An Annotated Checklist on the Introduced Aquatic Fauna of Two River Systems on

An Annotated Checklist on the Introduced

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INTRODUCTION

F

›Žœ‘ ŠŽ›ȱ šžŠ’Œȱ —ŸŠœ’ŸŽȱ ™ŽŒ’Žœȱ ǻǼȱ ‘ŠŸŽȱ ‹ŽŒ˜–Žȱ Šȱ

–Š“˜›ȱ •˜‹Š•ȱ Œ˜—ŒŽ›—ȱ Šœȱ ’—›˜žŒŽȱ œ™ŽŒ’Žœȱ ‹ŽŒ˜–Žȱ ’—ŸŠŽ›œȱ

Š—ȱ Œ˜–™Ž’˜›œȱ ˜ȱ •˜ŒŠ•ȱ ™˜™ž•Š’˜—œȱ Š—ȱ ŒŠžœŽȱ ŽŒ˜œ¢œŽ–ȱ

ŽœŠ‹’•’£Š’˜—ȱ ǻ¢Ž›œǰȱ ŘŖŗŖǼǯȱ —›˜žŒŽȱ œ™ŽŒ’Žœȱ ‘›ŽŠŽ—ȱ Š—ȱ

˜Ž—ȱŒŠžœŽȱ‘ŽȱŽ–’œŽȱ˜ȱ—Š’ŸŽȱœ™ŽŒ’ŽœǰȱŽœ™ŽŒ’Š••¢ȱ‘Žȱ›Š›Žȱœ™ŽŒ’Žœȱ

Š—ȱ œ™ŽŒ’Žœȱ Ž—Š—Ž›Žȱ ‹¢ȱ ˜‘Ž›ȱ ‘›ŽŠŽ—’—ȱ ŠŒ˜›œȱ ǻ˜‹•Ž—£ǰȱ ŗşşŖDzȱ˜¢•ŽȱǭȱŽ’¢ǰȱŗşşŘDzȱ•ŠŒ”ǰȱŗşşśDzȱ–’‘ȱǭȱž’—ǰȱŗşşŜǼǯȱȱ—ȱ

Š’’˜—ȱ˜ȱ‘’œǰȱœ˜–ŽȱŠ•’Ž—ȱ’œ‘ȱœ™ŽŒ’Žœȱ‘ŠŸŽȱ‹ŽŽ—ȱ”—˜ —ȱ˜ȱ’—ŽŒȱ

•˜ŒŠ•ȱ’œ‘ȱœ™ŽŒ’Žœȱ ’‘ȱ‘Ž’›ȱ™Š›Šœ’ŽœȱŠœȱ‘Žȱ‘˜œȬ™Š›Šœ’ŽȱŒ˜–™•Ž¡ȱ

’œȱ Ž¡Ž—Žȱ ˜ȱ ‘Žȱ •˜ŒŠ•ȱ œ™ŽŒ’Žœȱ ŒŠžœ’—ȱ ‘Žȱ —Š’ŸŽȱ ™˜™ž•Š’˜—ȱ

˜ȱ ŽŒ•’—Žȱ ǻ Š••’ȱ Žȱ Š•ǯǰȱ ŘŖŖśǼǯȱ —ȱ Ž›–œȱ ˜ȱ Œ˜—›’‹ž’˜—ȱ ˜ȱ •˜œœȱ ˜ȱ

‹’˜’ŸŽ›œ’¢ȱ’—ȱ›Žœ‘ ŠŽ›ȱŽŒ˜œ¢œŽ–œǰȱȱ’œȱŒ˜—œ’Ž›ŽȱœŽŒ˜—ȱ

˜ȱ ‘Š‹’Šȱ Žœ›žŒ’˜—ȱ ǻŠœŠ•ǰȱ ž—Šǰȱ ›˜ŽœŽǰȱ Š’••¢ǰȱ Š—ŠŒ’˜ǰȱ ǭȱ

‹Š¢Š—’ǰȱŘŖŖŝǼǯ

—ȱ‘Žȱ‘’•’™™’—ŽœǰȱȱŠ›Žȱ”—˜ —ȱ˜ȱŒ˜–™ŽŽȱ’—ȱŽ›–œȱ˜ȱ˜˜ȱ

Š—ȱ œ™ŠŒŽȱ  ’‘ȱ ‘Žȱ —Š’ŸŽȱ œ™ŽŒ’Žœȱ ǻŠ—˜œȬ˜›“Šǰȱ ŘŖŖŘDzȱ ‘ŠŸŽ£ǰȱ

Žȱ•ŠȱŠ£ǰȱŠ—˜‘Š›ǰȱŠž•Š¢Š—ǰȱǭȱŠ›Š—Š—ǰȱŘŖŖŜǼǯȱ‹˜žȱŗŞŗȱ

œ™ŽŒ’Žœȱ ǻŘŞȱ Š–’•’ŽœǼȱ Š›Žȱ ”—˜ —ȱ ˜ȱ ‘ŠŸŽȱ ‹ŽŽ—ȱ ’—›˜žŒŽȱ ’—˜ȱ

‘Žȱ Œ˜ž—›¢ȱ ǻŠŠžŠ—ǰȱ ŘŖŖŝǼǯȱ ‘ŽœŽȱ ’—Œ•žŽȱ ’œ‘ȱ ǻŒ’›ŒŠȱ şřƖǼǰȱ

–˜••žœ”œȱǻŘǯŜŝƖǼȱŠ—ȱœ˜–ŽȱŒ›žœŠŒŽŠ—œǰȱ›˜œǰȱŠ—ȱž›•ŽœȱǻŒ’›ŒŠȱ ŚƖǼǯȱŠœŠ•ȱŽȱŠ•ǯȱǻŘŖŖŝǼȱŽœ’–ŠŽȱ‘Šȱ˜ȱ‘Žȱŗśşȱœ™ŽŒ’Žœȱ˜ȱŽ¡˜’Œȱ

’œ‘Žœȱ ’—›˜žŒŽȱ ’—ȱ ‘Žȱ Œ˜ž—›¢ǰȱ Šȱ •ŽŠœȱ řşȱ œ™ŽŒ’Žœȱ  Ž›Žȱ Š‹•Žȱ

˜ȱ ŽœŠ‹•’œ‘ȱ ’—ȱ ‘Žȱ  ’•ǯȱ ‘ŽœŽȱ Ž¡˜’Œȱ ˜›Š—’œ–œȱ  Ž›Žȱ ™›’–Š›’•¢ȱ

žœŽȱ Šœȱ ˜›—Š–Ž—Š•ȱ ˜›ȱ ŠšžŠ›’ž–ȱ ™Žœȱ ǻŝŜƖǼǰȱ Šœȱ ˜˜ȱ ǻŘŗƖǼȱ Š—ȱ

Šœȱ ‹’˜•˜’ŒŠ•ȱ Œ˜—›˜•ȱ ǻŘƖǼȱ ǻŠŠžŠ—ǰȱ ŘŖŖŝǼǯȱ —ȱ Š•‘˜ž‘ȱ –˜œȱ

˜ȱ ‘ŽœŽȱ œ™ŽŒ’Žœȱ  Ž›Žȱ ‹›˜ž‘ȱ ’—ȱ ˜›ȱ ‘Žȱ ˜›—Š–Ž—Š•ȱ ’—žœ›¢ȱ ǻŠšžŠ›’ž–ȱ Š—ȱ ™Žȱ ›ŠŽǼǰȱ ‘Ž’›ȱ ’—Œ’Ž—Š•ȱ ›Ž•ŽŠœŽȱ ’—ȱ ‘Žȱ  ’•ȱ

Š™™ŽŠ›œȱ•Žœœȱœ’—’’ŒŠ—ȱ‘Š—ȱ‘ŽȱŸ˜•ž–Žȱ›Ž•ŽŠœŽȱ‹¢ȱ‘Žȱ’œ‘Ž›’Žœȱ

Š—ȱ ŠšžŠŒž•ž›Žȱ ǻž••’—ǰȱ Š•˜–Š›Žœǰȱ ŠœŠ•ǰȱ Ž¢ǰȱ ǭȱ Šž•¢ǰȱ ŗşşŝDzȱ

ŠœŠ•ȱŽȱŠ•ǯǰȱŘŖŖŝǼǯȱ—›˜žŒ’˜—ȱ˜ȱŽ¡˜’Œœǰȱ™Š›’Œž•Š›•¢ȱ’œ‘ȱœ™ŽŒ’Žœǰȱ  Šœȱ ‘’‘Žœȱ ž›’—ȱ ‘Žȱ ŗşŝŖœȱ Šœȱ Šȱ ›Žœž•ȱ ˜ȱ ‘Žȱ ˜ŸŽ›—–Ž—Ȃœȱ

ŠŽ–™ȱ ˜ȱ Šž–Ž—ȱ ˜˜ȱ Š—ȱ ’œ‘Ž›’Žœȱ ™›˜žŒ’˜—ȱ ǻŠŠžŠ—ǰȱ ŘŖŖŝǼǯ

˜Œž–Ž—Š’˜—ȱ ˜ȱ ’—›˜žŒŽȱ Š•’Ž—ȱ œ™ŽŒ’Žœȱ ’—ȱ ‘Žȱ ’œŠ¢Šœǰȱ

‘’•’™™’—Žœȱ ’œȱ ’—ŠŽšžŠŽǯȱ ‘’œȱ ™Š™Ž›ȱ ŠŽ–™œȱ ˜ȱ Šž–Ž—ȱ

’—˜›–Š’˜—ȱ˜—ȱȱ›˜–ȱ ˜ȱ–Š“˜›ȱ›’ŸŽ›ȱœ¢œŽ–œǰȱ‘ŽȱŠ˜ȱ’ŸŽ›ȱ

’—ȱŽ›˜œȱ’œ•Š—ȱŠ—ȱ‘ŽȱŠ•Šž›ȱ’ŸŽ›ȱ’—ȱŠ—Š¢ȱ’œ•Š—ǯȱȱ J. LINAUGO, E. ALCALA, A. BUCOL, C.C. MENES, ET AL

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MATERIALS AND METHODS Study sites

‘Žȱ œž¢ȱ ˜ŒžœŽȱ ˜—ȱ  ˜ȱ –Š“˜›ȱ ›’ŸŽ›ȱ œ¢œŽ–œǰȱ ‘Žȱ Š˜ȱ ’ŸŽ›ȱ ’—ȱ

Ž›˜œȱŠ—ȱŠ•Šž›ȱ’ŸŽ›ȱ’—ȱŠ—Š¢ǰȱ‘’•’™™’—Žœȱǻ’ž›ŽȱŗǼǯȱŠ˜ȱ’ŸŽ›ȱ

’œȱŠȱŝŜȬ”–ȱ•˜—ȱ›’ŸŽ›ȱœ¢œŽ–ȱ‘Šȱ›ŽŒŽ’ŸŽœȱœŽŸŽ›Š•ȱ›’‹žŠ›’ŽœȱŒ˜–’—ȱ

›˜–ȱ ˜ȱ•Š›Žȱ–˜ž—Š’—ȱ›Š—Žœȯ˜ž—ȱŠ—•Š˜—ȱ’—ȱ‘Žȱœ˜ž‘ŽŠœȱ

Š—ȱ˜›‘ȱŽ›˜œȱ˜ž—Š’—ȱŠ—Žȱ’—ȱ‘Žȱ—˜›‘ŽŠœǯȱȱ›Š’—œȱ’—˜ȱ ž’–Š›Šœȱ›Š’ǰȱ•˜ŒŠŽȱŠȱ‘Žȱ—˜›‘ Žœȱ™˜›’˜—ȱ˜ȱŽ›˜œȱœ•Š—ǯȱ

