CONG NGHE 177(01)

Nguyan Vii Bao va Dig I ap chi K.HUA HUL fi: CONG NGHE 177(01): 7 9 -

N G H I E N c u t ; M O T S O D A C D I E M P H A N L O A I C A C C H I T H U O C T O N G Gardenieae'uO C A P H E ( R u b i a c e a e ) T R O N G M O T S O H E S I N H T H A I Of V I E T N A M

Nguyen VO Bao'*, NguySn Thi M a i ^ Trln The Bach*

'Tru&ng Dai hQc Su-pham ~ DH Thai Nguyen 'Trudng Dgi hgc Ndng Lam - DH Thdi Nguyen, ^Vi?n Sinh thai vd Tdi nguyen smh vol TOM T A T

Cac chi thupc tong Danh danh (Gardenieae) trong hg Ca phe d Viet Nam dugc lua chon va sap xSp dya trfin he thflng ciia K. Schmann (1891) va bo sung ciia T. Chen & c6ng sy (2011), Theo ha thflng, tflng Danh danh (Gardenieae) thuflc phan hg Cinchonoideae - ho Ca phe (Rubiaceae) a Viet Nam gflm 19 chi. Cic chi thuflc tflng D^nh danh (Gtjn:/e«;eae)-ho C& pha phan bo hau khap ca nude, trong do, cic dai dien gSp nhifiu d miin Trung va Nam bfl, phan bo trong cac rirng thua va rung thir sinh d cac do cao khac nhau tuy loai. Ra hoa, kSt qua vao cac thdi diem khac nhau trong nam. Ve gia trj su dung, mgt so loai thugc chi dugc sii dung lam thuoc.

Til- k h i a : Cinchonoideae, Gardenieae, Rubiaceae, Ddnh ddnh, Cd phe DAT V A N DE

Phan loai hgc thuc vat dugc coi la mfit trong nhirng nganh khoa hgc quan trgng trong sinh hgc NhQng ket qua dang tin cay ciia phan loai thyc vgt da ddng gdp vao sy thanh cong cho nhieu nganh khoa hgc khac n h u sinh thai hgc, y hgc, dugc hgc, nfing h g c , . . . [ 4 ] . Ho Ca ph6 (Rubiaceae) vdi khoang 13,000 loai [6], duac phan bd trong 620 chi, hon 40 tfing, 6 Viet Nam, theo cac tai lieu da cdng bo mdi nhat ve hg Ca phe (Rubiaceae) cho t h i y , hg nay co khoang tren 90 chi va khoang 430 loai, phan bd rgng k h i p ca n u d e . . . [1], [2], [3], [7].

Ndi dung bai bao gdp phan bo sung va hoan chinh vdn tai lieu ve phan logi hg Ca phe (Rubiaceae) ndi chung va tong Danh danh {Gardenieae) noi rieng, phyc vu cho cac nghien ciiu sau hon trSn cac mat k h i c nhau ciia hg nay va cung cap cac can cu khoa hgc cho cdng trinh phan logi thuc vat va bien sogn thuc vat chi hg C a phe d Viet Nam; la c a sd khoa hgc phyc vu thiet thyc cho cac nganh n h u y - d u g c , tai nguySn thyc vat,...

DOI T U O N G v A P H U O N G PHAP NGHIEN C U X J

Boi tugng nghien clhi

' Tel 0973 987507. Email: nguyenvubao@dhspln edu vi

Cac chi thugc tdng Danh danh {Gardenieae) trong hg Ca phe (Rubiaceae) trong mdt so he sinh thai d Viet Nam bao gdm cac tieu ban khd dugc luu gifl tai cac phdng tieu ban trong nude, cac loai sdng trong mpt so he sinh thai t y nhien dugc thu thap qua cac cugc dieu tra thuc dja.

