NOTE ON DENTEX THUNBERGI
17. Genus SPARUS Linnaeus
Sparus Artedi,GeneraPiscium, 1738, p. 35.
SparusLiNN^us, Syst.Nat., ed.10, 1758, p.277(aurata,pagrus, etc.).
ChrysophrysCuvier, RegneAnim., ed.1, 1817, p.272(aurata).
AurataRisso,EuropeMeridionale, 1826, p.356 {semilunata=aurata)
.
Spams Fleming, British Animals, 1828, p. 211 (restricted to the gilthead, Sparusaurata).
IChrysoblephus Swainson, Nat. Hist. Class. Anim., vol. 2, 1839, p. 221 {gih- hiceps); (medianmolarsverylarge; preopercularlimbscaly).
PagrichthysBleeker, 1860 (driedspecimen,with oneanal epineremoved).
Ty2:)e.
—
Sparusaurata Linnasus.This genus agrees ingeneral with Pagrus, but the species are olive
and
silvery in color, never red,and
the teeth are in broader bands, theupper molarsinthree ormore
series.The
scales are smallerthan in Pagrus, Pagrosomus,and
Argyropsand
thebody
is still deeper.NO. 1875.
THE JAPANESE SPARID.^— JORDAN AND
TH03IPS0N.581 The
parietal (epiotic) crests of thecranium are welldeveloped,much
as in Pagrus pagrus. Species rather
numerous on
the coasts of theOld World from England
toJapan.The
group called ChrysoUephus,from South
Africa, with themedian
molars enlarged, is probably generically distinct.The
genus Calamus,from
tropical America, resemblesSparus
externally, but differs remarkably in the enlargedand
hollow interhaemal spine.(sparus, aTtdpor, the ancient
name, from
aTza'cpoj, togasp.) Keyto species,a^. Snout very blunt, theprofileapproachingthe vertical; anal raysIII, 11; dorsal rays
XI
orXIII,13; scalesabout60; rowsofolivaceousstripesalong theseriesof scales aries,27.
a^. Snoutmoreorlessacute,theprofileoblique, anal raysIII,8; scales45to 54.
b^. Bodyratherdeep, thedepth2J inbody length; dorsalrays XI, 12; scales45;
second anal spine If in head; olivaceous spotsalong the series of scales; a darkspotat originoflateral line, lowerfinsyellow latus, 28.
b^. Bodymoderatelyelongate,thedepth2| in length; dorsalraysXI,12; scales 54;
secondanalspine 2^inhead; nodistinctstreaks of spotsalongrowsof scales;
colordark; analfinpartlyblack swinhonis,29.
27.SPARUS ARIES (TemminckandSchlegel).
HTODAI(frontporgy); HEDAI(grunt porgy).
"fSpariissarbaForskIl, Descr. Anim., 1775, p.31(Red Sea).
"iChrysophrys sarbaCuvierandValenciennes,Hist.Nat.Poiss.,vol.6,1830, p.102 (Islede France).—RuppELL, N.W. Fische,1835, p. 110, pi. 28, fig. 1(Red Sea).—GtJNTHER, Cat. Fish. Brit. Mus., vol. 1, 1859, p. 488 (Red Sea).—
Kner,Reise Novara, Fisch., 1860, p. 88.
—
Klunzinger,Syn.FischeRoth.
Meer.,Verh. z.b. Ges.Wien, vol. 20, 1870, p. 759 (Red Sea).—Day,Fishes India,1875, p. 142, pi. 34,fig. 6 (Madras).—Sauvage, Poiss. Madag., 1891, p. 195,p\.25a, fig.3(Lagunesofeasterncoast ofMadagascar).
SparussarbaJ onBAii and Seale, Fishes Hongkong,Proc. Davenport Acad.Sci., vol.10, 1905, p.10 (notsynonymy).
ISparusbufonitesLacepede, Hist.Nat.Poiss., 1803,pp. 141, 143,pi.26, fig.3.
?SparuspsittacusLacepede,Hist. Nat.Poiss., 1803,pp. 141, 143.
IChrysophrys chrysargyraCuvier and Valenciennes, Hist. Nat. Poiss., vol. 6, 1830, p. 107 (Vizagapatam).