‘Žȱœž¢ȱœ’Žœȱ’—ȱŠ˜ȱ’ŸŽ›ȱŠ—ȱ‘Ž’›ȱŒ˜››Žœ™˜—’—ȱ‹ŠœŽȱ›ŽŽ›Ž—ŒŽȱ

™˜’—œǰȱœŠ›’—ȱ›˜–ȱ‘Žȱž™™Ž›ȱœ›ŽŒ‘ȱȱŠ—ȱ˜’—ȱ˜ —œ›ŽŠ–ȱŠ›ŽDZȱ

’’˜ȱŠ–‹Š—Š˜ǰȱŠ›Š—Š¢ȱž–Š•’œ”’œǰȱž—’Œ’™Š•’¢ȱ˜ȱ˜—ȱŠ•ŸŠ˜›ȱ

Ž—Ž’Œ˜ȱ ǻŗŖǚȱ řŗǯŝŜŖȇȱ ǰȱ ŗŘřǚȱ ŗŘǯŞśŚȇȱ Ǽǰȱ ž›˜”ȱ ›‹Š—ǰȱ Š›Š—Š¢ȱ

˜™Ž£ȬŠŽ—Šǰȱž—’Œ’™Š•’¢ȱ˜ȱž›Œ’ŠȱǻŗŖǚȱřřǯŚśŖȇȱǰȱŗŘřǚȱŖŚǯŗŚŖȇȱǼǰȱȱ

Š›Š—Š¢ȱŠ–œ’Žǰȱž—’Œ’™Š•’¢ȱ˜ȱž›Œ’ŠȱǻŗŖǚȱřřǯŘřŚȇȱǰȱŗŘřǚȱŖŘǯŗŚřȇȱ ǼǰȱŠ—ȱ‹Ž ŽŽ—ȱŠ›Š—Š¢ȱŠȬŠœŠ—ǰȱŠ˜ȱ’¢ȱŠ—ȱŠ›Š—Š¢œȱŠ™˜—ȱ

Š—ȱ‹Š¢ǰȱž—’Œ’™Š•’¢ȱ˜ȱž•ž™Š—Š—ȱǻŗŖǚȱřŗǯŘŖŚȇȱǰȱŗŘŘǚȱśŖǯŘŜŖȇȱ Ǽǯȱ‘ŽȱŽ•ŽŸŠ’˜—ȱ˜ȱ‘Žȱž™™Ž›–˜œȱœŠ’˜—ȱǻŠ’˜—ȱŗǰȱž–Š•’œ”’œǼȱ’œȱ

Š‹˜žȱřŘŖȱ–ŽŽ›œȱŠ‹˜ŸŽȱœŽŠȱ•ŽŸŽ•ȱŠ—ȱ’œȱŠ‹˜žȱŜŖȱ”–ȱŠ Š¢ȱ›˜–ȱ‘Žȱ

•˜ Ž›–˜œȱœŠ’˜—ȱǻŠ’˜—ȱŚǰȱŠŸŠ—ȬŠŠœŠ—Ǽǯȱ

‘Žȱœž¢ȱœ’Žœȱ’—ȱŠ•Šž›ȱ’ŸŽ›ȱŠ—ȱ‘Ž’›ȱŒ˜››Žœ™˜—’—ȱŒ˜˜›’—ŠŽœȱ

˜›ȱ‹ŠœŽȱ›ŽŽ›Ž—ŒŽȱ™˜’—œǰȱœŠ›’—ȱ›˜–ȱ‘Žȱž™™Ž›ȱœ›ŽŒ‘ȱ˜ȱ‘Žȱ›’ŸŽ›ǰȱ

Š›Žȱ‘Žȱž™™Ž›ȱŠ•’—˜ȱŠ’˜—ȱǻŗŗǚȱŖşǯŘŖŞȇȱǰȱŗŘŘǚȱŘşǯŖŗŘȇȱǼǰȱ‘Žȱ–’Ȭ

œ›ŽŠ–ȱ ˜›˜‹˜›˜Ȭž•Š‹˜Ȭž’Š—ȱ Š’˜—ȱ Šȱ ’—•Žȱ ǻŗŗǚȱ ŖŗǯŝŖŖȇȱ ǰȱ ŗŘŘǚȱřşǯŚřřȇȱǼǰȱŠ—ȱ‘Žȱ•˜ Ž›–˜œȱŽœžŠ›’—ŽȱœŠ’˜—ȱŠȱŽŠ—ŽœȱǻŗŖǚȱ

Figure 1. Satellite images: Black-and-white satellite image of Bago River showing location of the four stations and inset map of the Philippines (upper left). Black-and-white satellite view of upper Jalaur River showing the locations of the upper Calinog, mid-Dingle, and Leganes stations (upper right). Colored satellite image showing the respective positions of the Jalaur and Bago River mouths on the islands of Panay and Negros (lower large map).

INTRODUCED AQUATIC FAUNA OF TWO RIVER SYSTEMS

141 ŚŝǯřŚřȇȱǰȱŗŘŘǚȱřŞǯŗŚśȇȱǼǯȱ‘Žȱž™™Ž›–˜œȱœŠ’˜—ȱ‘ŠœȱŠ—ȱŽ•ŽŸŠ’˜—ȱ˜ȱ

Š‹˜žȱŞŖȱ–ŽŽ›œȱŠ‹˜ŸŽȱœŽŠȱ•ŽŸŽ•ȱŠ—ȱ’œȱŠ‹˜žȱşŖȱ”–ȱŠ Š¢ȱ›˜–ȱ‘Žȱ

•˜ Ž›–˜œȱ œŠ’˜—ȱ ǻ˜ž›ŒŽDZȱ ˜˜•Žȱ Š›‘ǰȱ ŘŖŗŖǼǯȱ Š•Šž›ȱ ‹Šœ’—ȱ ‘Šœȱ Šȱ

˜Š•ȱŠ›ŽŠȱ˜ȱŗǰŞŘŝȱœšžŠ›Žȱ”’•˜–ŽŽ›œȱǻ ˜—£Š•ŽœǰȱŗşŞŚǼǯȱ‘Žȱ›’ŸŽ›ȱ›Š’—œȱ

’—˜ȱ‘Žȱ•˜’•˜ȱ›Š’ǯ

Š–™•’—ȱ Šœȱ˜—Žȱžœ’—ȱŠȱŸŠ›’Ž¢ȱ˜ȱŽŒ‘—’šžŽœǯȱ‘ŽœŽȱ’—Œ•žŽȱ

‘Žȱ •˜ŒŠ•ȱ ꜑’—ȱ ŽŠ›ȱ ŒŠ••Žȱ taonȱ Š—ȱ ꜑’—ȱ ŽŒ‘—’šžŽȱ ŒŠ••Žȱ pahubasȱǻ’ž›ŽȱŘǼǯȱ‘Žȱ˜›–Ž›ȱ’œȱŠ—ȱ’—’Ž—˜žœȱ›Š™ȱžœŽȱ˜ȱŒŠ™ž›Žȱ

–ŠŒ›˜‹Ž—‘˜œȱ ’—ȱ Š˜ȱ ’ŸŽ›ȱ  ‘’•Žȱ ‘Žȱ •ŠĴŽ›ȱ ’œȱ Šȱ ꜑’—ȱ ŽŒ‘—’šžŽȱ

Ž–™•˜¢Žȱ ’—ȱ Š•Šž›ȱ ’ŸŽ›ȱ ‘Šȱ ’—Ÿ˜•ŸŽœȱ ›¢’—ȱ ž™ȱ ›’ŸŽ›ȱ œŽŒ’˜—œǯȱ

—˜‘Ž›ȱ •˜ŒŠ•ȱ –Ž‘˜ȱ žœŽȱ ˜›ȱ Œ˜••ŽŒ’—ȱ ꜑ǰȱ –ŠŒ›˜’—ŸŽ›Ž‹›ŠŽœǰȱ

Š—ȱ˜‘Ž›ȱ›’ŸŽ›’—ŽȱŸŽ›Ž‹›ŠŽœȱǻŽǯǯǰȱ›˜œȱŠ—ȱž›•ŽǼȱ Šœȱpanulo or

•ŽŠ—’—ȱ–Ž‘˜ǯȱŠ–™•’—ȱ Šœȱ˜—ŽȱŠ•˜—ȱ‘Žȱ–Š’—ȱ›’ŸŽ›ȱŒ‘Š——Ž•ǰȱ’—ȱ

›’‹žŠ›¢ȱœ›ŽŠ–œȱ“˜’—’—ȱ‘Žȱ–Š’—ȱŒ‘Š——Ž•ǰȱŠ—ȱ’—ȱœ–Š••ȱ™˜˜•œȱŠ•˜—ȱ

‘Žȱ›’ŸŽ›ȱ‹Š—”œǯȱŽ›’Žœȱ˜ȱœŠ–™•’—ȱ›ž—œȱ Ž›ŽȱŒ˜—žŒŽȱž›’—ȱ‘Žȱ  ŽȱŠ—ȱ›¢ȱœŽŠœ˜—œȱ˜ȱ˜™’–’£ŽȱŒ˜••ŽŒ’˜—ȱŠ—ȱ˜ȱŒ˜–™Š›ŽȱœŽŠœ˜—Š•ȱ

ŸŠ›’Š’˜—ǯ

Figure 2. PahubasDÀVKLQJWHFKQLTXHLQYROYLQJVWUHDPGU\LQJ$SRUWLRQRIWKLVULYHU-DODXU River in Calinog station) is being prepared for drying using rocks. The rock barrier functions to UHFKDQQHOWKHVWUHDPÁRZWRWKHULJKWODWHUFDXVLQJWKHSRUWLRQRQWKHOHIWVLGHWRGU\XS7KH GU\SRUWLRQRIWKHVWUHDPLVODWHUVXEMHFWHGWRJOHDQLQJDQGÀVKFROOHFWLRQTaon basket traps are laid down in preparation for their deployment.

J. LINAUGO, E. ALCALA, A. BUCOL, C.C. MENES, ET AL

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Bago Jalaur

Table 1.

List of Introduced Species Found in Bago and Jalaur River Systems and Their Ecological Impact and Status of Establishment in the River.