P h u v n g phap nghien ciiu

P h u a n g phap dung de nghien cuu phan loai cac chi thugc tdng Danh danh hg Ca phe (Rubiaceae) d Viet Nam la p h u a n g phap hinh thai so sanh dua vao dac diem hinh thai cua c a quan sinh dudng va chii yeu dua vao c a quan sinh san de nghien cdu.

Tham gia cac chuyen di thyc dja d V u d n Qudc gia, Khu bao ton thien nhien,,.. thu thgp mau vat va nghien ciiu dac diem hinh thai cung nhu sinh thai.

Mo ta mgt so dSc diem phan loai, phan bo, sinh hgc, sinh thai va gia trj sii dung ciia cac chi thugc tfing Danh danh trong mdt sd h?

sinh thai d Vet Nam, K E T QUA N G H I E N C U U

Lira chon he thong thich hop cho phan lo^i tong Danh danh {Gardenieae) - ho Ca phe (Rubiaceae) a Viet Nam

Chiing tdi da lya chgn he th6ng ciia K.

Schmann (1891) [7] va bd sung cQa Chen &

Nguyen Vu Bao vd Dtg Tgp chi KHOA HOC & CONG NGH? 'lyyi).

cgng sy (2011) [4] de ap dyng nghien ciiu phan logi tflng Danh diinh d Viet Nam vi day

!a h? thfing day dii, sii dung phucmg phap chinh la hinh thai so sanh de xac dinh mfii quan he gifla cac taxon va xac dinh vj tri ciia cac taxon, phu hgp vdi nguyen tic phan loai hien dai. Theo he thdng nay, tfing D^nh danh {Gardenieae) thufic phan hg Cinchonoideae - hg Ca phe (Rubiaceae) d Viet Nam gdm 19 chi.

Khoa dinh loai cac chi thuoc tong Danh danh {Gardenieae)

IA. Baunhuy mgt d 2A, Dinh noan try gifla 1. Ceriscoides 2B, Dmh noan ben 3A. So lugng no3n 2-6 2. Gardenia 3B. So lugng noan nhieu 4A. Bau do 2 la noan lam thanh 2. Kailarsenia

48. Bau do 3 la noan Iam thanh 4, Dioecresis

IB. Baunhuy hai 6 5A. Noan nhieu 6A. Bao phan dinh gdc 7A. Dai khdng mang Ifing 5. Aidia

7B. Dai mang long mm 6, Oxyceros 6B. Bao phan dinh lung 8A. Canh mang gai 9A. Hoa mgc don dgc 7, Fagerlindia 9B. Hoa mgc thanh cum 1 OA. Cum hoa d nach 1^

8. Randia

lOB. Cum hoa d dinh canh 9. Catunaregam 8B, Canh khflng mang gai

11 A, Cym hoa d dinh canh 10. Tarenna 11B. Cum hoa d nach la I2A. Noan bahang 11. Brachytome I2B. Noan hai hang 12. Morindopsis 5B. Noan 2-10 13A. Dinh phdi gifla vach 14A, Hoa mgc d dinh canh U.Alleizetta 14B. Hoa mpc d nach la 15A. La don mgc doi 14. Diplospora

15B. La don moc cheo hinh chfl thgp 15. Scyphiphora

13B. Dinh phdi d ndc vach I6A. Bao phan cd long, noSn 6-10 16. Hyptianthera

i6B. Bao phan khdng Ifing, noan 2-6 17A. Hoa mau 4, noSn 6 17. Hypobathrum 17B. Hoa mau 5 18A. Noan 3-4 18. Xantonnea 18B. Noan 2 19. Xantonneopsis

Dac diem hinh thai ^ng Danh danh {Gardenieae), ho Ca phe (Rubiaceae) d Viet Nam

* Deing song

Dang song cua cac chi thugc tflng Danh danh d Viet Nam thudng gap la dang cay than go {Aidia, Diplospora,,,.), 6 mdt so chi co mgt so loai dgng sdng la ciy than bui (cac chi Fagerlindia, Gardenia,...), da so la than go dung {Diplospora, Tarenna, Aidia,...), it khi la than bo hoSc trudn {Oxyceros).