ChrysophrysariesTemminckand Schlegel, FaunaJaponica, 1843, p. 67, pi. 31 (Nagasaki).—Bleeker, Verh. Bat. Gen., vol. 26, 1857, p. 87 (Nagasaki).—
GtJNTHER,Cat. Fish. Brit.Mus.,vol.1,1859, p.489(China).—Steindachner and Doderlein, Beitr. Fische Japan's, II, Denkschr. kais. Akad. Wiss.
Wien,vol. 48, 1883, p.18 (Tokyo, Tango).
Sparus ariesJordan and Snyder, Check List,Ann. Zool. Jap., vol.3,pts.2and
3,1901, p. 79(Yokohama).
SparuslatusJordan andEvermann, Proc.U. S.Nat. Mus.,vol. 25, 1903, p.350 (Giran,Formosa).—JordanandRichardson,FishesFormosa, Mem.Carne- gieMus.,vol.4,No.4,1909. (NotofHouttuyn,notPagrits latusRichardson.)
HaUtat.—
Coasts of southernJapan and
China. Possiblyidentical withSparus
sarla Forskalof theRed
Seaand
East Indian region.Description of nine specimens
from Wakanoura,
onefrom Naga-
saki,three
from Tokyo, two from Hongkong, and
onefrom Moreton
Bay, Queensland, Austraha.Body
lengths rangingup
to 250mm.
582 PROCEEDINGS OF THE NATIONAL
MVSEU3I. TOL.41.Head
3^ inbody
length; depth 2|-; eye 4-4^ in head; depth of preorbital 3^ to 4^; D.XI
or XII, 13; A. Ill, 11; scales in longi- tudinal series 56 to 64, in transversebetween
insertions of dorsaland
anal 7 or 8/13 or 14; gill-rakers 6+
8.Dorsal profile arched
more
stronglyfrom
snout to dorsal than posteriorly;mouth
low, close to straight ventral profile; lowerjaw
included; snout short, rounded, slightly overhanging premaxillaries,its profile approaching the vertical at tip.
Eyes
wellbelow
upper surfaceof head, interorbitalregionarched stronglyfrom
side to side, 3 in head; frontals prominent.Snout
2{ in head, maxillary 2-|,ending under anterior half of pupil. Preorbitals broader than eye in adults. Preopercular
margin
entire, flexible.Jaws
powerful, heavy.Teeth
in upperjaw
in four rows of molars, outer rounded,Fig.9.— Sparusaries.
small; inner largest; third consisting posteriorly of
two
very large molars; fourthand
innermost ending at middle ofjaw. Three rows ofmolarsinlowerjaw, outermostsmallest, rounded, secondenlarging posteriorly, toform
three very large molars, the last largest; thirdrow
endingbefore middleofjaw.Three
incisor teethon
either side ofboth
jaws in front, setsomewhat
rodent-like.Third
and
fourth dorsal spines longest, 2J to 2| in head; last 2§in head, slightly longer than succeeding ray,
which
is longer than the last rays.Anal
spines similar to dorsal spines, the secondand
third equaland
ofsame
length as the latter.Anal
rays rapidly decreasing in length to last,which
is half that of first. Fin outline straight. Pectorallong, reachinganal, equal tohead
plus two-thirds of eye diameter. Ventrals 1| to IJ inhead. Caudal nearly as long ashead.NO. 1875.
THE JAPANESE SPARID^—JORDAN AND THOMPSON. 583
Scales
smooth
to touch, cycloid, present as a sheath at bases of dorsaland
anal, presenton
cheeks in five or six rows. Lateral linebut
little arched.Color of alcoholic specimens uniformly silvery, unsealed portions of
head
dusky, the iris yellow.Peritoneum
black, gill cavity lining clear.A
small blackspot at beginningof lateralline,insome
speci-mens
obsolete; analfinmore
or less dusky.This species is generally
common
in southern Japan,and may be
know^n at onceby
its deep body, blunt snout,and
long anal fin.The
eye inSparus
sarha is describedand
figured asmuch
larger than inSparus
aries (3 to 3| inhead
in S. sarha,and
4 to A^ in S.aries).
The
anterior profileismuch
steeperand more convex
inthe latter, apparently, thanisshown
in the plates ofSparussarha.The
specimensfrom Hongkong
recorded asSparus
sarhaby Jordan and
Scale are quiteidenticalwiththeJapanesespecimens.The
dentitionis the same,
and
not as described. If the Japaneseform
should prove inseparablefrom
that of the East Indies, itwould
stand asSparus
sarha.{aries, ram.)