Species Common Name River Adverse Establishment Impact

•Š›’’ŠŽ

•Š›’Šœȱ‹Š›ŠŒ‘žœȱ Š•”’—ȱŠęœ‘ȱ ȱȱȱƸȱ ȱȱȱƸȱ Žœ^ŗȱ ȱȱȱȱȱȱȱȱȱȱŽœ^ŗ

‘Š——’ŠŽ

‘Š——Šȱœ›’ŠŠȱ —Š”Ž‘ŽŠȱ ȱȱȱƸȱ ȱȱȱƸȱ —”—˜ —^śȱȱȱȱȱȱȱȱȱ—”—˜ —^ś

˜ŽŒ’•’’ŠŽ

Poecilia sphenops ˜••¢ȱ ȱ ȱȱȱƸȱ ȱȱȱƸȱ —”—˜ —^śȱȱȱȱȱȱȱȱȱ—”—˜ —^ś Poecilia reticulata ž™™¢ȱ ȱ ȱȱȱƸȱ ȱȱȱƸȱ Žœ^ŗȱ ȱȱȱȱȱȱȱȱȱŽœ^Ř Xiphiphorus helleriȱ ›ŽŽ—ȱ ˜›Š’•ȱ ȱȱȱƸȱ ȱȱȱƸȱ Žœ^Řȱ ȱȱȱȱȱȱȱȱȱŽœ^Ř

¢™›’—’ŠŽ

¢™›’—žœȱŒŠ›™’˜ȱ ˜––˜—ȱŠ›™ȱ ȱȱȱƸȱ ȱȱȱƸȱ Žœ^ŗȱ ȱȱȱȱȱȱȱȱȱȱȱȱȱȱŽœ^ŗ Danio rerio Ž‹›Šȱ’œ‘ Ƹȱ ȱȱȱȬȱ —”—˜ —^śȱȱȱȱȱȱȱȱ—”—˜ —^ś

’Œ‘•’ŠŽ

Oreochromis niloticus ’•Žȱ’•Š™’Šȱ ȱȱȱƸȱ ȱȱȱƸȱ Žœ^ŗȱ ȱȱȱȱȱȱȱȱȱȱȱȱȱȱŽœ^ŗ Oreochromis ˜£Š–‹’šžŽ

mossambicus ’•Š™’Š Ȭȱ ȱȱȱƸȱ Žœ^ŗȱ ȱȱȱȱȱȱȱȱȱȱȱȱȱȱŽœ^ŗ

œ™‘›˜—Ž–’ŠŽ

Trichogaster

trichopterusȱ ‘›ŽŽȬœ™˜ȱ ˜ž›Š–’ȱ ȱȱȱȬȱ ȱȱȱƸȱ ȱ

˜›’ŒŠ›’’ŠŽ

Pterygoplichthys

ȱȱȱȱ’œ“ž—Œ’Ÿžœȱ Š’•ę—ȱŠęœ‘ȱ ȱȱȱƸȱ ȱȱȱȬȱ —”—˜ —^śȱȱȱȱȱȱȱ—”—˜ —^ś

–™ž••Š›’ŠŽ

Pomacea canaliculataȱ ˜•Ž—ȱ™™•Žȱ—Š’•ȱ ȱȱȱƸȱ ȱȱȱƸȱ Žœ^řȱ ȱȱȱȱȱȱȱȱȱȱȱȱȱŽœ^ř

—’˜—’ŠŽ

›’œŠ›’Šȱ™•’ŒŠŠȱ ˜•Žȱ›˜ —ȱžœœŽ•ȱ ȱȱȱƸȱ ȱȱȱƸȱ —”—˜ —śȱȱȱȱȱȱȱȱȱȱȱȱ—”—˜ —^ś

›’˜—¢Œ‘’ŠŽ

Pelodiscus sinensis ‘’—ŽœŽȱ˜Ȭœ‘Ž••Žȱž›•ŽȱȱȱȱƸȱ ȱȱȱȬȱ —”—˜ —śȱȱȱȱȱȱȱȱȱȱȱȱ—”—˜ —^ś

ž˜—’ŠŽ

Bufo marinus

(Rhinella marina) Š—Žȱ˜Šȱ ȱȱȱƸȱ ȱȱȱƸȱ Žœ^Śȱ ȱȱȱȱȱȱȱȱȱȱȱȱȱŽœ^Ś

Š—’ŠŽ

Rana (Hylarana) ˜––˜—ȱ ›ŽŽ—

erythraeaȱȱ ȱȱȱȱ›˜ȱ ȱ ȱȱȱƸȱ ȱȱȱƸȱ —”—˜ —Ŝȱȱȱȱȱȱȱȱȱȱȱȱ—”—˜ —^Ŝ

Total 16 sp. 14 13

ŗŸŽ›œŽȱ’–™ŠŒȱŠ—ȱŽœŠ‹•’œ‘–Ž—ȱǻ‹ŠœŽȱ˜—ȱ›˜ŽœŽȱŠ—ȱŠž•¢ǰȱŘŖŖŜǼ

؍ŸŽ›œŽȱ’–™ŠŒȱŠ—ȱŽœŠ‹•’œ‘–Ž—ȱǻ‹ŠœŽȱ˜—ȱ‘Ĵ™DZȦȦŽ—ǯ ’”’™Ž’Šǯ˜›Ȧ ’”’Ȧ ›ŽŽ—ȏœ ˜›Š’•Ǽ

řȱŸŽ›œŽȱ’–™ŠŒȱŠ—ȱŽœŠ‹•’œ‘–Ž—ȱǻ‹ŠœŽȱ˜—ȱ‘Ĵ™DZȦȦŽ—ǯ ’”’™Ž’Šǯ˜›Ȧ ’”’Ȧ˜–ŠŒŽŠȏŒŠ—Š•’Œž•ŠŠǼ

ŚȱŸŽ›œŽȱ’–™ŠŒȱŠ—ȱŽœŠ‹•’œ‘–Ž—ȱǻ‹ŠœŽȱ˜—ȱ‘Ĵ™DZȦȦŽ—ǯ ’”’™Ž’Šǯ˜›Ȧ ’”’ȦŠ—Žȏ˜ŠǼ

ś—”—˜ —ȱ˜›ȱ—˜ȱ’—˜›–Š’˜—

Ŝȱ—”—˜ —ȱ˜›ȱ—˜ȱ’—˜›–Š’˜—ȱǻ’Žœ–˜œȱŽȱŠ•ǯǰȱŘŖŖŜǼ

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Š–™•Žœȱ Š›Žȱ Œž››Ž—•¢ȱ Ž™˜œ’Žȱ ’—ȱ ’••’–Š—ȱ —’ŸŽ›œ’¢Ȭǯǯȱ ˜—£Š•Žœȱ žœŽž–ȱ ˜ȱ Šž›Š•ȱ ’œ˜›¢ȱ ǻȬ Ǽǰȱ ž–ŠžŽŽȱ

’¢ǯȱ Š¡˜—˜–’Œȱ ’Ž—’’ŒŠ’˜—ȱ ˜••˜ Žȱ ••Ž—ȱ ǻŗşşŗǼǰȱ Š›Ž—’ȱ ǻŘŖŖŗǼǰȱŠ››’œ˜—ȱǭȱŽ—˜žȱǻŘŖŖŗǼǰȱ˜œœȱǻŘŖŖŗǼȱ˜›ȱ’œ‘ŽœȱŠ—ȱ•ŒŠ•Šȱ ǭȱ›˜ —ȱǻŗşşŞǼȱŠ—ȱŽ›™ ŠŒ‘ȱ‘’•’™™’—ŽœȱǻŘŖŖşǼȱ˜›ȱŠ–™‘’‹’Š—œȱ

Š—ȱ›Ž™’•Žœǯȱ

RESULTS

ȱ˜Š•ȱ˜ȱŗŜȱ’—›˜žŒŽȱœ™ŽŒ’Žœȱ Ž›Žȱ˜‹œŽ›ŸŽȱ’—ȱŠ˜ȱŠ—ȱŠ•Šž›ȱ

’ŸŽ›ȱœ¢œŽ–œȱǻŠ‹•ŽȱŗǼǯȱ‘ŽœŽȱ’—Œ•žŽȱŗŗȱœ™ŽŒ’Žœȱ˜ȱ’œ‘ǰȱ ˜ȱœ™ŽŒ’Žœȱ

˜ȱ›˜ȱŠ—ȱ˜—Žȱœ™ŽŒ’Žœȱ˜ȱž›•ŽǰȱŠ—ȱ ˜ȱœ™ŽŒ’Žœȱ˜ȱ–˜••žœ”œǯȱ‘Žȱ

—ž–‹Ž›œȱ ˜ȱ ’—›˜žŒŽȱ œ™ŽŒ’Žœȱ ˜‹œŽ›ŸŽȱ ’—ȱ ‘Žȱ  ˜ȱ ›’ŸŽ›œȱ  Ž›Žȱ

Š‹˜žȱ‘ŽȱœŠ–ŽȱǻŗŚȱŠ—ȱŗřǰȱ›Žœ™ŽŒ’ŸŽ•¢Ǽǯȱ‘Žȱ™›Ž˜–’—Š—ȱŽ¡˜’Œœȱ

˜‹œŽ›ŸŽȱ ’—ȱ ‘Žȱ ›’ŸŽ›œȱ  Ž›Žȱ ’œ‘Žœȱ ǻŗŗȱ œ™ŽŒ’ŽœǼȱ Œ˜–™˜œŽȱ ˜ȱ œ’¡ȱ

ŠšžŠ›’ž–ȱ œ™ŽŒ’Žœȱ Š—ȱ ’ŸŽȱ ’œ‘Ž›¢ȱ ˜›ȱ ŠšžŠŒž•ž›Žȱ œ™ŽŒ’Žœǯȱ ˜–Žȱ

˜ȱ‘ŽȱŠšžŠ›’ž–ȱœ™ŽŒ’Žœȱ‘ŠŸŽȱ‘’œ˜›’Žœȱ˜ȱŽ•’‹Ž›ŠŽȱ’—›˜žŒ’˜—ȱ

‹¢ȱ˜ŸŽ›—–Ž—ȱŠŽ—Œ’Žœȱ˜ȱŒ˜—›˜•ȱ–˜œšž’˜ȱ•Š›ŸŠŽȱ’—ȱ›Žœ‘ ŠŽ›ȱ

Ž—Ÿ’›˜—–Ž—œǯȱ ’¡ȱ ˜ȱ ‘Žȱ ŗśȱ œ™ŽŒ’Žœȱ ˜ȱ Ž¡˜’Œœȱ ‘ŠŸŽȱ —˜ȱ ”—˜ —ȱ

’—˜›–Š’˜—ȱ ˜›ȱ ‘Ž’›ȱ ŽŒ˜•˜’ŒŠ•ȱ ›˜•Žȱ Š—ȱ ŒŠ™ŠŒ’¢ȱ ˜ȱ ŽœŠ‹•’œ‘ȱ

’—ȱ˜ž›ȱ›’ŸŽ›ȱœ¢œŽ–œȱǻŠ‹•ŽȱŗǼǯȱ˜ȱ’—›˜žŒŽȱœ™ŽŒ’Žœȱ˜ȱŒ›Š‹ȱ˜›ȱ

œ‘›’–™ȱ Ž›Žȱ˜ž—ȱ•’Ÿ’—ȱ’—ȱ‘Žȱ›’ŸŽ›œǯ

Notes on species FAMILY CLARIIDAE

Clarias batrachusȱǻŠ•”’—ȱŠęœ‘Ǽ

Š›•¢ȱŠŒŒ˜ž—œȱ˜ȱ‘Žȱœ™ŽŒ’Žœȱ’—ȱ‘Žȱ‘’•’™™’—Žœȱ’—Œ•žŽȱ‘Šȱ˜ȱŽ››Žȱ ǻŗşŘŚ‹Ǽǯȱȱ—Ž˜¢™Žȱ‘Šœȱ‹ŽŽ—ȱŽœ’—ŠŽȱ‹¢ȱȱŠ—ȱ˜ĴŽ•ŠȱǻŘŖŖŞǼȱ˜ȱ

