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E-mail: ziaaddini@vru.ac.ir
زا ﻲﺧﺮﺑ و ﻲﻧاﻮﺧاﺮﻓ رﺎﺘﻓر ﻪﺴﻳﺎﻘﻣ ﻲﮔﮋﻳو
،ﻲﺘﺴﻳز يﺎﻫ ﻪﺳ رد
ﺖﻴﻌﻤﺟ توﺎﻔﺘﻣ ﻲﻳﺎﻴﻓاﺮﻐﺟ
مﺮﻛ رﺎﻧا هﺎﮔﻮﻠﮔ
، رد ﻲﻫﺎﮕﺸﻳﺎﻣزآ ﻂﻳاﺮﺷ
ﻲﻨﻳﺪﻟاءﺎﻴﺿ يﺪﻬﻣ
*1
زﺎﺴﻧاﺪﻠﮔ ﻦﻴﺴﺣﺪﻴﺳ ، ،2
رﻮﺷﺎﻋ ﺪﻤﺣا
3ي ﻠﻋ و ﻲ ﻢﺳﺎﻗ ﺎﺿر
4رﻮﭘ
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1998)
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+( # ( 3&, 00 %.!
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(Howse et al.,
1998)
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/( 8)
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)
Howse et al., 1998
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# 3
DE
FG %&( 67"
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Delisle & Royer, 1994
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%&' L( + M& 80 %') (
( !. %:;' N@" +( - 6 >*
O' %& ( P0 ) (= 6H4 %')
&7
% ( ! ) )
Calatayud et al., 2007
( . <0
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M&
# %0 S@' )
W2X %G= <0
Q&
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)
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( .
Arn et al.
(1983)
)
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L!
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(Casimero et al., 1999;
Kawazu & Tatsuki, 2002; Mazor & Dunkelblum, 2005; Zhao et al., 2007)
.
& 2 9]
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6^;= %') ( &)Q !\ ( V3 6, 2
1.Calling behavior
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Yousefi et al., 2004; Mozaffarian et al., 2007)
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1976a; Al-rubeai, 1987; Baker et al., 1991; Vetter et al., 1997)
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(1997)
(
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(2007)
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&
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(2007)
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.
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