28. SPARUS LATUSi Houttuyn.
KAIDSTT.
Sparus latus Houttuyn, Holl. Maat. Wet. Haarlem,
XX,
Deel. 2, 1782, (Nagasaki).tSparus hasta Bloch and Schneider, Syst. Ichth., 1801, p. 275 (Coromandel CoastandIndia).
Chrysophrys hasta Gunther, Cat. Fish. Brit.Mus., vol. 1, 1859, p. 490 (Japan) (part).
—
Steindachner and Doderlein,Beitr.Fische Japan's,II,Denkschr.
kais.Acad.Wiss. Wien, 1883, p. 17(Tokyo).
ICoius datnia Hamilton Buchanan, Fish. Ganges, 1822, p. 88, pi. 9, fig. 29 (Ganges).
SparusdatniaBleeker,Versl. kon.Acad.Wet.,2Rks.,vol.11,1876, p.5, pi.2
. (Nagasaki, notCalcutta).
—
Bleeker,AtlasIchth., vol.8,p. 109 (Nagasaki).
^Sparus berda Cuvier and Valenciennes, Hist. Nat. Poiss., vol. 6, 1830, p. 113 (Pondicherry).
—
Jordan and Richardson, Check List Philippine IslandBur. ofSci., 1, Manila, 1910, p. 31.
SparusberdaRichardson,Ichth.ChinaandJapan,1846, p.240 (Canton).
—
Jordan and Evermann, Proc. U. S. Nat. Mus., vol. 25, 1903, p. 350 (Formosa).
—
Jordan and Richardson,FishesFormosa,Mem.Carnegie Mus.,vol.4,No.4, 1909, p. 189(Formosa), (notSparus berdaForskai, 1775).
Chrysophrys longispinis Cuvier and Valenciennes, Hist. Nat. Poiss., vol. 6, 1830, p. 116 (Japan), (not type,fromCalcutta).
ICalamaraRussell,FishesofVizagapatam,vol.1,1803, p.63, pi.92 (Coromandel).
IChrysophrys calamaraCuvier andValenciennes, Hist. Nat.Poiss., vol.6,1830, p.117(Java,Malabar).—Cantor,Cat.Malay.Fish, 1850, p.48 (SeaofPinang).
Chrysophrys auripesRichardson,Ichth.China andJapan,1846, p.241 (Canton).
Chrysophrys xanthopoda Richardson, Ichth. China and Japan, 1846, p. 241 (Canton).
IThefollowingisthesubstanceofHouttuyn's accountof Sparuslatus:"I havecalledthisspecies tawhichthescalesare placed instripeslengthwise'widesea-brassen'becauseit isoneofthe widestofthe family,thespecunen3inches long,an inch and ahalfwide. Color yellowish, silveryunderthescales.
D. XII,9;A.Ill,8."
584 PROCEEDINGS OF THE NATIONAL MUSEUM.
VOL.41.Chrysophrys schlegeli Bleeker, Nat. Tijd. Ned. Ind., vol. 6, 1854, p.400(paxt, epecimen with45scales),(Nagasaki); NieuweNalez.Verh. Bat. Gen.,vol.26, 1857,pp. 20 and 86 (Nagasaki).—Jordan and Snyder, Check List, Ann.
Zool. Jap., vol.3,pts. 2and3, 1901, p.80(part).
SparuschrysopterusKishinouye,Zool.Mag. Tokyo,vol.19, 1907, p.327 (Kiusiu, Shikoku, Inland SeaandPacific coast ofCentralHondo).
Habitat.
—
SouthernJapan and
China.Described
from
a specunen 155mm.
longfrom Kobe and
one 190mm.
longfrom Wakanoura,
besides four specimens 100 to 108mm.
in
body
lengthfrom
thelatter place.Head
SJ inbody
length; depth ofbody
2|; eye 4^ in head;depth
of preorbital 6;secondanalspine 1^inhead; D. XI,11or12;A.Ill,8;
scales in lateralline45, intransverseseries 5 or6/12; gill rakers 6
+
9 or 10.z..,<^'
Fig.10.—Sparuslatus.