ŠŸ˜’ȱŒ˜—žœ’˜—ȱ ’‘ȱ‘›ŽŽȱ˜‘Ž›ȱœ™ŽŒ’Žœȱ•’œŽȱž—Ž›ȱ‘ŽȱœŠ–Žȱ—Š–Žǯȱ

ȱ ’œȱ Œ˜—œ’Ž›Žȱ Šȱ ȃ›žŽȄȱ ›Žœ‘ ŠŽ›ȱ ꜑ȱ ˜ž—ȱ ’—ȱ ‘’•’™™’—Žȱ ›’ŸŽ›ȱ

œ¢œŽ–œȱǻŽ››ŽǰȱŗşŘŚŠDzȱŠ›ž–‹Š—ŠǰȱŘŖŖŘǼǯ

‘Žȱ œ™ŽŒ’Žœȱ Š™™ŽŠ›œȱ ˜ȱ ‹Žȱ Š’›•¢ȱ Œ˜––˜—ȱ ’—ȱ œŠ’˜—œȱ ŗȱ ˜ȱ řȱ ’—ȱ

Š˜ȱ’ŸŽ›ȱŠ—ȱŠ•œ˜ȱ’—ȱŠ’˜—œȱŗȱŠ—ȱŘȱ’—ȱŠ•Šž›ȱ’ŸŽ›ǯȱ—ȱŒ˜—›Šœǰȱ

Šž•Š¢Š—ȱ ǻŘŖŖŗǼȱ ˜ž—ȱǯȱ ‹Š›ŠŒ‘žœȱ ’—ȱ ˜—•¢ȱ ˜—Žȱ ˜žȱ ˜ȱ ꟎ȱ ›’ŸŽ›œȱ

œž’Žȱ’—ȱ‘Žȱž‹’ŒȱŠ¢ȱ˜›ŽœȱŽœŽ›ŸŽǯȱ‘Žȱ Š•”’—ȱŒŠęœ‘ȱŽŒ‘ŽœȱŠȱ

‘’‘Ž›ȱ™›’ŒŽȱŒ˜–™Š›Žȱ˜ȱ˜‘Ž›ȱ›Žœ‘ ŠŽ›ȱœ™ŽŒ’ŽœȱŠ—ȱ’œȱœ˜ž‘ȱŠŽ›ȱ

ŠœȱŠȱŽ•’ŒŠŒ¢ȱ’—ȱ‘ŽȱŠ˜ȱ’ŸŽ›ȱŠ›ŽŠȱǻ’ž›ŽȱřǼǯȱȱ J. LINAUGO, E. ALCALA, A. BUCOL, C.C. MENES, ET AL

144

FAMILY CHANNIDAE Channa striata (Snakehead)

Žȱ˜‹Š’—Žȱ˜ž›ȱœŠ–™•Žœȱ›˜–ȱŠ’˜—ȱŘȱ’—ȱŠ˜ȱ’ŸŽ›ȱŠ—ȱŠ’˜—ȱŗȱ

’—ȱŠ•Šž›ȱ’ŸŽ›ǯȱ˜—œ’Ž›ŽȱŠȱ™˜Ž—’Š•ȱ™Žœȱǻ›˜ŽœŽȱǭȱŠž•¢ǰȱŘŖŗŖǼǰȱ‘Žȱ

œ—Š”Ž‘ŽŠȱ’œȱ—ŽŸŽ›‘Ž•ŽœœȱŒ˜••ŽŒŽȱŠœȱ˜˜ȱ‹¢ȱ•˜ŒŠ•œȱ’—ȱ‘ŽȱŠ›ŽŠǯȱȱ’œȱ

—˜ȱ•’œŽȱ’—ȱ‘Žȱ •˜‹Š•ȱ—ŸŠœ’ŸŽȱ™ŽŒ’ŽœȱŠŠ‹ŠœŽȱǻ Ǽǯȱȱ

FAMILY POECILIIDAE Poecilia sphenops (Molly)

‘’œȱ œ™ŽŒ’Žœȱ  Šœȱ ŒŠ™ž›Žȱ ˜Ž‘Ž›ȱ  ’‘ȱ ˜‘Ž›ȱ œ™ŽŒ’Žœȱ ǻŽǯǯȱAnabas testudineusȱ Š—ȱRhinogobiusǼȱ ’—ȱ Š˜ȱ Š—ȱ Š•Šž›ȱ ’ŸŽ›œȱ žœ’—ȱ ꗎȬ

–Žœ‘ȱ—Žœȱǻ’ž›ŽȱŚǼǯȱŽ—’ęŒŠ’˜—ȱ˜ȱP. sphenopsȱ ŠœȱŒ˜—ę›–Žȱ‹¢ȱ

›ǯȱ¢——ŽȱŠ›Ž—’ȱ˜ȱ‘Žȱ—’ŽȱŠŽœȱŠ’˜—Š•ȱžœŽž–ȱ˜ȱŠž›Š•ȱ

’œ˜›¢ȱǻǼǯ

P. sphenopsȱ Šœȱ’—›˜žŒŽȱ’—ȱ‘Žȱ‘’•’™™’—Žœȱ˜ȱŒ˜—›˜•ȱ–˜œšž’˜ȱ

•Š›ŸŠŽȱǻŠ–™•˜—ŠȱŽȱŠ•ǯǰȱŘŖŖŝǼǯȱ‘Ž¢ȱŠ›Žȱ˜Ž—ȱ˜ž—ȱ’—ȱœ–Š••ȱœ›ŽŠ–œȱ

Š—ȱŒŠ—Š•œǯȱž•œȱ˜ȱ‘’œȱœ™ŽŒ’ŽœȱŠ›ŽȱŽ¡›Ž–Ž•¢ȱŠ›Žœœ’ŸŽȱŠ—ȱŠĴŠŒ”ȱ

Figure 3. :DONLQJFDWÀVKClarias batrachus, from Bago River.

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˜‘Ž›ȱ꜑ǰȱœ‘›Ž’—ȱꗜȱŠ—ȱœ˜–Ž’–Žœȱ”’••’—ȱ‘Ž–ǯȱ Poecilia reticulata (Guppy)

‘’œȱœ™ŽŒ’Žœȱ’œȱžœžŠ••¢ȱŒŠ™ž›Žȱ˜Ž‘Ž›ȱ ’‘ȱ˜‹’ŽœȱŠ—ȱP. sphenops

žœ’—ȱꗎȱ–Žœ‘ȱœŒ˜˜™ȱ—Žœȱ’—ȱŠ˜ȱŠ—ȱŠ•Šž›ȱ’ŸŽ›ȱœ¢œŽ–œǯȱ‘Ž¢ȱ

‘ŠŸŽȱ‹ŽŽ—ȱ˜‹œŽ›ŸŽȱ’—ȱœ›ŽŠ–ȱ›’‹žŠ›’ŽœȱŽŽ’—ȱ’—˜ȱ‘Žȱ–Š’—ȱ›’ŸŽ›ȱ

Œ‘Š——Ž•ȱŠ—ȱ’—ȱœ–Š••ȱ’–™˜ž—–Ž—œȱ˜›ȱ™˜˜•œȱŠ•˜—ȱ‘Žȱ›’ŸŽ›ǯȱȱ

‘Žȱœ™ŽŒ’Žœȱ Šœȱ’—›˜žŒŽȱ’—ȱ‘Žȱ‘’•’™™’—Žœȱ˜›ȱ–˜œšž’˜ȱŒ˜—›˜•ȱ ǻŠœŠ•ȱŽȱŠ•ǯǰȱŘŖŖŝǼǯȱȱ’œȱ”—˜ —ȱ˜ȱŽŠȱ‘ŽȱŽœȱ˜ȱ—Š’ŸŽȱ꜑ȱœ™ŽŒ’Žœȱ

Š—ȱœŽ›ŸŽœȱŠœȱ‘˜œȱ˜›ȱ‘Žȱ™Š›Šœ’’Œȱ—Ž–Š˜ŽǰȱŠ–Š••Š—žœȱŒ˜Ĵ’ǰȱŠ—ȱ

‘Žȱœ’Š—ȱŠ™Ž ˜›–ȱBothriocephalus acheilognathiȱ’—ȱŠ Š’’ȱǻ•›ŽŽǰȱ ŘŖŖŖǼǯȱ

Xiphiphorus helleri (Green Swordtail)

‘Žȱ œ™ŽŒ’Žœȱ  Šœȱ ŒŠ™ž›Žȱ žœ’—ȱ ꗎȬ–Žœ‘ȱ —Žœȱ ’—ȱ Š˜ȱ Š—ȱ Š•Šž›ȱ

’ŸŽ›œǯȱXiphophorus helleriȱ ŒŠ—ȱ ‹Žȱ ’œ’—ž’œ‘Žȱ ‹¢ȱ ‘Žȱ ™›ŽœŽ—ŒŽȱ ˜ȱ

Šȱ œ ˜›Ȭ•’”Žȱ Ž¡Ž—œ’˜—ȱ ˜—ȱ ‘Žȱ ŒŠžŠ•ȱ ę—ȱ ˜ȱ –Š•Žœȱ ǻ’ž›Žȱ ŜǼǯȱ ‘Žȱ

œ™ŽŒ’Žœȱ’œȱ—Š’ŸŽȱ˜ȱŽ—›Š•ȱ–Ž›’ŒŠȱ‹žȱ‘Šœȱ‹ŽŽ—ȱ’—›˜žŒŽȱ˜ȱ–Š—¢ȱ

Œ˜ž—›’ŽœȱŠ›˜ž—ȱ‘Žȱ ˜›•ȱ‘›˜ž‘ȱ‘ŽȱŠšžŠ›’ž–ȱ›ŠŽǯȱ•‘˜ž‘ȱ

Figure 4. Poecilia sphenops from Bago River.