Canh non cd nhieu tiet dien khac nhau, thudng khflng mang Iflng hogc cd thi mang 80

NguySn Vii Bao vd Dtg Tap chi KHOA HQC & CONG NGHE 177(01): 79-84 Idng min gap d mfit so loai thudc chi

Gardenia va Tarenna, cd doi khi canh non mang long mau nau nhu loai Diplospora fasciculata c6 mau xam Diplospora esculenla

(thugc chi Diplospora).

Mgt sd loai thugc cac chi nhu Diplospora, Oxyceros, Randi... canh mang gai. Cac gai vdi nhieu hinh dgng va kich thudc khac nhau, dang thang dai.

Hlnh: Ceriscoides howii Lo, 1-2. Cdnh mang hoa vd gai; 3 Ld; 4. Hoa; 5

Trdng bd doc, 6. Bdu nhuy cdt ngang:

y.Qud[4]

* La: Cac chi trong tfing Danh danh thudng CO la dan, mgc dfii, co cuong vdi chieu dai khac nhau, co mgt sd trudng hgp la mgc cheo hinh chir thap {Hyptianthera). Phien la cd nhieu hinh dgng khac nhau, be mat ciia phien la da sfi nhan, khflng mang long,

* Ld kem: Cac chi deu cd la kem, gifla cac chi cd su khac nhau ve hinh dang, kich thudc va thdi gian tdn tgi. Da so cac chl, la kem thudng rdi va cd hinh tam giac {Alleizelta,...) La kem thufic cac chi; Fagerlindia, Oxyceros thudng rung sdm, mgt so chi {Alleizelta) thudng rung mugn hoac khdng ryng.

* C^im hoa: Hoa thudng mgc thanh cym vdi nhieu dang khac nhau hoac mgc dan ddc {Fagerlindia, Gardenia,...). Cac hoa thudng mgc d dinh canh (Alleizelta,...) hogc mgc d nach cac la {Aidia. Randia,...)...

* Hoa: Hoa thudng la hoa mau 5 {Aidia...), doi khi mau 4. Da sd la hoa ludng tinh

{Fagerlindia), mgt so la hoa don tinh. Sd lugng la dai thudng la 4-5, phan dudi ciia cac la dai thudng dinh vdi nhau tao nen nhieu hinh dang khac nhau. Dai thudng tdn tgi d tren qua va lau rung nhu trong cac hoa thufic mfit sd chi Alleizelta, hoac co the rung sdm.

Trang hoa cd sfi lugng 4-5, cac la trang thudng dinh vdi nhau d phan gdc tao thanh cac ong trang khac nhau ve mau sac, kich thudc cung nhu hinh dang. Trimg thudng xep xoan d trong ny.

Sd lugng nhi da so la 5 {Aidia, Alleizelta.,...) hoac nhieu {Gardenia)-, chi nhi thudng ngan hoac rat ngan. Dau nhuy thudng chia liim hai thiiy (Aidia,...) tho ra ngoai dng trang; voi nhuy manh vii ngan; bau nhuy mdt d (Aidia,. .) hoac bau hai 8 sfi lugng noSn 2-10 (Diplospora) hoac nhieu (Brachytome) dinh noan thudng theo dang dinh noan try gifla hoac dinh noan ben.

* Qud: Da sd qua thudng la qua dan, mpng, CO nhiSu hinh dang khac nhau nhu hinh tron, hinh cau den hinh elip. Dai thudng khfing ryng va tdn tgi lau tren qua hoac sdm ryng.