Dorsal profileof
head
nearly straight to occiput,body
profilemost
strongly arched anteriorly, sometimes slightly concave behind eyes.Snout more
pointed thaninS.aries, theupperprofilenot approaching thevertical;lowerjaw
notincluded, upperjaw
notoverhung by
tipof snout; preorbital narrow, not as broad as eye is long. Interorbital space arched moderately. Maxillary ending under center of pupil, 2^ in head; preopercle very minutely serrated, not flexible. Teeth in jaws only; lowerjaw
withtwo
rows posteriorly, both of rounded molars; innerrow
broadest save posteriorly,where
it is replaced b}^much
smaller ones; bothbecoming
granular anteriorly.A
third innerrow
of small rounded molars present along anterior halves of each mandible.Upper jaw
with three rows of rounded molars in front, theouterconical anteriorly,themiddlerow
extending onlyhalfway
to snout,where
it is replacedby
anotherrow
inside of the third, allbecoming
granular anteriorly. Six conical canines present in frontineachjaw.NO.1875.
THE JAPANESE SPARTD.^— JORDAN AND THOMPSON. 585
Thirdand
fourthdorsalspineslongest, If to 2 inhead,lastspine2^in head. Doreal rays but little longer than the latter.
Anal
spines very strong,heavy,thesecond strongest, 1| to1^in head, third2Jto 2| (in adults). First raymuch
shorterthan secondspinebut equal to third, tip of second spine nearly coinciding wdth that of last raywhen
supine, or extendingbeyond
its middle. Pectorals long, falcate, reaching third anal spine, equal tohead
plus eye diameter.Ventrals IJ in head. Caudal forked, nearly as long ashead.
Scalesveryslightlyctenoidbelow lateralline,presentas low sheath at bases of vertical fins; ro\\'^
on
cheek 6 or7. Lateral linemore
stronglyarched anteriorly thanin S. aries.Color of alcoholic specunens uniform, save for indistinct small blotch at upperangleofgillopening. Doreal
margined
stronglyand somewhat
shaded below with black.Anal
lightly stippled with black; afamt dusky
shade at base. Caudalmargined
with black.Scale rows indicated
by
longitudinal stripes, but not as prominently as inSparusaries.A
darkspot atorigin of lateralline.Peritoneum
silvery. Gill cavityliningcolorless. Inlifesilvery with dark
brown
streaks;fins
and
lower lobeofcaudal bright golden.It is very questionable whether this fish extends into the Indian Saas, or
beyond
Formosa. Bleeker considered it identical with specimensfrom
Calcutta, but says that there is a difference in thenumber
of doi-salspines, 12 in the Indian, 11 in Japanese. His firstdescriptionofS. sclilegeli
from Japan was
composite, according to his later paper.^The
characters were certainly takenfrom
the species herecalled 8. swinlionis,and
notfrom
the presentspecies.However,
he applied thename
S. sclilegeli to the latter,and
identified it with Indianspecimens.He
decided that;S'. hastawas
distinctfrom
either, as it probably is.On
this basis his Indiansynonymy
hasbeenworked
out. Sparus herda of Forekal,from
theRed
Sea, is not thesame
as our fish, ifwe may
judgeby
Riippell's figure,and
all pub- lished descriptionsfrom
that locality. Steindachnerand
Doderlein say2 that Ghrysophrys cuvieri, schlegeli, datnia,and
Jiasta of Bleeker are thesame
species in all probability, but the evidence available is all against thisview.A
carefulcomparison ofactualspecimensfrom
alllocalities willbenecessaryto decide the
number
of species present.We
adopt for the Japanese fish thename Sparus
latus.Hout-
tuyn's careless description has three distinct characters of the spe- cies,thedeep body, the rowsof olivespotson
therowsofscales,and
the presence of rays III, 8 in the anal. Sparus aries has the anal III, 11. Sparusswinlionishasno
distinct rows of spots.The name
Sparus herda,
from
theRed
Sea, is older than Houttuyn'sname, and
itmay
prove, after all, that thetwo
species are identical, although the present evidence points to the contrary. If the pre->Versl.kon.Acad.Wet.,vol.11,1876, p.5. aBeitr., vol.2,1883.