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Figure 5. *DPEXVLDDIÀQLVDOVRNQRZQDVPRVTXLWRÀVKZDVLQWURGXFHGLQWKH3KLOLSSLQHVDV DTXDULXPÀVKDQGDVFRQWURODJDLQVWPRVTXLWRODUYDH

Figure 6. Xiphiphorus helleri, or Green Sword-tail was introduced to the Philippines as aquarium ÀVK7KLVPDOHVSHFLPHQLVGLVWLQJXLVKHGE\LWVH[WHQGHGVZRUGOLNHWDLO

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•’Ĵ•Žȱ’—˜›–Š’˜—ȱ’œȱ”—˜ —ȱ˜ȱ‘Žȱœ™ŽŒ’Žœȱ˜žœ’Žȱ’œȱ—Šž›Š•ȱ‘Š‹’Šǰȱ

’œȱ’—›˜žŒ’˜—ȱ˜ȱ˜‘Ž›ȱ‘Š‹’Šœȱ‘Šœȱ‹ŽŽ—ȱœžœ™ŽŒŽȱ˜ȱŒ˜—œ’žŽȱŠȱ

‘›ŽŠȱ˜ȱ‘Žȱ—Š’ŸŽȱ꜑ȱ™˜™ž•Š’˜—ȱǻ ǰȱŘŖŗŖǼǯ

FAMILY CYPRINIDAE Cyprinus carpio (Common Carp)

Š–™•Žœȱ ˜ȱ ‘’œȱ œ™ŽŒ’Žœȱ  Ž›Žȱ Œ˜••ŽŒŽȱ ›˜–ȱ ‹˜‘ȱ Š˜ȱ Š—ȱ Š•Šž›ȱ

’ŸŽ›œȱ ǻ’ž›Žȱ ŝǼǯȱ Ž››Žȱ ǻŗşŘŚ‹Ǽȱ ŠŒŒ˜ž—Žȱ ’œȱ ’—›˜žŒ’˜—ȱ ’—ȱ ‘Žȱ

‘’•’™™’—Žœȱ’—ȱŗşŗśǯȱ •˜‹Š•ȱ—ŸŠœ’ŸŽȱ™ŽŒ’ŽœȱŠŠ‹ŠœŽȱŒ’ŽȱŒŠ›™œȱŠœȱ

Šȱ–Š“˜›ȱŒŠžœŽȱ˜›ȱ‘ŽȱŽŒ•’—Žȱ’—ȱŠšžŠ’Œȱ–ŠŒ›˜™‘¢ŽœȱŠ—ȱ‘Žȱ—Šž›Š•ȱ

ŽŒ˜œ¢œŽ–ȱ ǻ •˜‹Š•ȱ —ŸŠœ’ŸŽȱ ™ŽŒ’Žœȱ ŠŠ‹ŠœŽǰȱ ŘŖŗŖǼǯȱ ‘Žȱ ŠŸŽ›œŽȱ

ŽěŽŒœȱ˜ȱŒŠ›™œȱ’—ȱŸŠ›’˜žœȱ™Š›œȱ˜ȱ‘Žȱ ˜›•ȱ‘ŠŸŽȱ‹ŽŽ—ȱ’œŒžœœŽȱ‹¢ȱ

ŸŠ›’˜žœȱŠž‘˜›œȱǻŠ–‹›Š—˜ȱŽȱŠ•ǯǰȱŗşşşDzȱ›•’—‘ŠžœȱǭȱŽ‘—Ž›ǰȱŘŖŖřǼǯ

Figure 7. Zebra Fish Danio rerio is an introduced species with no apparent invasive record on the environment.

Danio rerio (Zebra Fish)

‘›ŽŽȱ œ™ŽŒ’–Ž—œȱ  Ž›Žȱ Œ˜••ŽŒŽȱ ’—ȱ Š˜ȱ ’ŸŽ›ȱ ǻŠ’˜—ȱ Řȱ ’—ȱ ˜™Ž£Ȭ

ŠŽ—ŠǼȱžœ’—ȱꗎȬ–Žœ‘ȱ—Žœȱǻ’ž›ŽǯȱŞǼǯȱȱŽ ȱ’—’Ÿ’žŠ•œȱ Ž›ŽȱŠ•œ˜ȱ

Œ˜••ŽŒŽȱ ’—ȱ Šȱ Œ›ŽŽ”ȱ —ŽŠ›ȱ ‘Žȱ Ž—›Š•ȱ ‘’•’™™’—Žȱ ŸŽ—’œȱ ˜••ŽŽȱ ǻǼȱŒŠ–™žœȱ’—ȱž›Œ’ŠǰȱŠȱŽ ȱ”’•˜–ŽŽ›œȱŠ Š¢ȱ›˜–ȱŠ’˜—ȱŘǯȱ‘Žȱ

œ™ŽŒ’Žœȱ ’œȱ ‹Ž•’ŽŸŽȱ ˜ȱ ‘ŠŸŽȱ ‹ŽŽ—ȱ ’—›˜žŒŽȱ ‘›˜ž‘ȱ ‘Žȱ ŠšžŠ›’ž–ȱ

›ŠŽǯȱ˜ȱ™›’˜›ȱ’—˜›–Š’˜—ȱ˜—ȱ‘Žȱœ™ŽŒ’ŽœȂȱ’—›˜žŒ’˜—ȱ’—ȱŽ›˜œȱ’œȱ J. LINAUGO, E. ALCALA, A. BUCOL, C.C. MENES, ET AL

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ŠŸŠ’•Š‹•Žǯȱ‘’œȱœ™ŽŒ’Žœȱ Šœȱ—˜ȱ˜‹œŽ›ŸŽȱ’—ȱŠ•Šž›ȱ’ŸŽ›ǯȱ

‘ŽȱŽ‹›Šȱ’œ‘ȱ’œȱŒ˜—œ’Ž›Žȱ‘Š›–•Žœœȱ‹¢ȱ›˜ŽœŽȱŠ—ȱŠž•¢ȱǻŘŖŗŖǼȱ

Š—ȱ’œȱ—˜ȱ•’œŽȱŠœȱŠ—ȱ’—ŸŠœ’ŸŽȱœ™ŽŒ’Žœȱ’—ȱ‘Žȱ •˜‹Š•ȱ—ŸŠœ’ŸŽȱ™ŽŒ’Žœȱ

ŠŠ‹ŠœŽȱǻ ǼȱŠ—ȱŠœŠ•ȱŽȱŠ•ǯȱǻŘŖŖŝǼǯȱ

FAMILY CICHLIDAE

Oreochromis niloticus (Nile tilapia)

‘’œȱœ™ŽŒ’Žœȱǻ’ž›ŽȱşǼȱ ŠœȱŒ˜••ŽŒŽȱ›˜–ȱŠ••ȱœŠ’˜—œȱ˜ȱ‘Žȱ ˜ȱ›’ŸŽ›ȱ

œ¢œŽ–œȱŠ—ȱŠ™™ŽŠ›œȱ˜ȱ‘ŠŸŽȱŽœŠ‹•’œ‘Žȱ’œŽ•ȱ’—ȱŠ••ȱœž¢ȱœŠ’˜—œȱ’—ȱ

‘Žȱ›’ŸŽ›ȱœ¢œŽ–œȱ˜ȱŽ›˜œȱŠ—ȱŠ—Š¢ǯȱ‘Žȱœ™ŽŒ’Žœȱ˜™œȱ‘ŽȱŒŠŒ‘Ȭ™Ž›Ȭ

ž—’ȱŽě˜›ȱ˜›ȱ–˜œȱ꜑’—ȱŽŠ›œȱŠ—ȱ’œȱ›Žž•Š›•¢ȱŒ˜—œž–Žȱ•˜ŒŠ••¢ȱ

™Š›’Œž•Š›•¢ȱ’—ȱŠ’˜—ȱŘȱ’—ȱŠ•Šž›ȱ’ŸŽ›ǯ

—›˜žŒ’˜—ȱ ˜ȱ ‘’œȱ œ™ŽŒ’Žœȱ ’—˜ȱ ‘Žȱ ‘’•’™™’—Žœȱ  Šœȱ ˜—Žȱ

‹ŽŒŠžœŽȱ ˜ȱ ’œȱ ‘’‘ȱ ›Ž™›˜žŒ’ŸŽȱ ›ŠŽȱ Š—ȱ ›Ž•Š’ŸŽȱ ŽŠœŽȱ ˜ȱ Œž•ž›Žȱ ǻ’žŽ›Ž˜ȱǭȱ ’Š—’ǰȱŘŖŖśǼǯȱœȱŽěŽŒ’ŸŽȱ–˜ž‘Ȭ‹›˜˜’—ȱ›Ž™›˜žŒ’ŸŽȱ

œ›ŠŽ¢ȱŠ••˜ œȱ’ȱ˜ȱ’—Œ›ŽŠœŽȱ’—ȱ—ž–‹Ž›œȱŠȱŠȱ›ŠŽȱ‘ŠȱŒ˜–™Ž’’ŸŽ•¢ȱ

˜ŸŽ› ‘Ž•–œȱ—Š’ŸŽȱœ™ŽŒ’Žœȱ™˜™ž•Š’˜—œȱǻ’žŽ›Ž˜ȱǭȱ ’Š—’ǰȱŘŖŖśǼǯȱ

‘Žȱ ꜑ȱ ’œȱ Œ˜—œ’Ž›Žȱ Šȱ ‘’‘•¢ȱ ’—ŸŠœ’ŸŽȱ œ™ŽŒ’Žœȱ ‘Šȱ ™•ŠžŽœȱ Šȱ

ŸŠ›’Ž¢ȱ˜ȱŽŒ˜œ¢œŽ–œȱŽœ™ŽŒ’Š••¢ȱ’—ȱ‘Žȱ›˜™’Œœȱǻ ǰȱŘŖŗŖǼǯȱ‘Ž¢ȱŒŠžœŽȱ

Figure 8. Cyprinus carpioRU&RPPRQ&DUSLVDFRPPRQO\ÀVKHGVSHFLHVLQWKHXSVWUHDP portions of Bago and Jalaur Rivers.

INTRODUCED AQUATIC FAUNA OF TWO RIVER SYSTEMS

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Š•Š•ȱ ‹•˜˜–ȱ Š—ȱ Žž›˜™‘’ŒŠ’˜—ȱ ˜ȱ œ–Š••ȱ ™˜—œȱ Š—ȱ œ•˜ Ȭ̘ ’—ȱ

›’ŸŽ›ȱ ŠŽ›œȱ ‘Ž—ȱŠ›–Žȱž—Ž›ȱ’—Ž—œ’ŸŽȱŒ˜—’’˜—œȱǻ’žŽ›Ž˜ȱǭȱ ’Š—’ǰȱŘŖŖśǼǯ

Oreochromis mossambicus (Mozambique Tilapia)

‘’œȱœ™ŽŒ’Žœȱ ŠœȱŒŠ™ž›Žȱžœ’—ȱ꜑—ŽœȱŠ—ȱ꜑’—ȱ•’—Žœȱǻ‘˜˜”ȬŠ—Ȭ

•’—ŽǼȱ’—ȱœ’—’ęŒŠ—ȱ—ž–‹Ž›œȱ’—ȱ˜ —œ›ŽŠ–ȱœŠ’˜—ȱ˜ȱŠ•Šž›ȱ’ŸŽ›ȱ‹žȱ  Šœȱ—˜ȱ˜‹œŽ›ŸŽȱ’—ȱŠ˜ȱ’ŸŽ›ǯȱȱ

O. mossambicusȱ’œȱŠȱŽž›¢˜™’ŒȱŠ—ȱŸŠ’•Žȱœ™ŽŒ’Žœȱ˜ȱ’œ‘ȱ”—˜ —ȱ

˜›ȱ’œȱŒŠ™ŠŒ’¢ȱ˜ȱ’—ŸŠŽȱ‘Š‹’Šœȱ˜ȱ˜‘Ž›ȱ’œ‘ȱœ™ŽŒ’ŽœȱǻŽȱ˜˜›ȱ ǭȱ›ž˜—ǰȱŗşŞŞǼǯȱ‘Žȱ–˜ž‘Ȭ‹›˜˜’—ȱ‘Š‹’ȱ˜ȱ‘Žȱœ™ŽŒ’ŽœȱŠ••˜ œȱ