* Hgt: Sfl lugng hat khac nhau gifla cac chi thay ddi tii' mpt den nhieu hat, cd dang hinh trai xoan, Ifii, det, co gdc canh, nhan, giau ngi nhu. Kich thudc hgt thay ddi tiiy Ioai fiac diem cac chi thuoc tong Danh danh {Gardenieae)^ hg Ca phe (Rubiaceae) Viet Nam Gen. 1. CERISCOIDES (Benth. & Hook.f.) Tirveng.

Cay byi hogc cay gd nhd. Cd khoang 11 loiii phan bfi d mgt so nude nhu Trung Quflc, Viet Nam. Ra hoa thang 10-4 (nam sau).

Mgc d trong cac thung lung, d do cao 400- 500 m so vdi muc nude bien.

Gen. 2. GARDENIA J.Ellis DANH DANH, NAM HUONG

Cay bui, ddi khi la cay gd nhd. Tren the gidi CO khoang 60-200 Ioai, d Viet Nam 13 loai.

Ve gia tri su dyng: G. augusta dimg lam canh hay G. lucida diing tri benh ngoai da va diet rufii va sau bg. Re cua G. augusta va G.

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Nguyen Vu Bao vd Dtg Tap chl KHOA HOC & C 6 N G NGHE stenophuylla clifla sdt li, cd tac dung Igi tieu,

chfla benh ngoai da, dau mat, dai dudng, thap khdp, dau dau. Qua G. augusta, G.

stenophuylla mau vang cam nhupm thiic an, chu'a viem gan, vimg da, viem ket mgc, loet mieng, dau rang, chay mau cam, dai ra mau.

Mgc rai rac trong rirng d dp cao 800 m, trong cac riing thii sinh, rflng thua.

Gen. 3. KAILARSENU Tiiveng. - DANH DANH Da so la ciy bui, canh non co Idng mau vang hoe. Cd khoang 10 loai phan bfi trong rimg nhi?t ddi chau A, Nude ta g5p 1 loai Kailarsenia godefroyana (Kurze) Tirv. Ra hoa thang 2-3, cd qua thang 5-12.

Gen.4.DIOECRESCIS Tirveng-NANH HEO Cay gfl, vd cac cay gfl gia co mau xam den, gai tu hop thanh cum tren nhflng canh chuyen biet phan gdc. C) Viet Nam cd mpt Ioai Dioecrescis erythroalada (Kurz) Tirv. Gfl cd mau vimg, diing lam can ndng cy. Mii cay dimg dl nhugm. Mgc riii rac trong rung thua.

Ra hoa thang 5-7, qua thang 11-12.

Gen. 5. AIDLA Lour. - TA HAY, GANG.

Ciiy go nho, khflng cd gai, mpc thing diing, cao tu 4-18 m. Tren thS gidi 40-50 loai, d Vi?t Nam gap dai dign 5 loai, phan bd chii ySu d BIC Kgn,,,, Gfl mau vang cd the lam dd mgc thdng thudng. Hoa, la va vd than dung udng thay tra, vd ding chfla benh sflt ret.

Mgc rai rac trong cac rung thir sinh, rflng thua. Ra hoa qua viio thang 4-12.

Gen. 6. OXYCEROS Lour. - GANG Cay bui hogc cay gd leo, canh khdng mang long nhung mang cac gai mgc doi. Cd khoang 10 loai, d nude ta gap dgi dien ciia 4 loai.

Phan bfi d Lang Son... Lii siiy kho tren than hSm udng giai kbit.

Mpc riii rac trong rirng d dp cao 500-800 m, dgc theo rupng, rgch; gap nhieu d cac tinh phi'aNam nude ta.

Gen. 7. FAGERLINDIA Tirveng. - GANG Cay gfl hogc cay byi, cd nhiSu nhanh, cac canh phan nhanh kieu ludng phan, hoac cay 82

gd nhd cao 1 -2 m, cd gai thing dai 0,6-2 cm, canh khflng mang Idng, Fagerlindia la mot chi nhd vdi khoang 10 loai. CJ nude ta gap dgi dien cua 3 Ioai phan bo d San La,....

