The Permian ostracods described here are from the top of the Episkopi Formation (Dorashamian) (USNM locality on Hydra Island, Greece (Sohn and Kornicker, 1998). We thank Elizabeth Harrison-Nelson (NMNH) for general assistance, Molly Ryan (NMNH) for letter plates and Jack Korytowski (Smithsonian Institution Press) for final editing and preparation of the manuscript for publication.The known range of Paleomyodocopina is Devonian to Permian, whereas the known range of Myodocopine is Silurian to Recent.
The absence of post-Paleozoic Paleomyodocopina is herein interpreted as an indication that the suborder became extinct at the end of the Permian. Most synapomorphies that define extant members are not shell characters, but the fossil shells included in Myodocopa have shell characters like those of the extant Myodocopa. Key to the Superfamilies of the Paleomyodocopina (The key includes characters from most members of each superfamily).
Due to the pronounced rostrum of Sulcuna lepus, the type species, the genus is preferably included in the Cypridinelliformidae. COMPARISONS.—The carapace of Cypridinelliforma differs from Cypridellina and Sulcuna in that the anteroventral arch does not extend beyond the tip of the rostrum; differs from Cypridella and Sulcuna in lacking a nuchal furrow. The cross ribs forming the reticulations at the posterior end of the carapace are visible only in the holotype (Fig. la.c).
COMPARISONS.—The new species differs from Cypridinelliforma emmaensis in having a straight posterodorsal margin. The lack of the node places that genus in the Myodocopina, rather than in the Paleomyodocopina. Shells of the Giganto-cypridinini longer than 4 mm. COMPOSITION.—This tribe includes about 21 Holocene genera and three genera known only as fossils.
Because many Paleozoic species have been incorrectly referred to Cypridina (e.g., Jones and Kirkby, 1874), a discussion of the genus is included. DISCUSSION – A fundamental problem with this genus is the inadequate description of the species. 20 SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY the absence of the incisor accounted for error in observation of.
Poulsen mentioned the difficulty in referring species to the genus Cypridina due to the very incomplete description of C.
21 these forms. Their shape and outline is comparable with Devo-
COMMENTS. The surface ridges of many genera of "fingerprint" ostracods occurring in the Entomozoacea of Sylvester-Bradley (1961:Q388) resemble those of Asteropella kalkei Kornicker, 1986b (fig. 56). These genera are Bertillonella Stewart and Hendrix, 1945; EntomoprimitiaKummeww referred the families within which these genera are usually placed to the Halocypridina and Cladocopina. We have similar reservations about referring these genera to the Myodocopina, and they will not be considered further here.
Key to the Genus of Asteropteroninae (Key only applies to taxa reported from the Paleozoic and/or Mesozoic). COMPOSITION.—This genus includes Triadocypris spitzbergensis Weitschat, 1983a, from the Triassic of Spitzbergen and a new species from the Permian. DIAGNOSIS.-Carapace oval in lateral outline, small rostrum with small incisures; inner lamella narrow and calcified; with numerous radial pore channels.
Scar pattern in the adductor muscles consisting of two diagonal rows of scars (based on the diagnosis of the shield in Weitschat, 1983a:314). DISCUSSION. The shape of the carapace, the small rostrum, the lack of a deep incisure and the convex dorsal margin of the left valve visible in dorsal view (Figure 15b) of T pax are very similar to carapaces of species of the extant genus Actinoseta jonesi (Kornicker, 1981, pis. 62a, 63a,e); this suggests that the genus Triadocypris should be referred to the Asteropteroninae. It is not known whether fossil specimens of Triadocypris have the posterodorsal tooth-and-socket structures present on recent species of Actinoseta (see Kornicker 1981, pis. 57e,f, 58a,b).
1, 2 of Triadocypris spitzbergensis in Weitschat (1983b: 127) stated that both illustrations are from specimen number. GPIHM referred this species to a new family, Triadocypridinae, but because this is based on soft parts of the unique fossil, we do not think it is practical to use the new family in identifying fossils. DESCRIPTION. Carapace oval in lateral view with slightly protruding rostrum and with small incisure just ventral to halfway up the valve (Figure I5a,d). In dorsal and ventral views, the carapace is widest at % valve length measured from the anterior end of the valve; shield tapered anteriorly to widest part.
DIAGNOSIS.—Carapace oval in lateral view, with short rostrum and rounded posteriorly without caudal process. Paratype, USNM 496728, complete specimen (valves crooked), length unknown: d, lv; e, dv, anterior to right;/ vv, anterior to right. 66a,b,d, 67a) suggested that the genus be referred to the Asteropteroninae. The shells of known species of Siveterella lack the posterodorsal tooth-and-socket structures present on recent species of Actinoseta (Figures 17a, 20a,c).
HOLOTYPE.—USNM 496730, RV
29 Eocypridina radiata (Jones and Kirkby, 1874)
A low swelling in anterodorsal quadrant of USNM 496714 (Figure 2la,b) is tentatively interpreted as not a node similar to that in the Paleomyodocopina.). REMARKS.—Differences in the width and curvature of the rostrum of the Permian specimens suggest that several species may be included in this taxon. On a new genus and species of Cypridinacea (Ostracoda) from the Upper Carboniferous of the Netherlands.
Monograph of the Marine and Freshwater Ostracoda of the North Atlantic and Northwest Europe. Travaux des Laboratoires de Geologie de la Faculte des Sciences de Lyon, new series of plates 1-3. A myodocopid ostracod with preserved appendages from the Upper Jurassic of the Volga River region (USSR).
In John Young and James Armstrong, editors, On the Carboniferous fossils of the west of Scotland, their range and vertical distribution with a general catalog of the fossils and their mode of occurrence, and an index to the principal localities. Ostracods of the Upper Pennsylvanian and Lower Permian Strata of Kansas, III: Bairdiidae (Concluded), Cytherellidae, Cypri- dinidae, Entomoconchidae, Cytheridae, and Cypridae.