’ȱ˜ȱ—ž›ž›ŽȱŠ—ȱŒŠ››¢ȱ’œȱ¢˜ž—ȱ˜ŸŽ›ȱ•˜—ȱ’œŠ—ŒŽœȱŠ—ȱ’—ŸŠŽȱ

‘Š‹’Šœȱ Š›ȱ ›˜–ȱ ‘Žȱ ˜›’’—Š•ȱ œ’Žȱ ˜ȱ ’—›˜žŒ’˜—ȱ ǻ˜œŠȬ’Ž›ŒŽǰȱ ŘŖŖřǼǯȱ œȱ ™˜Ž—’Š•ȱ ‘›ŽŠȱ ˜ȱ ‘Žȱ —Š’ŸŽȱ œ™ŽŒ’Žœȱ ’œȱ ™Ž›ŒŽ’ŸŽȱ ‹¢ȱ  Š¢ȱ˜ȱŒ˜–™Ž’’˜—ȱ˜›ȱ˜˜ȱŠ—ȱ—Žœȱœ™ŠŒŽǯȱžŸŽ—’•Žœȱ‘ŠŸŽȱ‹ŽŽ—ȱ

˜Œž–Ž—Žȱ ˜ȱ ŽŽȱ ˜—ȱ ˜‘Ž›ȱ ’œ‘ȱ ǻŽȱ ˜˜›ȱ Žȱ Š•ǯǰȱ ŗşŞŜǼǯȱ ‘’œȱ

œ™ŽŒ’Žœȱ’œȱŒž››Ž—•¢ȱŒ˜—œ’Ž›ŽȱŠȱ™Žœǯȱ—ȱŠ Š’ȁ’ǰȱ‘’œȱœ™ŽŒ’Žœȱ’œȱ

œžœ™ŽŒŽȱ˜ȱ‹ŽȱŠȱ‘›ŽŠȱ˜ȱ—Š’ŸŽȱœ™ŽŒ’ŽœȱœžŒ‘ȱŠœȱœ›’™Žȱ–ž••Žȱ ǻMugil cephalusǼȱ ǻœŽŽȱ Š—Š••ǰȱ ŗşŞŝDzȱ ŽŸ’Œ”ǰȱ ŗşşŗǼǯȱ ‘Žȱ œ™ŽŒ’Žœȱ

‘Šœȱ‹ŽŽ—ȱŒ˜—œ’Ž›ŽȱŠȱ–Š“˜›ȱŒ˜—›’‹ž’—ȱŠŒ˜›ȱ’—ȱ‘ŽȱŽŒ•’—Žȱ˜ȱ

‘ŽȱŽœŽ›ȱ™ž™’œ‘ȱǻ¢™›’—˜˜—ȱ–ŠŒž•Š›’žœǼȱ’—ȱ‘ŽȱŠ•˜—ȱŽŠȱŠ›ŽŠȱ ǻ˜ž›Ž—Š¢ȱǭȱ˜‹’—œǰȱŗşŞşǼǯȱȱ—ȱŠ’’˜—ȱ˜ȱ‘’œǰȱŽ›Š’ŒŠ’˜—ȱ˜ȱ‘Žȱ

Figure 9. Oreochromis niloticus or Nile Tilapia is a ubiquitous species that has proliferated and established in Bago and Jalaur Rivers.

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œ™ŽŒ’Žœȱ‘Šœȱ‹ŽŽ—ȱœžŽœŽȱ˜—ȱŠ›Š ŠȱŠ—ȱŠž›žȱ‹ŽŒŠžœŽȱ˜ȱ‘Ž’›ȱ

™˜Ž—’Š•ȱ‘›ŽŠȱ˜ȱ‘Žȱ•˜ŒŠ•ȱœ™ŽŒ’ŽœȱŠ—ȱ‘ŽȱŽŒ˜œ¢œŽ–ȱǻ•›ŽŽǰȱ ŘŖŖŖǼǯ

FAMILY LORICARIIDAE

Pterygoplichthys disjunctivusȱ ǻ˜ž‘ȱ –Ž›’ŒŠ—ȱ Š’•ę—ȱ Šęœ‘Dzȱ ȃ“Š—’˜›ȱ꜑ȄǼ

’¡ȱ•Š›Žȱ–Šž›Žȱ’—’Ÿ’žŠ•œȱ–ŽŠœž›’—ȱřŖȬřśȱŒ–ȱ Ž›ŽȱŒŠ™ž›Žȱ‹¢ȱ

’••—Žœȱ’—ȱŠ˜ȱ’ŸŽ›ȱŠȱŠ›Š—Š¢ȱ˜™Ž£ȬŠŽ—Šȱ’—ȱž›Œ’Šǯȱ‘ŽȱŒŠ™ž›Žȱ

˜ȱ‘’œȱœ™ŽŒ’Žœȱ’œȱ‹Ž•’ŽŸŽȱ˜ȱ‹Žȱ‘Žȱꛜȱ›ŽŒ˜›ȱ˜›ȱ‘Žȱ’œŠ¢Šœȱ›Ž’˜—ǯȱ

‘ŽȱŽ—žœȱ‘Šœȱ‹ŽŽ—ȱ›Ž™˜›Žȱ’—ȱŠ–‹˜Š—Šȱ’—ȱ’—Š—Š˜ȱŠ—ȱ™Š››’ǰȱ

ŠŠ¢Š—ȱ’—ȱ˜›‘Ž›—ȱž£˜—ȱǻ‘ŠŸŽ£ȱŽȱŠ•ǯǰȱŘŖŖŜǼǯȱ

•‘˜ž‘ȱ ‘Žȱ œ™ŽŒ’Žœȱ ‘Šœȱ —˜ȱ ‹ŽŽ—ȱ ›ŽŒ˜––Ž—Žȱ ˜›ȱ

Œ˜—œž–™’˜—ȱ ‹¢ȱ ‘ŠŸŽ£ȱ Žȱ Š•ǯȱ ǻŘŖŖŜǼǰȱ ’œ‘Ž›˜•”ȱ ’—ȱ Š˜ȱ Œ˜••ŽŒȱ

‘’œȱ’œ‘ȱ˜›ȱ˜˜ǯȱ

FAMILY BELONTIDAE

Trichogaster trichopterus (Three-spot Gourami)

—•¢ȱ˜—Žȱ’—’Ÿ’žŠ•ȱ ŠœȱŒ˜••ŽŒŽȱ’—ȱŠ•Šž›ȱ’ŸŽ›ȱžœ’—ȱœŒ˜˜™ȱ—Žǯȱ‘’œȱ

œ™ŽŒ’Žœȱ’œȱŠȱ—Š’ŸŽȱ˜ȱ–Š’—•Š—ȱœ’Šǰȱ™Š›’Œž•Š›•¢ȱ’—ȱ‘ŽȱŽ”˜—ȱ›ŽŠǯȱ

Figure 10. Cristaria plicata was commonly found in Bago and Jalaur Rivers, including sections attached to the rivers like irrigation canals.

INTRODUCED AQUATIC FAUNA OF TWO RIVER SYSTEMS

151 Ž‘Ž‹Ž›ǰȱŒŠ‘Š—ǰȱǭȱ’••Ž›ȱǻŘŖŗŖǼȱŒ’ŽȱŽŸ’Ž—ŒŽȱ˜—ȱ‘Žȱ’—›˜žŒ’˜—ȱ

˜ȱ‘’œȱœ™ŽŒ’Žœȱ’—ȱ—’ŠǰȱŠ’ Š—ǰȱ˜›‘ȱŽœŽ›—ȱžœ›Š•’Šǰȱ‘’•’™™’—Žœǰȱ

Š™žŠȱŽ ȱ ž’—ŽŠǰȱ˜ž‘Ž›—ȱ›’ŒŠȱŠ—ȱ˜–’—’ŒŠ—ȱŽ™ž‹•’Œǯȱ

‘Žȱœ™ŽŒ’Žœȱ Šœȱ’—›˜žŒŽȱ’—ȱ‘Žȱ‘’•’™™’—Žœȱ™›’–Š›’•¢ȱ˜ȱŒ˜—›˜•ȱ Aedes aegyptiȱ •Š›ŸŠŽȱ ǻŠ–™•˜—Šǰȱ ’–Šǰȱ ž—‘Šǰȱ ǭȱ Š—Š—Šǰȱ ŘŖŖŝǼǯȱ

ž•’Š—˜ǰȱ žŽ››Ž›˜ȱǭȱ˜—šž’••˜ȱǻŗşŞşǼȱꛜȱ›ŽŒ˜›Žȱ‘ŽȱŽœŠ‹•’œ‘–Ž—ȱ

˜ȱ‘Žȱœ™ŽŒ’Žœȱ’—ȱ‘Žȱ‘’•’™™’—Žœǯȱ‘Žȱ™˜Ž—’Š•ȱ‘›ŽŠȱ˜ȱ‘Žȱœ™ŽŒ’Žœȱ

˜ȱ‘Žȱ—Š’ŸŽȱ™˜™ž•Š’˜—ȱ‘Šœȱ‹ŽŽ—ȱŠĴ›’‹žŽȱ˜ȱ‘Žȱœ™ŽŒ’ŽœȂȱŽ››’˜›’Š•ȱ

—Šž›Žȱ Š—ȱ Š›Žœœ’ŸŽȱ ‹Ž‘ŠŸ’˜›ǰȱ ‹Ž’—ȱ Š—ȱ ˜™™˜›ž—’œ’Œȱ ŒŠ›—’Ÿ˜›Žȱ ǻŽ‹‹ǰȱŠž‘Š—ǰȱǭȱ—˜ĴǰȱŘŖŖŝǼǯȱȱ