Mgc rai rac nhftng nai dat khfi can coi (F.

Depauperata) hay mgc d cac trang trong tren vung da vdi. Ra hoa vao ciic thdi diem khac nhau tiiy tung loai.

Gen. 8. RANDIA L. - GANG GAI Cay than gd hoac than byi, thudng khfing mang long. Canh non khdng cd gai hogc c6 gai nhd {R. fasciculata), ddi khi Idn diii 1-5 cm (R. dasycarpa), cd khi che dfli R.

uliginosa. Trdng liim hang rao phfi bien d cac lang que {R.spinosa). Thiln dimg sic nude cho phu nft sau sinh de uong cho mau khde.

Thudng ra hoa vao cac thdi diem khac nhau, Mgc rai rac trong cac rirng thii sinh, rirng thudng xanh hay trong cac rimg thua, riing coi, tren dit can cfii.

Gen. 9. CATUNAREGAM Wolf. GANG TRAU, GANG TU HU

Cty bui hogc cay go nhd, canh non ngin, thudng mang cac gai. Cd khoang 5-10 loai, phan bo d mdt sd nude nhu Trung Quoc, Lao,... 0 nude ta phan bo d cac tinh nhir Tuyen Quang.

Mgc d trong rflng ven cac con sudi, tren doi hoac cac sudn nfli, cac thung IQng, d dg cao 1,600 m so vdi myc nude bien.

Gen. 10. TARENNA Gaertn. - TREN, CUT CHUOT, N H A U RUNG

Cay byi hogc cay gfi nhd, canh non co cgnh hogc vudng, than non cd bdn canh vuong, ciinh non dgp. S6 lugng loiii Idn vdi khoang 3.700 loai, d nude ta g3p dai dien ciia 17 loai.

Phan bo d Thai Nguyen,... Ngoai ra cdn ggp cr An Dg, Xri Lanka, Gfl dugc su dung trong xay dyng tgm, dong do mpc thdng thudng, lam bao bi. Rl giii dip chfta viet thuang. LS T.hoaensis dimg udng thay tra.

Mgc rai rac trong rung d do cao 300-1,400 tn, trong ckc rimg thudng xanh, mgc ven cac rung, dya rach. Thai gian ra hoa va tao qua khac nhau tuy tflng loai.

Nguyen Vii Bao vd Dtg Tgp chl KHOA HOC & CONG NGH$ 177(01): 79- Gen.ll. BRACHYTOME Hook. f. -DOAN

THIET, D O A N NHA

Ciiy byi hoSc cay gd nhd. La mgc doi ho3c thudng chum 3, La kem ben hogc ryng sau mdt thdi gian, rdl hogc dinh xung quanh cuflng va CO hinh tam giac. La mdt chi nhd, co 4 loiii trong khu vyc Ddng Nam A. C) nude ta gap dai dien hai loai: B. hirlelleta va B.

wallichii phan bo d Lao Cai (Sa Pa).

Mgc rai rac trong cac rimg thii sinh, dp cao 800 m (B. hirlelleta). Ra hoa vao thing 1-3.

Gen. 12. MORINDOPSIS Hook. f. - SONG NHAU Cay byi hogc cay gd nhfl, khdng mang long va gai, don ti'nh khac gfic. O nude ta, ggp dgi dien cua mgt loai M. capillaris Kurz, phan bd d Tay Ninh, Ddng Nai,...

Ra hoa vii ket qua vao thiing 5-6.

Gen.l3,ALLEIZETTELLAPit-AILpI, ALAY Cay byi dung, dfii khi dang than leo, khflng mang gai. Co hai loai dgi dien gap d Viet Nam va Trung Quoc, d nude ta chi gap mpt loai Alleizelta rubla - day la loai dgc huu hep cua Viet Nam, mdi chi gap d diem lay mau chuan thudc tinh Lam Dfing. La ngudn gen hiem, quy va dgc dao.

