.

38. Woldstedtius pallidus Balueva & Lee, 2016 연붉은두점뭉툭맵시벌

Figures 37-41. Habitus of Diplazontinae spp. (37) Woldstedtius holarcticus (Diller 1969, Syrphoctonus), (38) Woldstedtius karafutensis (Uchida 1957), (39) Woldstedtius nigrosignatus Balueva & Lee, 2016, (40) Woldstedtius pallidus Balueva & Lee, 2016, (41) Woldstedtius yokohamensis (Uchida 1930).

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Woldstedtius pallidus Balueva & Lee, 2016b: 395. Type: ♀, TL: Korea; TD: YNU.

Material examined. [Korea] GB: 1♀, Cheongdo-gun, Gakbuk-myeon, Namsan-3ri, MT, 13–26.iv.2009; 1♀, Gyeongsan-si, Dae-dong, Yeungnam University, 6–20.v.2013.

coll. J.W. Lee; GW: 1♀, Donghae-si, Samhwa-dong, Mureung valley, MT, 29.iv–26.v.2007; 1♀, Wonju-si, Heungeom-myeon, Maejiri 234, Yeonse Univ., Wonju campus, MT, 4–24.v.2007, coll. J.W. Lee; JB: 1♀, Jeongeup- si, Ssangam-dong, Dapgok-ri, MT, 19.v.2005, coll. K.B.

Kim; 1♀, Jeongeup-si, Ibae-myeon, Deungcheon-ri, Wetland, MT, 21.vi.2005, coll. J.H. Han.

39. Woldstedtius yokohamensis (Uchida, 1930) 검정다리두점뭉툭맵시벌

Homocidus yokohamensis Uchida, 1930: 258. Type: ♀; TD:

HU.

Material examined.[Korea]CB: 1♀, Chungju-si, Sangmo- myeon, Samunri, Sinseon Falls, Mt. Worak National Park, s-72, 21.v.2006, M.K. Yun; GB: 1♀, Cheongdo-gun, Unmun-myeon, Mt. Unmun, MT, 6 –19.vi.2008, coll. J.W.

Lee; GW: 1♀, Donghae-si, Samhwa-dong, Mureung valley, MT, 21.iv–6.v.2007, coll. J.W. Lee; JB: 1♀, Jeongeup-si, Naejang-dong, Yeonja-sinseongbong 20.v.2004, coll. K.B. Kim; 1♀, Jeongeup-si, Ibae-myeon, Deungcheon- ri, Wetland, MT, 21.vi.2005, coll. J.H. Han.

Remark. Naejangsan National Park (New record), Woraksan National Park (New record).

Key to the genera of South Korean Diplazontinae 1. Notaulus distinct on front part of mesoscutum, also the

apical edge of tergites 3-7 not concave. Areolet absent

··· 2 – Notaulus absent or indicate by weak rugosity, if present (in Campocraspedon), in which genus the apical edge of tergites 3–7 are concave. Areolet present or absent ··· 3 2. Spiracle of third tergite on epipleurum. Face polished, sometimes weakly punctate, with two vertical impressions between clypeus and antennal socket. Flagellum of male with tyloids Tergites 1–3, each without a postmedian transverse impression ··· Promethes Förster – Spiracle of third tergite on tergite. Face mat, lacking two vertical impressions between clypeus and antennal socket.

Flagellum of male without tyloids. Tergites 1–3, each with a postmedian transverse impression. Hind tibia black with white band medially ··· Diplazon Nees 3. Face polished, or if mat then spiracle of third tergite on epipleurum. Areolet absent ··· 4 – Face mat; spiracle of third abdominal segment on tergite.

Areolet present or absent ··· 6 4. Spiracle of propodeum and first tergite enlarged, with pale

raised margin. Spiracle of second tergite on tergite.

··· Bioblapsis Förster – Spiracle of propodeum and first tergite not enlarged.

Spiracle of second tergite on epipleurum ··· 5 5. Metasoma in female distinctly elongate and strongly compressed, hind edges of tergites 3–8 concave. Face usually lacking two vertical impressions between clypeus and antennal socket ··· Eurytyloides Nakanishi – Metasoma in female not elongate and not compressed, if compressed then hind edges of tergites 3–6 not concave.

Face usually with two vertical impressions between clypeus and antennal socket. ···Sussaba Cameron 6. Tergites 3–7 concave on hind edge. Metasoma elongate, usually projecting noticeably beyond wings. Second tergite (often third and fourth) longer than wide. ··· 7 – Tergites 3–7 truncate or slightly convex on hind edge.

Metasoma usually not elongate. Second tergite usually wider than long, or as long as wide. ··· 8 7. Areolet absent. Metasoma of female depressed.

··· Campocraspedon Uchida – Areolet present. Metasoma of female compressed.

··· Phthorima Förster 8. Apex of clypeus with a median notch, impressed near apical edge. Areolet often present. ··· 9 – Apex of clypeus roundly lobed apically or truncate, but not impressed near apical edge (if apex of clypeus notched, then propodeum strongly carinate). Areolet absent. ··· 14 9. First tergite and basal 0.25 of second tergite usually with strong dorsal longitudinal carinae. Areolet usually present.

Hind tibia black with white basal part.

··· Enizemum Förster – First and second tergites usually without distinct dorsal longitudinal carinae. Areolet present or absent. Hind legs different colors, but not black with white basal part.

··· 10 10. Prepectal carina usually broadly interrupted ventrally behind fore coxae. Clypeus usually flat or with apico- lateral impressions. ··· Syrphoctonus Förster – Prepectal carina complete ventrally. Clypeus with preapical impression. Female with mesopleuron usually entirely black. ··· Homotropus Förster

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11. Propodeum without carinae. Dorsal longitudinal carinae of first tergite absent or weak. Hind tibia black white at base. Clypeus with apical margin thin, with a basal elevation that makes it flat or even convex in profile, relatively broad and short, with median vertical groove.

··· Woldstedtius Carlson – Propodeum with well-developed carinae. Median dorsal carinae of first tergite strong. ··· Syrphophilus Förster

Acknowledgments

This work was supported by a grant from the National Institute of Biological Resources (NIBR), funded by Graduate Program for the Undiscovered Taxa of Korea (NIBR201722202).

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