MOLLUSCA PELECYPODA Cristaria plicata

‘’œȱ ‹’ŸŠ•ŸŽȱ ǻ’ž›Žȱ ŗŖǼȱ ’œȱ Œ˜••ŽŒŽȱ ‹¢ȱ •˜ŒŠ•ȱ ›Žœ’Ž—œȱ ˜›ȱ ˜˜ȱ

’—ȱŠ˜ȱŠ—ȱŠ•Šž›ȱ’ŸŽ›œǯȱ‘Ž¢ȱŠ›Žȱ‘Š›ŸŽœŽȱ‹¢ȱ‘Š—ȱ‹¢ȱ’ŸŽ›œȱ

Žšž’™™Žȱ  ’‘ȱ ˜•Žœȱ Š—ȱ —Žȱ ‹Šœǯȱ Š›ŸŽœ’—ȱ ˜ȱ ‘Žȱ œ‘Ž••ȱ

Š™™ŽŠ›œȱ˜ȱ‘ŠŸŽȱŠȱœŽŠœ˜—Š•ȱŒ¢Œ•Žȱ‘Šȱ’œȱ’ŒŠŽȱ‹¢ȱ‘Žȱœ‘Š••˜ —Žœœȱ

Š—ȱ ŠŒŒŽœœ’‹’•’¢ȱ ˜ȱ ‘Žȱ œ‘Ž••ȱ ‘Š‹’Šȱ ˜›ȱ Œ‘Š——Ž•ȱ ‹˜˜–ǯȱ ž›’—ȱ

‘’‘ȱ œ›ŽŠ–ȱ •˜ ȱ ǻ•˜˜’—Ǽǰȱ ‘Žȱ œ‘Ž••œȱ Š›Žȱ ˜Ž—ȱ œ™Š›Žȱ ›˜–ȱ

Œ˜••ŽŒ’˜—ȱ Š••˜ ’—ȱ ‘Ž–ȱ ˜ȱ ›˜ ȱ ’—ȱ —ž–‹Ž›œȱ ǻ‘’œȱ œ™ŽŒ’Žœȱ ˜›–ȱ

Œ•žœŽ›Žȱ •Š¢Ž›œȱ ˜ȱ ŽŠȱ Š—ȱ •’ŸŽȱ œ‘Ž••œȱ ’—ȱ œŠ—¢ȱ Š›ŽŠœǼȱ ‹ŽŒŠžœŽȱ

˜ȱ‘Žȱ’’Œž•¢ȱ’—ȱŠŒŒŽœœ’—ȱ‘Ž–ȱŠ—ȱ‘ŽȱŠ™™Š›Ž—ȱ™˜˜›ȱšžŠ•’¢ȱ

˜ȱ ‘Žȱ œ‘Ž••ȱ –ŽŠȱ ‘Šȱ Ž—œȱ ˜ȱ ŠŒŒž–ž•ŠŽȱ œ’•ȦœŠ—ȱ ž›’—ȱ ‘Žȱ

™Ž›’˜ǯȱ˜ŒŠ•ȱŠŒŒ˜ž—œȱ’—’ŒŠŽȱ‘Šȱ‘Žȱœ‘Ž••œȱ‹ŽŒ˜–Žȱ–˜›Žȱ–ŽŠ¢ȱ

Š—ȱ™Š•ŠŠ‹•Žȱž›’—ȱ‘Žȱ›¢ȱœŽŠœ˜—ǯȱ

‘Žȱ œ™ŽŒ’Žœȱ ‘Šœȱ ‹ŽŽ—ȱ ’—›˜žŒŽȱ ’—ȱ ˜‘Ž›ȱ ›Žœ‘ ŠŽ›ȱ œ¢œŽ–œȱ

’—ȱŽ›˜œȱ›’Ž—Š•ǰȱ’—Œ•ž’—ȱŠ•Š—Š—ȱŠ”Žȱ’—ȱ’Š˜—ȱŠ—ȱŠ•Š—˜ȱ

–Š••ȬŠŽ›Ȭ–™˜ž—–Ž—ȱ’—ȱŠ–‹˜Š—ž’Šǯȱ GASTROPODA

Pomacea canaliculata (Golden Apple Snail)

ŽȱžœžŠ••¢ȱ˜‹œŽ›ŸŽȱ‘’œȱœ—Š’•ȱ’—ȱ›’ŸŽ›ȱ‹Š—”œȱŠ—ȱ›’ŒŽȱ™Š’ŽœǯȱPomacea canaliculataȱ Šœȱ˜›’’—Š••¢ȱ’—›˜žŒŽȱ›˜–ȱ˜ž‘ȱ–Ž›’ŒŠȱ˜ȱ˜ž‘ŽŠœȱ

œ’ŠȱŠ›˜ž—ȱŗşŞŖǰȱŠœȱŠȱ•˜ŒŠ•ȱ˜˜ȱ›Žœ˜ž›ŒŽȱŠ—ȱŠœȱŠȱ™˜Ž—’Š•ȱ˜ž›–Žȱ

Ž¡™˜›ȱ’Ž–ǯȱ‘Žȱ–Š›”Žœȱ—ŽŸŽ›ȱŽŸŽ•˜™ŽDzȱ‘Žȱœ—Š’•œȱŽœŒŠ™Žȱ˜›ȱ Ž›Žȱ J. LINAUGO, E. ALCALA, A. BUCOL, C.C. MENES, ET AL

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›Ž•ŽŠœŽǰȱŠ—ȱP. canaliculataȱ‹ŽŒŠ–ŽȱŠȱœŽ›’˜žœȱ™Žœȱ˜ȱ›’ŒŽȱ‘›˜ž‘˜žȱ

–Š—¢ȱŒ˜ž—›’Žœȱ˜ȱ˜ž‘ŽŠœȱœ’Šǯȱ—ȱ‘Žȱ‘’•’™™’—Žœǰȱ’ȱ’œȱŒ˜—œ’Ž›Žȱ

‘Žȱ —ž–‹Ž›ȱ ˜—Žȱ ›’ŒŽȱ ™Žœȱ Š—ȱ ‘Šœȱ ŒŠžœŽȱ ‘žŽȱ ŽŒ˜—˜–’Œȱ •˜œœŽœǯȱ

˜Ž—’Š•ȱ’–™ŠŒœȱ˜ȱ‘Žȱœ™ŽŒ’Žœȱ’—Œ•žŽȱŽœ›žŒ’˜—ȱ˜ȱ—Š’ŸŽȱŠšžŠ’Œȱ

ŸŽŽŠ’˜—ȱ •ŽŠ’—ȱ ˜ȱ œŽ›’˜žœȱ ‘Š‹’Šȱ –˜’ęŒŠ’˜—ȱ Š—ȱ Œ˜–™Ž’’˜—ȱ  ’‘ȱ—Š’ŸŽȱœ—Š’•œǯȱ—›˜žŒŽȱP. canaliculataȱ‘Šœȱ‹ŽŽ—ȱ’–™•’ŒŠŽȱ’—ȱ

‘ŽȱŽŒ•’—Žȱ˜ȱ—Š’ŸŽȱœ™ŽŒ’Žœȱ˜ȱ’•Šȱ™™•Žȱ—Š’•œȱ’—ȱ˜ž‘ŽŠœȱœ’Šǯȱ

˜›Ž˜ŸŽ›ǰȱŒ•’–Š’ŒȬŒ‘Š—Žȱ–˜Ž•’—ȱ‘Šœȱœ‘˜ —ȱ‘Šȱ‘Žȱœ™ŽŒ’Žœȱ‘Šœȱ

‘Žȱ™˜Ž—’Š•ȱ˜ȱœ™›ŽŠȱ˜ȱ˜‘Ž›ȱ™Š›œȱ˜ȱ‘Žȱ ˜›•ȱ‘Šȱ‘ŠŸŽȱ—˜ȱ‹ŽŽ—ȱ

’—ŽœŽǰȱ•’”Žȱ‘Žȱ›’ŒŽȬ›˜ ’—ȱŠ›ŽŠœȱ˜ȱ—’Šȱǻ ǰȱŘŖŗŖǼǯ

AMPHIBIA

FAMILY BUFONIDAE Bufo marinus (Rhinella marina)

˜––˜—ȱ ‘›˜ž‘˜žȱ ‘Žȱ ‘’•’™™’—Žœǰȱ ‘’œȱ œ™ŽŒ’Žœȱ  Šœȱ ’—›˜žŒŽȱ

ž›’—ȱ ‘Žȱ ŗşřŖœȱ ‹¢ȱ ‘Žȱ ž›ŽŠžȱ ˜ȱ •Š—ȱ —žœ›¢ȱ ǻǼȱ ˜ȱ Œ˜—›˜•ȱ

‹ŽŽ•Žœȱ ’—ȱ œžŠ›ŒŠ—Žȱ ꎕœȱ ǻ’Žœ–˜œǰȱ ’Žœ–˜œȱ ǭȱ ›˜ —ǰȱ ŘŖŖŜǼǯȱ

•‘˜ž‘ȱ ‘Žȱ œ™ŽŒ’Žœȱ ’œȱ Œ˜—œ’Ž›Žȱ Ž››Žœ›’Š•ǰȱ ™Š›ȱ ˜ȱ ’œȱ •Š›ŸŠ•ȱ •’Žȱ

Œ¢Œ•Žȱ’œȱ’—ȱŠšžŠ’ŒȱŽ—Ÿ’›˜—–Ž—œǯ

‘’œȱœ™ŽŒ’Žœȱ Šœȱ˜ž—ȱ’—ȱŠ••ȱœŠ–™•’—ȱœŠ’˜—œǰȱžœžŠ••¢ȱ’—ȱ™˜˜•œȱ

˜›–ŽȱŠ•˜—ȱ‘Žȱ›’ŸŽ›ȱ‹Š—”ǯȱ•‘˜ž‘ȱ›ŽŒ˜›œȱ›˜–ȱ˜‘Ž›ȱŒ˜ž—›’Žœȱ

’—’ŒŠŽȱ‘Šȱ‘Žȱœ™ŽŒ’Žœȱ‹ŽŒŠ–ŽȱŠȱ™Žœȱž—Ž›ȱ’—›˜žŒŽȱŒ˜—’’˜—œǰȱ

’œȱ’–™ŠŒȱ˜—ȱ‘’•’™™’—ŽȱŽŒ˜œ¢œŽ–œȱ’œȱ—˜ȱ‘ŠȱŠ™™Š›Ž—ȱǻǯȱ•ŒŠ•Šȱ

™Ž›œǯȱ˜‹œŽ›ŸǯǼǯȱ˜ ŽŸŽ›ǰȱ‘Žȱœ™ŽŒ’Žœȱ‘Šœȱ‹ŽŽ—ȱœžœ™ŽŒŽȱ˜ȱŒ˜–™ŽŽȱ

˜›ȱ ‹›ŽŽ’—ȱ œ’Žœȱ  ’‘ȱ ‘›ŽŽȱ ›˜ȱ œ™ŽŒ’Žœǰȱ Ž“Ž›ŸŠ›¢Šȱ ŒŠ—Œ›’Ÿ˜›Šǰȱ Polypedates leucomystax,ȱŠ—ȱŠ•˜ž•ŠȱŒ˜—“ž—ŒŠȱ—Ž›˜œŽ—œ’œȱ’—ȱ‘’•’™™’—Žȱ

•˜ •Š—ȱŠ›ŽŠœȱǻ˜œ‘’ǰȱŘŖŖśǼǯȱǯȱ•ŒŠ•Šȱǻ™Ž›œȱ˜‹œŽ›ŸǼǰȱ‘˜ ŽŸŽ›ǰȱŒ˜ž•ȱ

—˜ȱŒ˜—ę›–ȱ‘ŠŸ’—ȱ˜‹œŽ›ŸŽȱ‘’œȱŒ˜–™Ž’’˜—ȱ ’‘ȱ‘Žȱ•ŠĴŽ›ȱœ™ŽŒ’Žœȱ

˜—ȱŽ›˜œȱœ•Š—ǯȱȱ

RANIDAE

Rana (Hylarana) erythraea

Žȱ ˜‹œŽ›ŸŽȱ ‘Žȱ œ™ŽŒ’Žœȱ ’—ȱ ‹˜‘ȱ ›’ŸŽ›ȱ œ¢œŽ–œǰȱ ˜Ž—ȱ Ž—Œ˜ž—Ž›Žȱ

Š•˜—ȱ‘Žȱ‹Š—”œȱ˜ȱ‘Žȱ›’ŸŽ›œǰȱœ›ŽŠ–œǰȱŠ—ȱ›’ŒŽȱ™Š’ŽœǯȱŽœ™’Žȱ’œȱ  ’Žœ™›ŽŠȱ ’œ›’‹ž’˜—ȱ ’—ȱ ˜ž‘ŽŠœȱ œ’Šǰȱ ’Žœ–˜œȱ Žȱ Š•ǯȱ ȱ ǻŘŖŖŜǼȱ