Ra hoa va qua vao thang 2-3. Mgc rai rac dudi tan rimg ram nhiet ddi thudng xanh, miia mua am, d sudn mii, do cao 1400 m.

Gen. 14. DIPLOSPORA DC. - SONG TU", TA DE, DE

Cay bui hoac cay gd nhd, canh khfing mang gai. La dan, mgc ddi. Nude ta g3p dai dign ciia 2 loai D.singularis va D. vilidiflora phan bo d cic tinh nhu Lang San. Gfi cd the dimg trong xSy dyng tam va ddng do mgc thflng thudng,

Mgc d cac riing thu sinh ddfi cao 600-1200 m so vdi myc nude bien.

Gen.lS.SCYPHIPHORA C.F.Gaertn. - COI Cay bui ho3c cay gd nhd, khdng cd gai va Ifing, canh non hai vuong, hoi mgng nude, dfii khi cd rl cgc. La mgc doi, cheo hinh chft thap. Co mpt loai phan bo d Quang Ninh,...

Lii chiet xuat ndng dung de chfla b^nh dau dg day. Gap d cac rirng ngap mgn. Ra hoa qua gan nhu quanh nam.

Gen. 16. HYPTIANTHERA Wight & Am. - TANG NHI MOC, CO LA QUAN Cay byi hoac cay gfi nhd, khflng cd gai. La mgc ddi, xep cheo hinh chft thgp. Cd mfit vai loiii d Trung Quoc, An Do, Lao, Myanmar, Nepan, Thai Lan, O Viet Nam co mpt loai H.

slricta gap dai dien d HaNgi.

Ra hoa va ket qua thang 5-7.

Gen. 17. HYPOBATHRUM Blume - SU"A, HADE

Cay gd nhd hoac cay bui, than chinh thudng thang vdi cac canh thudng sap xep hlnh chft thgp. La mgc doi, phien la cd hinh xoan thon hoac hinh bau due, day va ciing. 0 chau A cd khoang 30 loai. Re dugc dflng phoi hgp vdi cac vi thuoc khac sac lay nude uong de dieu tri benh ghe cdc.

Mgc rai rac trong rirng hoac dya rach, Ra hoa vao thang 4-5 hogc ra hoa quanh nam.

Gen. 18. XANTONNEA Pierre ex Pit. - XAN TON, X U A N T O N , COM Cay gd nhd cao 0,6-4 m, hoac ciiy gfi Idn cao 6-25 m, canh khfing mang long, co canh hoac d?p, mau trang hogc xam trang. Nude ta gap dai dien cua 5 loai, phan bd d mpt so nai nhu:

Kon Tum. Go sir dyng trong xay dyng va ddng do mpc thflng thudng. Song d cac vung mii thap den dg cao 1200 m. Ra hoa thiing 11- 4 (nam sau) X. Coffeoides.

Gen. 19. XANTONNEOPSIS Pitard in Lecomte - XUAN T O N , X A N G TON Cay bui cao 1-4 m, hoiin toiin nhan, cilnh non cd hinh 4 cgnh vdi 4 ranh doc, Loai dSc hflu hep ciia Vi?t Nam, mdi chi ggp d mgt diem lay mau chuan thugc tinh Khanh Hda (Nha Trang). Dgi di?n duy nhat ciia z\\\ Xantonneopsis dac hftu hep d Trung bg Vi^t Nam, Mpc rai rac dudi tan rung rgm nhi$t ddi thudng xanh mua am, tren sudn mii dg cao khfing qua 600-700 m,

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Nguyan Vu Bao vd Dtg Tgp chf KHOA HQC & CONG NGHE 177(01): / 9 - S 4

Tren c a sd phan tich va so sanh mgt so he thong phan loai da biet, he thong cua K.