586 PROCEEDINGS OF THE NATIONAL MUSEUM.
vol.41.ceding species is called Sparus sarba, this one
may
wellbe
Sparus lerda.But
our material is inadequate to settle either question.We
consider Sparus chrysopterus Kishinouye identical with the present species as there isno
character givenby which
itmay
be distinguished.The
black spot ''in the anal fin near its base" isrepresented in our specimens
by
a slightdusky
shade. Mr. Kislii-nouye
gives the following translation ofthe original description:D. XI,II;A. Ill,8. Scales4or5-42-11. Pyloric coeca4. Vertebrse24. Height ofthebody2|inthetotallengthwithcaudal,andthe lengthof head4§ in the same.
Snoutobtuseandthe supraoccipital regionis alittle elevated. Incisors4-6 in each jaw, andmolars are weakly developed andsparsely distributedin3 irregular rows.
Thesecondspineofthe analverylong,longer thanthe lengthoftheinsertionofthe samefin. Grayish orgreenishgray at theback andsilvery white at theabdomen.
A
black spot respectively near theoriginofthelateallineandinthe analfinnearits base. Ventrals, anal,andthelower lobeofthecaudal yellow.Itistoldthatthisspeciesgrowstothelengthofabout36cm. Thisspeciesseemsto
spawnintheautumn.
Known
fromwarmerparts ofourcountry—
Kiushiu,Shikoku, InlandSea,andatthePacificcoast ofthecentral partofHondo.This species is generally
common m
southern Japan.Our
speci-mens
arefrom Wakanoura and Kobe.
(latus,broad.)
29.SPARUS SWINHONIS Gunther.
KURODAI (blackporgy).
Chrysophrys longispinis Temminck and Schlegel,Fauna Japonica,1843, p. 68, pi. 32 (Nagasaki).^
—
Richardson, Ichth. ChinaandJapan, 1846, p. 240 (on specimenofTemminckandSchlegel), (notofCuvierandValenciennes).
ChrysophrysschlegeliBleeker, Japan, Nat. Tijd. Ned.Ind., vol.6, 1854, p.400 (Nagasaki) (part, not of described specimen); Nieuwe Nalez. Verh. Bat.
Gen.,vol. 26, 1857, p.86(afterabove). (SeeVersl.kon.Acad.Wet.,2Rks., vol.11, 1876, p.7.)
Sparus sivinhonis GtJNTHEK, Ann. and Mag. Nat. Hist., 1874, p. 155 (Chifu in Chma).
Sparus schlegeli Bleeker, Versl. kon. Acad. Wet., 2 Rks., vol. 11, 1876, p. 2 (Nagasaki), (notofNieuweNalez.,p. 86).
—
Jordan and Snyder, CheckList,
Ann.Zool. Jap., vol.3,pts..2and3,1901, p. 80 (Japan),(in part).
Sparus longispinis Jordan and Richardson, Fishes Formosa, Mem. Carnegie Mus.,vol.4.No.4, 1909 (Takao).
Habitat.
—
CoastsofsouthernJapan and
China.Description of a specimen
from Kobe,
onefrom Wakanoura, and
12from
Tokyo.Body
lengthsfrom
60 to 225mm.
Head
3J in length of body, depth 2§; eye 4§ to 5 in head; depth of preorbital 6; D.XI,
12; anal III, 8; scales in longitudinal series 53 to 56 to last vertebra; in transverse seriesbetween
insertions of dorsaland
ofanal 7/15; gill-rakers 6+
8.Dorsal profilesimilar to that of Sparus latus; straight
from
snout to occiput;most
stronglyarchedanteriorly; sometimesslightly con- cave behind eyes.Snout somewhat
pointed, upperprofile not over- hanging premaxillaries.Lower jaw
not included, equal to upper.NO. 1875.
THE JAPANESE SPARIDJE— JORDAN AND THOMPSON. 687
Preorbitals not as broad as eyes. Interorbital space occasionally slightly gibbous. Maxillary 2| in head, ending
under
anterior half ofeye. Preoperclevery minutely serrated. Teethin jawsmuch
as in S. latus, in three rows above, intwo
below, allrounded
molars saveouterrow
above,which
issomewhat
incisorlike anteriorly,and
all
becoming
granular in front. Sixmoderate
canines present in fronton
each jaw.Dorsal spines longest in third
and
fourth, 2-2^ in head, last 2^ to 3J; dorsalrays ofnearlyequallength,about 2^inhead.Anal
spines ofmoderate
strength,between
those ofS. ariesand
S. latus; second contained 2 to 2^ in head, third 2| to 2f. First anal ray nearly equal to length of second spine,and
longer than third. Fin outline straight or but slightly concave, tip of second spine not extendingbeyond
middle of last ray,when
supine. Pectoral long, falcate, reaching to secondanal spine. Ventrals I3 inhead. Caudal nearly equaltohead..^
Fig.11.—Sparusswinhokis.