œžœ™ŽŒȱ‘Šȱ‘’œȱœ™ŽŒ’Žœȱ–Š¢ȱ‹Žȱ’—›˜žŒŽȱ’—ȱ‘Žȱ‘’•’™™’—ŽœȱžŽȱ˜ȱ INTRODUCED AQUATIC FAUNA OF TWO RIVER SYSTEMS

153

’œȱŠ‹œŽ—ŒŽȱ’—ȱŠ•Š Š—ȱŠ—ȱž•žȱŠ›Œ‘’™Ž•Š˜ǯȱ

REPTILIA

FAMILY TRIONYCHIDAE Pelodiscus (Trionyx) sinensis

‘’œȱ œ˜Ȭœ‘Ž••Žȱ ž›•Žȱ ǻ’ž›Žȱ ŗŗǼȱ ’œȱ ‹Ž•’ŽŸŽȱ ˜ȱ ‘ŠŸŽȱ ‹ŽŽ—ȱ

ŠŒŒ’Ž—Š••¢ȱ’—›˜žŒŽȱ‘›˜ž‘ȱ‘Žȱ™Žȱ›ŠŽǯȱ˜ ŽŸŽ›ǰȱŠŒŒ˜ž—œȱ

˜ȱ ‘Žȱ œ™ŽŒ’Žœȱ ‹Ž’—ȱ ‹›˜ž‘ȱ ’—ȱ ‹¢ȱ ŽŠ›•¢ȱ ‘’—ŽœŽȱ ›ŠŽ›œȱ ˜›ȱ

™Ž›œ˜—Š•ȱ Œ˜—œž–™’˜—ȱ ǻ˜˜Ǽȱ ’œȱ Š•œ˜ȱ ‹Ž’—ȱ Ž—Ž›Š’—Žȱ œ’—ŒŽȱ ‘Žȱ

œ™ŽŒ’Žœȱ’œȱŠ•œ˜ȱ˜ž—ȱ’—ȱ™•ŠŒŽœȱ”—˜ —ȱ˜›ȱ’œȱ’—Ž—œ’ŸŽȱŒ˜—ŠŒȱ ’‘ȱ

ŽŠ›•¢ȱ‘’—ŽœŽȱ›ŠŽ›œȱǻǯȱ—ǰȱ™Ž›œǯȱŒ˜––Ǽǯ

—ȱ ’—’Ÿ’žŠ•ȱ ˜ȱ ‘’œȱ œ™ŽŒ’Žœȱ  Šœȱ ŒŠ™ž›Žȱ ’—ȱ ‘Žȱ Š˜ȱ ’ŸŽ›ȱ

œŠ–™•’—ȱ œŠ’˜—ȱ ’—ȱ ˜™Ž£ȬŠŽ—Šȱ ǻž—’Œ’™Š•’¢ȱ ˜ȱ ž›Œ’ŠǼȱ ‹¢ȱ ‘Žȱ

›ŽœŽŠ›Œ‘ȱŽŠ–ȱ  ‘’•Žȱ •ŽŠ—’—ȱ ‘›˜ž‘ȱ Šȱ ›˜Œ”ȱ ™’•Žȱ ’—ȱ Šȱ ŠœȬ̘ ’—ȱ

™Š›ȱ˜ȱ‘Žȱ›’ŸŽ›ǯȱ•‘˜ž‘ȱ‘Žȱœ™ŽŒ’ŽœȱŠ™™ŽŠ›œȱ˜ȱ‹Žȱž—Œ˜––˜—ǰȱ˜Ž—ȱ

’–Žœȱ’ȱ’œȱŠŒŒ’Ž—Š••¢ȱŒŠ™ž›Žȱ‹¢ȱ꜑ȱ—ŽœǯȱŠ™ž›Žȱ’—’Ÿ’žŠ•œȱ

ŽŸŽ—žŠ••¢ȱŽ—ȱž™ȱŠœȱ˜˜ǯȱ‘’œȱœ™ŽŒ’ŽœȱŠ™™ŽŠ›œȱ˜ȱ‘ŠŸŽȱŽœŠ‹•’œ‘Žȱ’—ȱ

Š˜ȱ’ŸŽ›ȱ‹ŽŒŠžœŽȱ’ȱ’œȱ›Žž•Š›•¢ȱŒŠ™ž›Žȱ‘Ž›Žǯȱ‘’œȱž›•Žȱ›˜ œȱ˜ȱ

Šȱ•Š›Žȱœ’£Žǰȱœ˜–Ž’–Žœȱ›ŽŠŒ‘’—ȱřŖȱŒ–ȱ’—ȱŒŠ›Š™ŠŒŽȱ•Ž—‘ǯ

Figure 11. Pelodiscus sinensis, is a soft-shelled turtle found in Bago River. This species appears to have established in the area as indicated by regular capture of individuals.

J. LINAUGO, E. ALCALA, A. BUCOL, C.C. MENES, ET AL

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CONCLUSIONS

—˜›–Š’˜—ȱ ˜—ȱ ’—›˜žŒŽȱ ’—ŸŠœ’ŸŽȱ ŠšžŠ’Œȱ œ™ŽŒ’Žœȱ ǻǼȱ ’—ȱ

‘Žȱ ‘’•’™™’—Žœȱ Š™™ŽŠ›œȱ ˜ȱ ‹Žȱ Š’—’—ȱ Šœȱ –˜›Žȱ œž’Žœȱ Š›Žȱ ‹Ž’—ȱ

Œ˜—žŒŽǯȱ˜ ŽŸŽ›ǰȱœž’Žœȱ˜—ȱȱ’–™ŠŒœȱ˜—ȱ‘’•’™™’—ŽȱŠšžŠ’Œȱ

‘Š‹’Šœȱ Š—ȱ ŽŒ˜œ¢œŽ–œȱ Š›Žȱ ’—œžĜŒ’Ž—ǯȱ žŒ‘ȱ ’—ŠŽšžŠŒ¢ȱ ŠŽȱ

˜ȱ ‘Žȱ •ŠŒ”ȱ ˜ȱ ŒŠ™ŠŒ’¢ȱ ˜ȱ ŽŠ•ȱ  ’‘ȱ ȱ ‘Šœȱ Œ˜—›’‹žŽȱ ˜ȱ ‘Žȱ

™›˜•’Ž›Š’˜—ȱ˜ȱŽ¡˜’Œȱœ™ŽŒ’Žœǯȱ‘Ž›Žȱ’œȱŠȱ—ŽŽȱ˜›ȱ‘ŽȱŠšžŠŒž•ž›ŽȱŠ—ȱ

ŠšžŠ›’ž–ȱ›ŠŽȱ’—žœ›’Žœȱ˜ȱœ›’Œ•¢ȱŒ˜–™•¢ȱ ’‘ȱ‘Žȱ‹’˜œŠŽ¢ȱŠ—ȱ

šžŠ›Š—’—Žȱ›Žž•Š’˜—œǯȱ—ȱŠ’’˜—ȱ˜ȱ‘’œǰȱ˜ŸŽ›—–Ž—ȱŠŽ—Œ’Žœȱ•’”Žȱ

‘ŽȱŽ™Š›–Ž—ȱ˜ȱ›’Œž•ž›ŽȱǻǼȱŠ—ȱŽ™Š›–Ž—ȱ˜ȱ—Ÿ’›˜—–Ž—ȱ

Š—ȱ Šž›Š•ȱ Žœ˜ž›ŒŽœȱ ǻǼȱ œ‘˜ž•ȱ ™žȱ –˜›Žȱ Žě˜›ȱ ’—˜ȱ ‘Žȱ

–˜—’˜›’—ȱŠ—ȱŒ˜—›˜•ȱ˜ȱ‘ŽœŽȱŽ¡˜’Œȱœ™ŽŒ’Žœǯȱž›‘Ž›–˜›Žǰȱ™›’ŸŠŽȱ

Š—ȱ ˜ŸŽ›—–Ž—ȱ ŠŒŠŽ–’Œȱ ’—œ’ž’˜—œȱ ŒŠ—ȱ ™›˜Ÿ’Žȱ œ’—’ęŒŠ—ȱ

Œ˜—›’‹ž’˜—œȱ ’—ȱ ‘Žȱ –˜—’˜›’—ȱ Š—ȱ Œ˜—›˜•ȱ ˜ȱ ȱ ‘›˜ž‘ȱ ‘Ž’›ȱ

›ŽœŽŠ›Œ‘ȱ™›˜›Š–œǯȱŠœȱ‹žȱ—˜ȱ•ŽŠœǰȱž›‘Ž›ȱœž¢ȱŠ—ȱ–˜—’˜›’—ȱ

˜ȱȱŠ›Žȱ›ŽŒ˜––Ž—Žǰȱ™Š›’Œž•Š›•¢ȱ‘Žȱ•ŽŠœȱ”—˜ —ȱœ™ŽŒ’Žœǰȱœ˜ȱ‘Šȱ

™›˜™Ž›ȱŒ˜—›˜•ȱŠ—ȱ–Š—ŠŽ–Ž—ȱ‹¢ȱŠž‘˜›’’ŽœȱŒŠ—ȱ‹Žȱ˜—ŽǰȱŽœ™ŽŒ’Š••¢ȱ

˜ȱŽŽ›–’—Žȱ’ȱ‘Ž¢ȱŒ˜–™ŽŽȱ ’‘ȱ‘Žȱ—Š’ŸŽȱœ™ŽŒ’Žœȱ’—ȱ‘ŽȱŠ˜ȱŠ—ȱ

Š•Šž›ȱ’ŸŽ›ȱœ¢œŽ–œȱŠ—ȱ’—ȱ˜‘Ž›ȱ›’ŸŽ›ȱœ¢œŽ–œȱ˜ȱ‘ŽȱŒ˜ž—›¢ǯ

ACKNOWLEDGMENTS

ŽȱŠ›Žȱ›ŠŽž•ȱ˜ȱ‘Žȱ˜––’œœ’˜—ȱ˜—ȱ’‘Ž›ȱžŒŠ’˜—ȱ˜›ȱž—’—ȱ‘Žȱ›ŽœŽŠ›Œ‘ȱ

™›˜“ŽŒǯȱ‘Žȱœž¢ȱŠŒ”—˜ •ŽŽœȱ‘Žȱœž™™˜›ȱ™›˜Ÿ’Žȱ‹¢ȱ‘Žȱ•˜ŒŠ•ȱ˜ŸŽ›—–Ž—ȱž—’œȱ

˜ȱŠ˜ȱ’¢ǰȱž•ž™Š—Š—ǰȱ˜—ȱŠ•ŸŠ˜›ȱŽ—Ž’Œ˜ǰȱž›Œ’ŠǰȱŽŠ—Žœǰȱ’—•ŽǰȱŠ—ȱ

Š•’—˜ǯȱŽȱŠ›ŽȱŠ•œ˜ȱ‘Š—”ž•ȱ˜ȱ›œǯȱ—Ž•ȱ•ŒŠ•ŠȱŠ—ȱ›Ž—Œ’˜ȱŠŒ‘’ŒŠȱ˜ȱȬȱ ȱ˜›ȱž’’—ȱžœȱ‘›˜ž‘˜žȱ‘Žȱž›Š’˜—ȱ˜ȱ‘Žȱœž¢ǯ

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