Schumann (1891) [8] va bo sung cua Chen &

cfing sy (2011) [4] d u g c sir dung cho viec s i p xep vii mfi ta cac chi thugc tdng Danh danh {Gardenieae) — hp Cii phe d Viet N a m bao gflm 19 chi. Da sd cac dai dien cd dang than gfi, mgt sfi dang than bui; la dan cd cuong vdi chieu dai khac nhau; ciic loai deu cd la kem khac nhau ve hlnh dgng, kich thudc vii thdi gian ton tai; hoa thudng mpc thanh cym d dinh cilnh, nach lii, da sd mau 5 vii ludng tinh;

qua don, mgng vdi nhieu hinh dgng khac nhau va co dai thudng tfin tai tren qua, hat cd so lugng va kich thudc khac nhau tuy loai, giau ngi nhu. Cac loai trong cac chi thupc tflng Danh diinh (Gardenieae) - hp Ca phe phan bo hau khap ca nude, gap nhieu d mien Trung va Nam bg, Ciic dac diem sinh thai rat da dang d ca rflng thii sinh, rimg nguyen sinh.

Gia tri sfl dyng ciia cac loai thu^c tfing Danh

danh da dang, cd th^ dimg lam thuflc, cung c i p gS cho xay d y n g , lam d o gd.

TAI LIEU THAM KHAO 1. Nguyin TiSn Ban (1997), Cdm nang Ira cuuva nhdn biit cdc ho thuc vgt hgt kin & Viit Nam, Nxb Nflng n ^ i e p . Ha Ngi.

2. Nguyen Tien Ban (2005), "Rubiaceae", DanH Ivc cdc lodi thuc vdi Viit Nam, 3, tr. 82-156, Nxb Nong nghiep.

3. Pham Hoang Hfl (1999), "Rubiaceae-hp Ca phi", Cdy co Viet Nam, so 3, Nxb Tre thanh pho

Ho Chi Minh, fr, 187-236, ^ 4. Chen T. & al. (2011), "Rubiaceae". Flora of

China, Vol. 19 MBGpress.

5. Goevarts R,, et al, (2006), "Worid checklist of Rubiaceae", Royal Botanic Gardens, Kew, http://www.kew org/wcsp/rublaceae/

6. Linnaeus C. (1753), Species Plantarum, Vol I London.

7. Schumann K, (1891), "Rubiaceae", In A Engfer and K. Prantll [eds.]. Die naturlinchett Pflanzenfamilien 4: 1-156. Verlag von Vilhelm Engelmann, Leipzig, Germany.

S U M M A R Y

S T U D Y I N G O N S O M E C L A S S I F I C A T I O N C H A R A C T E R I S T I C S O F G E N E R A O F T R I B U S - Gardenieae (Rubiaceae)

IN S O M E E C O S Y S T E M S IN V I E T N A M

Nguyen Vu Bao'", Tran The Bach^ Nguyen Thi Mai^

TNU - Umversity ofEducation, ^Institute of Ecology and Biological Resources, , 'TNU- University of Agriculture and Forestry Classification system is built by Schumann (1891) and Chen et al. (2011) used to classify genera ofthe tribus Gardenieae (Rubiaceae) in Vietnam, and we found that they included 19 genera. We built a classification key and described morphological, ecological characteristics and of 19 genera o f t h e tribus Gardenieae (Rubiaceae) in Viemam, The 19 genera found in all studied areas, especially in centre and southern areas in Vietnam, Some species ofthe 19 genera are using as medicine, some provide wood for human being, etc.

Keywords: Cinchonoideae, Gardenieae. Rubiaceae, Danhdanh. Caphe

Ngdy nh^n bdi: 28/11/2017; Ngdyplidn biin: 20/12/2018; Ngdy duyft ddng: 30/01/2018 Tel- 0973 987507, Email ngiiyenvubao@dhsptn.edu v,

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