Scales slightly ctenoid below lateral line; not present
on
snout, preorbitals, jaws, interorbitalspace, orpreopercular margin; present as alow
sheath along bases of vertical fins,on
cheek in 7 or 8 rows.Color of alcoholic specimens uniform, those fi'om
Tokyo
with darker color, probably because of preservation.Rows
of scales indicatedby
prominent stripeson
cheeks as well ason
body. Ver-tical fins alldusky, dorsal
margined
with denser black, pectoralsand
ventralssomewhat
clearer; anal largely black.Peritoneum
silvery, gillcavityclear.Thisspeciesdiffers
from Sparus
latus ina greaternumber
of scales, lower dorsaland
anal spines,the latter not reachingbeyond
half the lengthof thelastray, beside several smaller pointsof difference.This well-marked species is very
common
in southernJapan and
China.Our
specimens arefrom Tokyo, Wakanoura, and Kobe.
It588 PROCEEDINGS OF THE NATIONAL
MU8EUBI. vol.41.has been confused with Sparus lotus under the
name
ofschlegeliand
with Sparus longispinisfrom
India, but it is distinctfrom
both,and
it is not yet recordedfrom
the East Indies.(Named
forMr. R. Swinhoe, consul atChifu.) 7.Family KYPHOSID^.
The
RUDDER
FISHES.Herbivorousfishes, withincisor teethonlyin the frontofthe jaws.
Body
oblong or elevated, withmoderate
or small scales, ctenoid or smooth.Mouth
moderate, withincisor-liketeethinthe frontofeach jaw;no
molars; teethon vomer and
palatmes present or absent;premaxillaries moderately protractile; preorbital rather narrow, sheathing the maxillary. Gill rakers moderate; pseudobranchise well developed; opercles entire. Gills 4, a slit behind the fourth;
gill
membranes
separate, freefrom
theisthmus; dorsalfincontinuous or divided, with 10 to 15 rather strongspines, the soft dorsalnaked
or scaly; anal with 3 spines; ventrals thoracic, the rays I, 5, an accessory scale at base; caudal lunate or forked; pectoral fin withall its rays branched. Intestinal canal elongate, with
few
ormany
pyloric caeca. Air bladder usually with 2 posterior horns. Verte- brae in ordinary or slightly increased
number,
24 to 28. Post-tem- poral ofnormal
percoid form, the stout forks not adnate to the cra- nium. Herbivorous shore fishes, feeding largelyon
green or olive algse; chiefly of the Mediterranean Seaand
the Pacific Ocean;most
of
them
valued as food.Keytogenera.
a'. GiRELLiNiE. Softpartofdoraaland analfinsnakedorpartlyscaled; headmore
orlessnaked; teethinbroadbands,allfreelymovable,none on vomer; pyloric caecavery numerous; vertebrae insomewhatincreasednumber. PacificOcean.
b}. Incisorsalltricuspid.
c^ Dorsal spines 14 or 15; eachjaw withaseries offlat,movable, tricuspidin- cisors,behindwhichiaa broadbandofsimilarsmallerones; dorsalcontinu-
ous, itsspineslow Girella,18.
a2. Kyphosin.e. Softparts of vertical fins closely scaled; teeth moreorlessfixed, usuallypresenton vomer, pyloriccaecanumerous.
(i '. Topofheadaswellassidesandjawsclosely scaled; broadbandsofteethbe-
hindtheincisors; villiformteethon vomer, palatinesandtongue; dorsal spineslow; incisorteethlanceolate,
e^. Incisorteethstrong,withhorizontal,backwardprojectingroots; softdor- salandanalnotmuchelevated.
f^. Incisorteeth well developed, eachwitha conspicuous horizontal pro- cessor root; caudalfinmoderate,aboutaslongasthehead, the outer raysnot3 timesas longas middlerays; junctionof gillmembranea
formingan angle Kyphosus,19.